455 lines
74 KiB
XML
455 lines
74 KiB
XML
<document id="28D13BB26B4F0C020EDA729C12437770" ID-DOI="10.5281/zenodo.275727" ID-GBIF-Dataset="1e79f840-6ced-48b5-ab20-94992c6f0a88" ID-ISSN="1175-5326" ID-Zenodo-Dep="275727" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460160971924" checkinUser="plazi" docAuthor="West, Rick C. & Nunn, Steven C." docDate="2010" docId="03CC87B17657FFBA9293FDF7FD38D511" docLanguage="en" docName="zt02362p043.pdf" docOrigin="Zootaxa 2362" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lyrognathus achilles West & Nunn, 2010, sp. nov." docType="treatment" docVersion="8" lastPageNumber="19" masterDocId="FFF5FFC9765DFFA89204FFFAFF92D456" masterDocTitle="A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae" masterLastPageNumber="43" masterPageNumber="1" pageNumber="11" updateTime="1698235793967" updateUser="plazi">
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<mods:titleInfo id="CB254EE7B977A1DD42659B90458D80AF">
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<mods:title id="3CDF76F794F7EFD101332A862818A13D">A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae</mods:title>
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<mods:name id="01B2D89AE6BEBCF00CAE0B85666567D6" type="personal">
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<mods:namePart id="71B706877D81A8DDDB16AC7003C50DD9">West, Rick C.</mods:namePart>
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<mods:name id="C22A40F2B85171FFE4F2424E717C092E" type="personal">
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<mods:role id="8C0CD5924A8D7115EADC8809727B5224">
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<mods:roleTerm id="3E8583E275BDDB8450C5426A9ABCCDF9">Author</mods:roleTerm>
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<mods:namePart id="B2BB681E1C1A3E333EB7509F15FB3323">Nunn, Steven C.</mods:namePart>
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<mods:typeOfResource id="290A1A6F4F5D5A7CE34B38580D2BB304">text</mods:typeOfResource>
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<mods:title id="4BD41156447D31A132E11D2F2B16A4DE">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="615296B8B7490A8E7D4038A492FB87DF">
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<mods:date id="A75913EC3F5A32F2444012F938F2DC58">2010</mods:date>
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<mods:detail id="276BA8A4F96FC49D65AD754EFAAC8E4F" type="volume">
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<mods:number id="5F84E8803F8BF5CADDE6BD7AB2659C63">2362</mods:number>
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<mods:start id="94729B34AEB707BEEE3EA4F5C79B9A05">1</mods:start>
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<mods:end id="CEFFFF969651DB0C95110A33CD23E7BC">43</mods:end>
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<mods:identifier id="870E45F88F2565EC8958F2C859EE002E" type="DOI">10.5281/zenodo.275727</mods:identifier>
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<mods:identifier id="468E410731CF525FDB9A1A9DAD5CFC90" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="B0427DF7A0F4E3585BC00BFC6D742B18" type="Zenodo-Dep">275727</mods:identifier>
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<treatment id="03CC87B17657FFBA9293FDF7FD38D511" ID-DOI="http://doi.org/10.5281/zenodo.6210152" ID-GBIF-Taxon="127654810" ID-Zenodo-Dep="6210152" LSID="urn:lsid:plazi:treatment:03CC87B17657FFBA9293FDF7FD38D511" httpUri="http://treatment.plazi.org/id/03CC87B17657FFBA9293FDF7FD38D511" lastPageId="18" lastPageNumber="19" pageId="10" pageNumber="11">
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<subSubSection id="C37F652C7657FFA29293FDF7FEF5D61C" pageId="10" pageNumber="11" type="nomenclature">
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<paragraph id="8BDA36A77657FFA29293FDF7FE6BD671" blockId="10.[151,505,525,586]" box="[151,505,525,551]" pageId="10" pageNumber="11">
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<heading id="D09281CB7657FFA29293FDF7FE6BD671" bold="true" box="[151,505,525,551]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
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<emphasis id="B911EAB57657FFA29293FDF7FE6BD671" bold="true" box="[151,505,525,551]" pageId="10" pageNumber="11">
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<taxonomicName id="4C654D247657FFA29293FDF7FE07D671" ID-CoL="3WTQH" box="[151,405,525,551]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
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<emphasis id="B911EAB57657FFA29293FDF7FE07D671" bold="true" box="[151,405,525,551]" italics="true" pageId="10" pageNumber="11">Lyrognathus achilles</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257CE7657FFA29398FDF7FE6BD671" box="[412,505,525,551]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BDA36A77657FFA29293FDCAFEF5D61C" blockId="10.[151,505,525,586]" box="[151,359,560,586]" pageId="10" pageNumber="11">
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(
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<figureCitation id="135E2A227657FFA2929BFDCAFEB4D61C" box="[159,294,560,586]" captionStart-0="FIGURES 13 – 14. 13" captionStart-1="FIGURES 15 – 19" captionStart-2="FIGURES 20 – 23" captionStart-3="FIGURES 24 – 28" captionStart-4="FIGURES 29 – 34" captionStart-5="FIGURES 35 – 40" captionStart-6="FIGURES 41 – 44. 41" captionStartId-0="11.[151,269,1971,1995]" captionStartId-1="12.[151,269,1580,1604]" captionStartId-2="13.[151,270,1796,1820]" captionStartId-3="14.[151,269,1606,1630]" captionStartId-4="16.[151,269,1915,1939]" captionStartId-5="17.[151,270,1248,1272]" captionStartId-6="18.[151,270,1327,1351]" captionTargetBox-0="[151,1436,306,1949]" captionTargetBox-1="[151,1436,194,1556]" captionTargetBox-2="[151,1436,343,1773]" captionTargetBox-3="[151,1434,455,1581]" captionTargetBox-4="[151,1434,746,1894]" captionTargetBox-5="[151,1435,194,1226]" captionTargetBox-6="[152,1434,384,1304]" captionTargetId-0="figure@11.[151,1436,306,1950]" captionTargetId-1="figure@12.[151,1436,194,1559]" captionTargetId-2="figure@13.[151,1436,343,1775]" captionTargetId-3="figure@14.[151,1436,455,1585]" captionTargetId-4="figure@16.[151,1436,746,1894]" captionTargetId-5="figure@17.[151,1436,194,1226]" captionTargetId-6="figure@18.[152,1435,384,1305]" captionTargetPageId-0="11" captionTargetPageId-1="12" captionTargetPageId-2="13" captionTargetPageId-3="14" captionTargetPageId-4="16" captionTargetPageId-5="17" captionTargetPageId-6="18" captionText-0="FIGURES 13 – 14. 13. Lyrognathus achilles sp. nov. 13. Holotype female in life. 14. Paratype male in life. Photos: R. C. West." captionText-1="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." captionText-2="FIGURES 20 – 23. Lyrognathus achilles sp. nov., holotype female: 20. Leg IV, dorsal view. 21. Leg IV, prolateral view. 22. Tar. IV, ventral view. 23. Spermathecae, dorsal view. Scale bar = 10 mm for 20, 21; 5 mm for 22; 2 mm for 23. Photos: S. C. Nunn." captionText-3="FIGURES 24 – 28. Lyrognathus achilles sp. nov., paratype female QM S 88047: 24. Left chelicerae, retrolateral view. 25. Cheliceral strikers, ectal view. 26. Basomedial cheliceral spine, prolateral view. 27. Left chelicerae, prolateral view. 28. Intercheliceral pegs, prolateral view. Scale bar = 5 mm for 24, 25, 27; 1 mm for 26, 28. Photos: S. C. Nunn." captionText-4="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." captionText-5="FIGURES 35 – 40. Lyrognathus achilles sp. nov., paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10 mm for 35; 2 mm for 38, 39; 1 mm for 36, 37, 40. Photos: S. C. Nunn." captionText-6="FIGURES 41 – 44. 41. Lyrognathus achilles sp. nov. Upper slopes of Gunung Nibung, Borneo, Indonesia. 42. Biotope, Gunung Nibung. 43. Biotope with arrow pointing to burrow entrance. 44. Detail of burrow entrance among foliage. Photos: 41 – 44: H. Yanto." httpUri-0="https://zenodo.org/record/275731/files/figure.png" httpUri-1="https://zenodo.org/record/275732/files/figure.png" httpUri-2="https://zenodo.org/record/275733/files/figure.png" httpUri-3="https://zenodo.org/record/275734/files/figure.png" httpUri-4="https://zenodo.org/record/275735/files/figure.png" httpUri-5="https://zenodo.org/record/275736/files/figure.png" httpUri-6="https://zenodo.org/record/275737/files/figure.png" pageId="10" pageNumber="11">Figs 13–44</figureCitation>
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,
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<figureCitation id="135E2A227657FFA29336FDCAFECCD61C" box="[306,350,560,586]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="10" pageNumber="11">114</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F652C7657FFA29293FD80FE83D689" pageId="10" pageNumber="11" type="description">
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<paragraph id="8BDA36A77657FFA29293FD80FE83D689" blockId="10.[151,1437,634,2004]" pageId="10" pageNumber="11">
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<emphasis id="B911EAB57657FFA29293FD80FF7BD6C2" bold="true" box="[151,233,634,660]" pageId="10" pageNumber="11">
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<typeStatus id="54DE88057657FFA29293FD80FF4DD6C2" box="[151,223,634,660]" pageId="10" pageNumber="11">Types</typeStatus>
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:
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</emphasis>
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female
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<typeStatus id="54DE88057657FFA29349FD80FE2BD6C2" box="[333,441,634,660]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
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,
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<accessionNumber id="9436AB447657FFA293C1FD80FDB6D6C2" box="[453,548,634,660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88049" pageId="10" pageNumber="11" type="EnaNcbi">S88049</accessionNumber>
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; male
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<typeStatus id="54DE88057657FFA29077FD80FD48D6C2" box="[627,730,634,660]" pageId="10" pageNumber="11" type="paratype">paratype</typeStatus>
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<accessionNumber id="9436AB447657FFA290E7FD80FCD0D6C2" box="[739,834,634,660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88050" pageId="10" pageNumber="11" type="EnaNcbi">S88050</accessionNumber>
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;
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<specimenCount id="9D63FD2E7657FFA2914AFD80FC25D6C2" box="[846,951,634,660]" pageId="10" pageNumber="11" type="female">2 female</specimenCount>
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<typeStatus id="54DE88057657FFA291BBFD80FBA0D6C2" box="[959,1074,634,660]" pageId="10" pageNumber="11" type="paratype">paratypes</typeStatus>
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<accessionNumber id="9436AB447657FFA2963EFD80FB04D6C2" box="[1082,1174,634,660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88047" pageId="10" pageNumber="11" type="EnaNcbi">S88047</accessionNumber>
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and
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<accessionNumber id="9436AB447657FFA296D5FD80FABDD6C2" box="[1233,1327,634,660]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88048" pageId="10" pageNumber="11" type="EnaNcbi">S88048</accessionNumber>
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, Gunung Nibung (
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<geoCoordinate id="EE5150607657FFA29300FD5AFECDD6EC" box="[260,351,672,698]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="925" value="-2.1166666">2°07' S</geoCoordinate>
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,
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<geoCoordinate id="EE5150607657FFA29369FD5AFE7BD6EC" box="[365,489,672,698]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="925" value="112.0">112°00' E</geoCoordinate>
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), Kotawaringin Barat, Borneo,
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<collectingCountry id="F37276377657FFA2916EFD5AFC76D6EC" box="[874,996,672,698]" name="Indonesia" pageId="10" pageNumber="11">Indonesia</collectingCountry>
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, col. H. Yanto,
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<date id="FFDB10677657FFA296A0FD5AFAABD6EC" box="[1188,1337,672,698]" pageId="10" pageNumber="11" value="2009-04-02">2 Apr. 2009</date>
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, QM.— examined.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F652C7657FFA292C2FD10FB5DD77C" pageId="10" pageNumber="11" type="etymology">
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<paragraph id="8BDA36A77657FFA292C2FD10FB5DD77C" blockId="10.[151,1437,634,2004]" pageId="10" pageNumber="11">
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<emphasis id="B911EAB57657FFA292C2FD10FEC5D752" bold="true" box="[198,343,746,772]" pageId="10" pageNumber="11">Etymology:</emphasis>
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After the mythical Greek warrior of Homer’s poem,
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<emphasis id="B911EAB57657FFA291F7FD10FBF0D755" box="[1011,1122,746,771]" italics="true" pageId="10" pageNumber="11">The Iliad</emphasis>
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, due to the unique tarsal scopula division: distinct proximal bald patch on ventral tarsi (
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<figureCitation id="135E2A227657FFA29170FCEAFC5BD77C" box="[884,969,784,810]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="10" pageNumber="11">Fig. 19</figureCitation>
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) this species displays.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37F652C7657FFA292C2FCCFFAD4D05C" pageId="10" pageNumber="11" type="diagnosis">
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<paragraph id="8BDA36A77657FFA292C2FCCFFAD4D05C" blockId="10.[151,1437,634,2004]" pageId="10" pageNumber="11">
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<emphasis id="B911EAB57657FFA292C2FCCFFEDAD719" bold="true" box="[198,328,821,847]" pageId="10" pageNumber="11">Diagnosis:</emphasis>
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Differs from both
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<taxonomicName id="4C654D247657FFA29028FCCFFD3ED718" box="[556,684,821,846]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="crotalus">
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<emphasis id="B911EAB57657FFA29028FCCFFD3ED718" box="[556,684,821,846]" italics="true" pageId="10" pageNumber="11">L. crotalus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C654D247657FFA290EDFCCFFCF6D718" box="[745,868,821,846]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="saltator">
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<emphasis id="B911EAB57657FFA290EDFCCFFCF6D718" box="[745,868,821,846]" italics="true" pageId="10" pageNumber="11">L. saltator</emphasis>
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</taxonomicName>
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in having intercheliceral pegs (not spines), and retrolateral basomedial cheliceral spines. Differs from
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<taxonomicName id="4C654D247657FFA2911DFCA0FC0FD725" box="[793,925,858,883]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="robustus">
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<emphasis id="B911EAB57657FFA2911DFCA0FC0FD725" box="[793,925,858,883]" italics="true" pageId="10" pageNumber="11">L. robustus</emphasis>
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</taxonomicName>
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in the undivided scopula for the full length of metatarsi, third claw absent on tarsi IV and swollen distal lateral lobes of the spermathecae. Differs from
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<taxonomicName id="4C654D247657FFA29780FC7AFF6CD7E8" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
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<emphasis id="B911EAB57657FFA29780FC7AFF6CD7E8" italics="true" pageId="10" pageNumber="11">L. lessunda</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257CE7657FFA29300FC5FFEF0D7E9" box="[260,354,933,959]" pageId="10" pageNumber="11" rank="species">
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<emphasis id="B911EAB57657FFA29300FC5FFEF0D7E9" bold="true" box="[260,354,933,959]" pageId="10" pageNumber="11">sp. nov.</emphasis>
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</taxonomicNameLabel>
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in the absence of “mushroom cap” medial lobes of the spermathecae. Differs from
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<taxonomicName id="4C654D247657FFA29730FC5FFA0ED7E8" box="[1332,1436,933,958]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
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<emphasis id="B911EAB57657FFA29730FC5FFA0ED7E8" box="[1332,1436,933,958]" italics="true" pageId="10" pageNumber="11">L. fuscus</emphasis>
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||
</taxonomicName>
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<taxonomicNameLabel id="A22257CE7657FFA29293FC30FF65D7B2" box="[151,247,970,996]" pageId="10" pageNumber="11" rank="species">
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<emphasis id="B911EAB57657FFA29293FC30FF65D7B2" bold="true" box="[151,247,970,996]" pageId="10" pageNumber="11">sp. nov.</emphasis>
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</taxonomicNameLabel>
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in the more pronounced anterior maxillary lobe, distal cream/white leg segment bands, larger size, color and uniform retrolateral tibiae IV with a penicillate brush (grading distally in
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<taxonomicName id="4C654D247657FFA2966CFC0AFB42D05F" box="[1128,1232,1008,1033]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
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<emphasis id="B911EAB57657FFA2966CFC0AFB42D05F" box="[1128,1232,1008,1033]" italics="true" pageId="10" pageNumber="11">L. fuscus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A22257CE7657FFA296D3FC15FAA6D05F" box="[1239,1332,1007,1033]" pageId="10" pageNumber="11" rank="species">
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||
<emphasis id="B911EAB57657FFA296D3FC15FAA6D05F" bold="true" box="[1239,1332,1007,1033]" pageId="10" pageNumber="11">sp. nov.</emphasis>
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</taxonomicNameLabel>
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).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F652C7657FFB992C2FBEFFC61D39C" lastPageId="17" lastPageNumber="18" pageId="10" pageNumber="11" type="description">
|
||
<paragraph id="8BDA36A77657FFA292C2FBEFFC53D079" blockId="10.[151,1437,634,2004]" box="[198,961,1045,1071]" pageId="10" pageNumber="11">
|
||
<emphasis id="B911EAB57657FFA292C2FBEFFEF3D079" bold="true" box="[198,353,1045,1071]" pageId="10" pageNumber="11">Description:</emphasis>
|
||
Female
|
||
<typeStatus id="54DE88057657FFA293C2FBEFFDBED079" box="[454,556,1045,1071]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
|
||
(
|
||
<figureCitation id="135E2A227657FFA29038FBEFFD03D079" box="[572,657,1045,1071]" captionStart="FIGURES 13 – 14. 13" captionStartId="11.[151,269,1971,1995]" captionTargetBox="[151,1436,306,1949]" captionTargetId="figure@11.[151,1436,306,1950]" captionTargetPageId="11" captionText="FIGURES 13 – 14. 13. Lyrognathus achilles sp. nov. 13. Holotype female in life. 14. Paratype male in life. Photos: R. C. West." httpUri="https://zenodo.org/record/275731/files/figure.png" pageId="10" pageNumber="11">Fig. 13</figureCitation>
|
||
) with body length: 50.57.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77657FFA292C0FBC0FB24D0C9" blockId="10.[151,1437,634,2004]" pageId="10" pageNumber="11">
|
||
Color (in life,
|
||
<figureCitation id="135E2A227657FFA29376FBC0FE59D002" box="[370,459,1082,1108]" captionStart="FIGURES 13 – 14. 13" captionStartId="11.[151,269,1971,1995]" captionTargetBox="[151,1436,306,1949]" captionTargetId="figure@11.[151,1436,306,1950]" captionTargetPageId="11" captionText="FIGURES 13 – 14. 13. Lyrognathus achilles sp. nov. 13. Holotype female in life. 14. Paratype male in life. Photos: R. C. West." httpUri="https://zenodo.org/record/275731/files/figure.png" pageId="10" pageNumber="11">Fig. 13</figureCitation>
|
||
): anteriorly orange/brown, grading medially to dark brown/black posteriorly and entirely, with distinct cream/white bands on all leg segments distally. Dorsal coxae, trochantera palest of all, light grey. Ventrally dark brown/black on sternum, coxae, with legs same as seen dorsally.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77657FFA292C0FB50FB4AD1B9" blockId="10.[151,1437,634,2004]" pageId="10" pageNumber="11">
|
||
Carapace (
|
||
<figureCitation id="135E2A227657FFA29348FB50FE3FD092" box="[332,429,1194,1220]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="10" pageNumber="11">Fig. 15</figureCitation>
|
||
): length 21.45, width 13.8 (width across anterior edge 9.45). Fovea 2.71 wide, procurved, deep, smaller in width than OT. Distance from anterior carapace to fovea 13.7. Carapace with 4 discernible hair
|
||
<typeStatus id="54DE88057657FFA2935AFB0FFE0DD159" box="[350,415,1269,1295]" pageId="10" pageNumber="11">types</typeStatus>
|
||
(C1 to C4):
|
||
<typeStatus id="54DE88057657FFA29038FB0FFDE2D159" box="[572,624,1269,1295]" pageId="10" pageNumber="11">type</typeStatus>
|
||
C1, long, needle-form pallids bordering carapace entirely (darker basally):
|
||
<typeStatus id="54DE88057657FFA29300FAE0FEA5D162" box="[260,311,1306,1332]" pageId="10" pageNumber="11">type</typeStatus>
|
||
C2, short blunt-tipped, basally dark grading to distally pallid and cover entire carapace evenly, but reasonably sparse:
|
||
<typeStatus id="54DE88057657FFA293AFFABAFE4CD10C" box="[427,478,1344,1370]" pageId="10" pageNumber="11">type</typeStatus>
|
||
C3, short, thin wavy pallids, form basis of carapace mat:
|
||
<typeStatus id="54DE88057657FFA29698FABAFB5DD10C" box="[1180,1231,1344,1370]" pageId="10" pageNumber="11">type</typeStatus>
|
||
C4 emerge from what resemble bothrial collars, long, distally spiniform, pallid and lie in uniform arrangement of (>5) along anterior edge of each radial groove, (>4) along anterior edge of each thoracic groove, (>5) along anterior edge of each sulcal groove, (>10) along anterior edge of each caput groove and line of>5 along medial caput ridge (between OT and fovea).>4 more located behind each side of OT, nearest median caput row.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77657FFA292C0FA00FCB1D34C" blockId="10.[151,1437,634,2004]" pageId="10" pageNumber="11">
|
||
Eyes (
|
||
<figureCitation id="135E2A227657FFA29315FA00FEFDD242" box="[273,367,1530,1556]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="10" pageNumber="11">Fig. 16</figureCitation>
|
||
): ocular tubercle; length 2.25, width 3.0. OT highest posteriorly behind AME, sloping gradually anteriorly. Anterior row transverse, posterior row recurved. Eyes in lengths/widths: AME: 0.71/ 0.70, ALE: 0.69/0.27, PLE: 0.49/0.25, PME: 0.46/0.27. Inter-ocular distances: AME–AME: 0.29, AME– ALE: 0.21, AME–PME: 0.18, ALE–ALE: 1.85, ALE–PME: 0.38, PME–PME: 1.51, PME–PLE: 0.19, PLE– PLE: 2.35, ALE–PLE: 0.26. OT hairs:>15 (C2) directly posterior to AME’s, 4–5 between PME and center of OT (directly behind each AME). 20 (C2) along median anterior OT surface. OT entirely covered in small, weak, pallid setae (C3), directionally pointing to center of OT, posteriorly and anteriorly. Lateral small pallids very sparse with no uniform direction/random pointing.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77657FFA392C0F8DFFC10D4AA" blockId="10.[151,1437,634,2004]" lastBlockId="11.[151,1436,152,252]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
|
||
Chelicerae (left,
|
||
<figureCitation id="135E2A227657FFA2938DF8DFFD80D369" box="[393,530,1829,1855]" captionStart="FIGURES 24 – 28" captionStartId="14.[151,269,1606,1630]" captionTargetBox="[151,1434,455,1581]" captionTargetId="figure@14.[151,1436,455,1585]" captionTargetPageId="14" captionText="FIGURES 24 – 28. Lyrognathus achilles sp. nov., paratype female QM S 88047: 24. Left chelicerae, retrolateral view. 25. Cheliceral strikers, ectal view. 26. Basomedial cheliceral spine, prolateral view. 27. Left chelicerae, prolateral view. 28. Intercheliceral pegs, prolateral view. Scale bar = 5 mm for 24, 25, 27; 1 mm for 26, 28. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275734/files/figure.png" pageId="10" pageNumber="11">Figs 24–28</figureCitation>
|
||
): length 9.8, width at base of each lobe 4.25 (at bases widest point, dorsally), height: 7.1. Ectal lyrate region a series of strikers (>95) in 8 horizontal rows (
|
||
<figureCitation id="135E2A227657FFA2962BF8B0FB1AD332" box="[1071,1160,1866,1892]" captionStart="FIGURES 24 – 28" captionStartId="14.[151,269,1606,1630]" captionTargetBox="[151,1434,455,1581]" captionTargetId="figure@14.[151,1436,455,1585]" captionTargetPageId="14" captionText="FIGURES 24 – 28. Lyrognathus achilles sp. nov., paratype female QM S 88047: 24. Left chelicerae, retrolateral view. 25. Cheliceral strikers, ectal view. 26. Basomedial cheliceral spine, prolateral view. 27. Left chelicerae, prolateral view. 28. Intercheliceral pegs, prolateral view. Scale bar = 5 mm for 24, 25, 27; 1 mm for 26, 28. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275734/files/figure.png" pageId="10" pageNumber="11">Fig. 25</figureCitation>
|
||
). Strongest and longest strikers on lowest rows. Each striker very stout, blade or scimitar-shaped, with long filiform ends. Cheliceral teeth, 12,>65 basomesal teeth. Intercheliceral pegs (>9 larger,>28 smaller) in tight cluster on basodorsal surface, very stout, 1/3 as wide as long (
|
||
<figureCitation id="135E2A227657FFA29071F840FD5FD382" box="[629,717,1978,2004]" captionStart="FIGURES 24 – 28" captionStartId="14.[151,269,1606,1630]" captionTargetBox="[151,1434,455,1581]" captionTargetId="figure@14.[151,1436,455,1585]" captionTargetPageId="14" captionText="FIGURES 24 – 28. Lyrognathus achilles sp. nov., paratype female QM S 88047: 24. Left chelicerae, retrolateral view. 25. Cheliceral strikers, ectal view. 26. Basomedial cheliceral spine, prolateral view. 27. Left chelicerae, prolateral view. 28. Intercheliceral pegs, prolateral view. Scale bar = 5 mm for 24, 25, 27; 1 mm for 26, 28. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275734/files/figure.png" pageId="10" pageNumber="11">Fig. 28</figureCitation>
|
||
). On retrolateral cheliceral surface, basomedially, with small cluster of short but strong spiniform setae (>5) and 1 spine pointing distad (
|
||
<figureCitation id="135E2A227656FFA39600FF62FBC9D4E4" box="[1028,1115,152,178]" captionStart="FIGURES 24 – 28" captionStartId="14.[151,269,1606,1630]" captionTargetBox="[151,1434,455,1581]" captionTargetId="figure@14.[151,1436,455,1585]" captionTargetPageId="14" captionText="FIGURES 24 – 28. Lyrognathus achilles sp. nov., paratype female QM S 88047: 24. Left chelicerae, retrolateral view. 25. Cheliceral strikers, ectal view. 26. Basomedial cheliceral spine, prolateral view. 27. Left chelicerae, prolateral view. 28. Intercheliceral pegs, prolateral view. Scale bar = 5 mm for 24, 25, 27; 1 mm for 26, 28. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275734/files/figure.png" pageId="11" pageNumber="12">Fig. 26</figureCitation>
|
||
). Surrounding these spines, notable patch of fine setae, dark, of medium length, blends into typical thicker pallid retrolateral cheliceral setae. Setae filiform, seem to emerge from hemispherical bases.
|
||
</paragraph>
|
||
<caption id="DF1A662F7656FFA39293F849FF43D3BD" httpUri="https://zenodo.org/record/275731/files/figure.png" pageId="11" pageNumber="12" targetBox="[151,1436,306,1949]" targetPageId="11">
|
||
<paragraph id="8BDA36A77656FFA39293F849FF43D3BD" blockId="11.[151,1436,1971,2027]" pageId="11" pageNumber="12">
|
||
<emphasis id="B911EAB57656FFA39293F849FECFD39D" bold="true" box="[151,349,1971,1995]" pageId="11" pageNumber="12">FIGURES 13–14.</emphasis>
|
||
13.
|
||
<taxonomicName id="4C654D247656FFA39388F84EFDF9D39D" box="[396,619,1972,1995]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB57656FFA39388F84EFDF9D39D" box="[396,619,1972,1995]" italics="true" pageId="11" pageNumber="12">Lyrognathus achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE7656FFA39075F849FD55D39D" box="[625,711,1971,1995]" pageId="11" pageNumber="12" rank="species">
|
||
<emphasis id="B911EAB57656FFA39075F849FD55D39D" bold="true" box="[625,711,1971,1995]" pageId="11" pageNumber="12">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
13. Holotype female in life. 14. Paratype male in life. Photos: R. C. West.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A662F7651FFA49293F9D6FCD4D2D2" httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="12" pageNumber="13" targetBox="[151,1436,194,1556]" targetPageId="12">
|
||
<paragraph id="8BDA36A77651FFA49293F9D6FCD4D2D2" blockId="12.[151,1436,1580,1668]" pageId="12" pageNumber="13">
|
||
<emphasis id="B911EAB57651FFA49293F9D6FECFD212" bold="true" box="[151,349,1580,1604]" pageId="12" pageNumber="13">FIGURES 15–19.</emphasis>
|
||
<taxonomicName id="4C654D247651FFA49360F9D7FDD1D212" box="[356,579,1581,1604]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="13" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB57651FFA49360F9D7FDD1D212" box="[356,579,1581,1604]" italics="true" pageId="12" pageNumber="13">Lyrognathus achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE7651FFA4904EF9D6FD32D212" box="[586,672,1580,1604]" pageId="12" pageNumber="13" rank="species">
|
||
<emphasis id="B911EAB57651FFA4904EF9D6FD32D212" bold="true" box="[586,672,1580,1604]" pageId="12" pageNumber="13">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10mm for 15, 17, 19; 2mm for 16, 18. Photos: S. C. Nunn.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA36A77651FFA492C0F948FA0ED384" blockId="12.[151,1437,1714,2002]" pageId="12" pageNumber="13">
|
||
Maxillae (
|
||
<figureCitation id="135E2A227651FFA49339F948FE0BD29A" box="[317,409,1714,1740]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="12" pageNumber="13">Figs 17</figureCitation>
|
||
,
|
||
<figureCitation id="135E2A227651FFA493A2F948FE5AD29A" box="[422,456,1714,1740]" captionStart="FIGURES 29 – 34" captionStartId="16.[151,269,1915,1939]" captionTargetBox="[151,1434,746,1894]" captionTargetId="figure@16.[151,1436,746,1894]" captionTargetPageId="16" captionText="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="12" pageNumber="13">29</figureCitation>
|
||
): prolaterally plano-convex, anterior lobe well pronounced, many cuspules (
|
||
<emphasis id="B911EAB57651FFA49756F948FAF4D29D" box="[1362,1382,1714,1739]" italics="true" pageId="12" pageNumber="13">></emphasis>
|
||
140) on inner basoventral surface.
|
||
<taxonomicName id="4C654D247651FFA4900EF922FDD1D2A4" box="[522,579,1752,1778]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="12" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
|
||
(
|
||
<figureCitation id="135E2A227651FFA49051F922FD7BD2A4" box="[597,745,1752,1778]" captionStart="FIGURES 29 – 34" captionStartId="16.[151,269,1915,1939]" captionTargetBox="[151,1434,746,1894]" captionTargetId="figure@16.[151,1436,746,1894]" captionTargetPageId="16" captionText="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="12" pageNumber="13">Figs 29, 30</figureCitation>
|
||
): bacilliform rods (>145) form a dense, ovoid patch on prolateral maxillary surface (2.85 long, 1.71 high), lowest row with up to>11 bacillae, longest rods medially in lowest row. Rods distally paddle-shaped (paddles up to 0.45 long, paddles with proximal gradation into shaft, extends for full shaft length) with short to medium length shafts (total rod length including paddles 0.69), largest rods with no distal blades. At widest point, lyrate patch 13 rows deep with smallest rods dorsally. Posterior end of patch truncate but rounded, anterior end rounded. Immediately above maxillary suture>45 elongated spines on anterior margin, rows disordered. Labium (
|
||
<figureCitation id="135E2A227651FFA49615F842FBFBD384" box="[1041,1129,1976,2002]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="12" pageNumber="13">Fig. 18</figureCitation>
|
||
): length 2.51, width 3.35.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77650FFA59293FF62FEE0D574" blockId="13.[151,1437,152,290]" pageId="13" pageNumber="14">
|
||
Many small cuspules (>560) along anterior 1/4 surface. Labium with 2 discernible hair
|
||
<typeStatus id="54DE88057650FFA596A9FF62FB7ED4E4" box="[1197,1260,152,178]" pageId="13" pageNumber="14">types</typeStatus>
|
||
(L1, L2);
|
||
<typeStatus id="54DE88057650FFA5976DFF62FA0ED4E4" box="[1385,1436,152,178]" pageId="13" pageNumber="14">type</typeStatus>
|
||
L1: long, reasonably dark spiniform, curving distally toward anterior, most concentrated along lateral borders. Setae clearly longest along anterior edge (>35);
|
||
<typeStatus id="54DE88057650FFA590DFFF18FC9DD4AA" box="[731,783,226,252]" pageId="13" pageNumber="14">type</typeStatus>
|
||
L2: 1/2 length of L1, interspersed with L1 but darker, more needle-form.
|
||
</paragraph>
|
||
<caption id="DF1A662F7650FFA59293F8FEFE87D30D" httpUri="https://zenodo.org/record/275733/files/figure.png" pageId="13" pageNumber="14" targetBox="[151,1436,343,1773]" targetPageId="13">
|
||
<paragraph id="8BDA36A77650FFA59293F8FEFE87D30D" blockId="13.[151,1437,1796,1883]" pageId="13" pageNumber="14">
|
||
<emphasis id="B911EAB57650FFA59293F8FEFECCD34A" bold="true" box="[151,350,1796,1820]" pageId="13" pageNumber="14">FIGURES 20–23.</emphasis>
|
||
<taxonomicName id="4C654D247650FFA59361F8FFFDD7D34A" box="[357,581,1797,1820]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB57650FFA59361F8FFFDD7D34A" box="[357,581,1797,1820]" italics="true" pageId="13" pageNumber="14">Lyrognathus achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE7650FFA59048F8FEFD31D34A" box="[588,675,1796,1820]" pageId="13" pageNumber="14" rank="species">
|
||
<emphasis id="B911EAB57650FFA59048F8FEFD31D34A" bold="true" box="[588,675,1796,1820]" pageId="13" pageNumber="14">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype female: 20. Leg IV, dorsal view. 21. Leg IV, prolateral view. 22. Tar. IV, ventral view. 23. Spermathecae, dorsal view. Scale bar = 10mm for 20, 21; 5mm for 22; 2mm for 23. Photos: S. C. Nunn.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA36A77650FFA692C0F870FD05D5C4" blockId="13.[151,1436,1930,2031]" lastBlockId="14.[151,1437,152,402]" lastPageId="14" lastPageNumber="15" pageId="13" pageNumber="14">
|
||
Sternum (
|
||
<figureCitation id="135E2A227650FFA59333F870FE1DD3F2" box="[311,399,1930,1956]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="13" pageNumber="14">Fig. 17</figureCitation>
|
||
): length 9.58, width 7.35, saddle-form; with 5 discernible hair
|
||
<typeStatus id="54DE88057650FFA59674F870FB3CD3F2" box="[1136,1198,1930,1956]" pageId="13" pageNumber="14">types</typeStatus>
|
||
(S1 to S5):
|
||
<typeStatus id="54DE88057650FFA5973FF870FAFFD3F2" box="[1339,1389,1930,1956]" pageId="13" pageNumber="14">type</typeStatus>
|
||
S1, longest, thick, darker basally, becoming pallid distally, entirely over surface, sparse:
|
||
<typeStatus id="54DE88057650FFA596C7F855FB6BD39F" box="[1219,1273,1967,1993]" pageId="13" pageNumber="14">type</typeStatus>
|
||
S2, elongate spiniforms, dark basally, distally pallid, on margins, most concentrated posteriorly:
|
||
<typeStatus id="54DE88057650FFA59669F82FFB0DD3B9" box="[1133,1183,2005,2031]" pageId="13" pageNumber="14">type</typeStatus>
|
||
S3, short spiniforms, entirely, sparse:
|
||
<typeStatus id="54DE88057653FFA69353FF62FE18D4E4" box="[343,394,152,178]" pageId="14" pageNumber="15">type</typeStatus>
|
||
S4, short, thin pallids, form basis of mat:
|
||
<typeStatus id="54DE88057653FFA6917AFF62FC23D4E4" box="[894,945,152,178]" pageId="14" pageNumber="15">type</typeStatus>
|
||
S5 dark basally, but apically pallid, short and blunt tipped, entirely cover in sparse disordered arrangement. Posteriorly between left and right coxae IV, border highly acuminate, lateral points slightly acuminate. Sigilla (
|
||
<figureCitation id="135E2A227653FFA691CBFF18FBBED4AA" box="[975,1068,226,252]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="14" pageNumber="15">Fig. 17</figureCitation>
|
||
): 3 pairs (not including labiosternal sigilla), posterior, large, right: length 1.38, width 0.75, left with length 1.35, width 0.65. Ovular, 1.76 apart, 1.65 to 1.7 from margin. Median pair, 1/2 size of posterior, similar shape and 0.7–1.0 of their length from sternal margin. Anterior pair large, as big as medials, somewhat obscured and border margin. Labiosternal sigilla largest, 1.6 times size of PSS.
|
||
</paragraph>
|
||
<caption id="DF1A662F7653FFA69293F9BCFA96D2CB" httpUri="https://zenodo.org/record/275734/files/figure.png" pageId="14" pageNumber="15" targetBox="[151,1434,455,1581]" targetPageId="14">
|
||
<paragraph id="8BDA36A77653FFA69293F9BCFA96D2CB" blockId="14.[151,1436,1606,1693]" pageId="14" pageNumber="15">
|
||
<emphasis id="B911EAB57653FFA69293F9BCFECFD208" bold="true" box="[151,349,1606,1630]" pageId="14" pageNumber="15">FIGURES 24–28.</emphasis>
|
||
<taxonomicName id="4C654D247653FFA69360F9BDFDD0D208" box="[356,578,1607,1630]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB57653FFA69360F9BDFDD0D208" box="[356,578,1607,1630]" italics="true" pageId="14" pageNumber="15">Lyrognathus achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE7653FFA6904CF9BCFD0FD208" box="[584,669,1606,1630]" pageId="14" pageNumber="15" rank="species">
|
||
<emphasis id="B911EAB57653FFA6904CF9BCFD0FD208" bold="true" box="[584,669,1606,1630]" pageId="14" pageNumber="15">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype female QM
|
||
<accessionNumber id="9436AB447653FFA69189F9BCFC71D20B" box="[909,995,1606,1629]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88047" pageId="14" pageNumber="15" type="EnaNcbi">S88047</accessionNumber>
|
||
: 24. Left chelicerae, retrolateral view. 25. Cheliceral strikers, ectal view. 26. Basomedial cheliceral spine, prolateral view. 27. Left chelicerae, prolateral view. 28. Intercheliceral pegs, prolateral view. Scale bar = 5mm for 24, 25, 27; 1mm for 26, 28. Photos: S. C. Nunn.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA36A77653FFA692C0F936FAF0D3F7" blockId="14.[151,1437,1740,2027]" pageId="14" pageNumber="15">
|
||
Legs: formula (length); IV, I, II, III: (width); IV, III, I, II. Leg
|
||
<collectingCountry id="F37276377653FFA69196F936FC2BD2B0" box="[914,953,1740,1766]" name="Russia" pageId="14" pageNumber="15">RF</collectingCountry>
|
||
~84.51. Leg lengths (fem., pat., tib., met., tar., total): palp: 9.46, 5.98, 6.74, 0.00, 5.75, 27.93. I: 13.21, 8.87, 9.98, 9.15, 5.34, 46.55. II: 11.64, 7.96, 8.48, 7.59, 5.22, 40.89. III: 9.64, 6.79, 6.78, 8.57, 5.01, 36.79. IV: 15.55, 8.44, 11.33, 14.16, 5.6, 55.08. Leg widths (fem., pat., tib., met., tar., total.): palp: 2.15, 2.71, 2.42, 0.00, 2.54, 9.82. I: 3.38, 3.37, 2.99, 2.34, 2.14, 14.22. II: 3.37, 3.26, 2.79, 2.35, 2.23, 14.00. III: 4.07, 3.57, 3.05, 2.33, 2.18, 15.20. IV: 4.08, 3.61, 4.05, 2.83, 2.49, 17.06.Tib. IV with strong retrolateral villous setal fringe, proximally to distally uniform–entire (
|
||
<figureCitation id="135E2A227653FFA696F8F87DFAC6D3F7" box="[1276,1364,1927,1953]" captionStart="FIGURES 20 – 23" captionStartId="13.[151,270,1796,1820]" captionTargetBox="[151,1436,343,1773]" captionTargetId="figure@13.[151,1436,343,1775]" captionTargetPageId="13" captionText="FIGURES 20 – 23. Lyrognathus achilles sp. nov., holotype female: 20. Leg IV, dorsal view. 21. Leg IV, prolateral view. 22. Tar. IV, ventral view. 23. Spermathecae, dorsal view. Scale bar = 10 mm for 20, 21; 5 mm for 22; 2 mm for 23. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275733/files/figure.png" pageId="14" pageNumber="15">Fig. 20</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77653FFA692C0F856FB7DD3BD" blockId="14.[151,1437,1740,2027]" pageId="14" pageNumber="15">
|
||
Scopula: met. and tar. I–III undivided, met. IV undivided, tar. IV (
|
||
<figureCitation id="135E2A227653FFA6961CF856FBE4D390" box="[1048,1142,1964,1990]" captionStart="FIGURES 20 – 23" captionStartId="13.[151,270,1796,1820]" captionTargetBox="[151,1436,343,1773]" captionTargetId="figure@13.[151,1436,343,1775]" captionTargetPageId="13" captionText="FIGURES 20 – 23. Lyrognathus achilles sp. nov., holotype female: 20. Leg IV, dorsal view. 21. Leg IV, prolateral view. 22. Tar. IV, ventral view. 23. Spermathecae, dorsal view. Scale bar = 10 mm for 20, 21; 5 mm for 22; 2 mm for 23. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275733/files/figure.png" pageId="14" pageNumber="15">Fig. 22</figureCitation>
|
||
) only partially divided proximally (by one row of sparse setae>5, nonexistent in distal 1/2). Met. I–IV, for full length.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0FF62FED2D58A" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Coxae: some small black thorns prolatero-dorsally, no thorns retrolaterally on I–IV. Coxae easily seen dorsally. Cox. I longest,
|
||
<emphasis id="B911EAB57652FFA793BDFF47FE4ED480" box="[441,476,189,214]" italics="true" pageId="15" pageNumber="16">ca.</emphasis>
|
||
1.2 times length of II. IV widest, as long as III, basally rectangular with rounded corners. Coxae with small ventral thorns prolaterally on I–IV. I–III ventrally with many long thick blunt setae proximally, pallid. No short black setae. IV with mixture of long thick blunt setae entirely, pallid intermixed with shorter thin pallid setae. Ventral surface of coxae I–IV gently sloping anteriorly. Retrolateral setation: I– III with median narrow light brush, IV entirely covered in short thin pallid setae. I–IV retrolaterally lack ventral ledge. Ventral measurements for coxae: palp–length 9.78, width 4.94; I–9.18, 4.16; II–7.53, 3.63; III– 6.24, 4.02; IV–7.53, 4.96. Trochantera: palp–length 2.85, width 2.9; I–3.31, 4.06; II–2.9, 3.88; III–3.26, 3.6; IV–4.14, 4.11.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0FE12FC4CD721" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Trichobothria: on all tarsi basal filiform field slightly wider than clavate field, merges evenly. Clavates on tar. I in distal 1/2 (very dark), long filiforms only in basal 2/3, shorter filiforms intermixed with clavates distally. Clavate extent on tar. II–IV
|
||
<emphasis id="B911EAB57652FFA79057FDC8FDE3D61D" box="[595,625,562,587]" italics="true" pageId="15" pageNumber="16">cf.</emphasis>
|
||
I, in distal 1/2. Shorter filiforms for length. Short epitrichobothrial field on I shorter than clavates, increasing in length proximally. Tarsal organ evident on legs II–IV under stereo microscope (up to
|
||
<date id="FFDB10677652FFA793C2FD87FE64D6C1" box="[454,502,637,663]" pageId="15" pageNumber="16" value="1964-10">x64</date>
|
||
), coniform without noticeable ridges. On tar. I, entire field 4.56 long and 1.32 wide distally, to 1.85 wide proximally. On tar. IV, field 4.78 long and 0.89 wide distally, increasing to 1.95 proximally. Metatarsi: trichobothrial field not evident laterally or proximally in any form. Tibia: on tib. I prolaterally and proximally, short, possibly clavate trichobothria noted (>8 per
|
||
<quantity id="4C9D9B427652FFA7963FFD17FB0ED751" box="[1083,1180,749,775]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="mm" value="0.25">0.25mm</quantity>
|
||
squared) in L-shaped patch 0.25 long, 1.25 wide, clearly pallid:>6 filiforms in semicircular pattern proximally to rest of patch. On tib. IV prolaterally and proximally, short dark clavates (>7 per
|
||
<quantity id="4C9D9B427652FFA79170FCC2FC47D704" box="[884,981,824,850]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="15" pageNumber="16" unit="mm" value="0.25">0.25mm</quantity>
|
||
squared) in patch 1.66 long, 0.22 wide distinct:>7 filiforms in semicircular pattern proximally to rest of patch.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0FC78FCD4D794" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">Spines: met. I with 1 DV, met. II with 1 DV, 1 DPV, 1 DRV, met. III with 2 DV, 1DPV, 1 DRV, 1 DD, 1 DPL and met. IV with 2 DV, 1DPV, 1 DRV, 1 DD, 1 DPD.</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0FC37FBCED7B1" blockId="15.[151,1437,152,1970]" box="[196,1116,973,999]" pageId="15" pageNumber="16">Claws: unarmed paired claws on all legs. Reduced third claw absent on leg IV.</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0FC08FE97D161" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Abdomen (
|
||
<figureCitation id="135E2A227652FFA7934FFC08FE35D05A" box="[331,423,1010,1036]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="15" pageNumber="16">Fig. 19</figureCitation>
|
||
): ovular, elongated, yellow brown (in alcohol); with 4 discernible hair
|
||
<typeStatus id="54DE88057652FFA79702FC08FAD7D05A" box="[1286,1349,1010,1036]" pageId="15" pageNumber="16">types</typeStatus>
|
||
(A1 to A4): Dorsally with 3 hair
|
||
<typeStatus id="54DE88057652FFA793CDFBE2FD99D064" box="[457,523,1048,1074]" pageId="15" pageNumber="16">types</typeStatus>
|
||
:
|
||
<typeStatus id="54DE88057652FFA79013FBE2FDDBD064" box="[535,585,1048,1074]" pageId="15" pageNumber="16">type</typeStatus>
|
||
A1 long, dark blunt tipped, uniform:
|
||
<typeStatus id="54DE88057652FFA79602FBE2FBABD064" box="[1030,1081,1048,1074]" pageId="15" pageNumber="16">type</typeStatus>
|
||
A2 mid-length, also dark, but distally spiniform, unlike
|
||
<typeStatus id="54DE88057652FFA793CEFBC7FE6FD001" box="[458,509,1085,1111]" pageId="15" pageNumber="16">type</typeStatus>
|
||
A1:
|
||
<typeStatus id="54DE88057652FFA7903DFBC7FDFED001" box="[569,620,1085,1111]" pageId="15" pageNumber="16">type</typeStatus>
|
||
A3 form dense mat, mid-length, uniformly dark, needle-form (some stand erect, some posteriorly, but share similar morphology). Longest hairs (
|
||
<typeStatus id="54DE88057652FFA79647FB98FBEBD02A" box="[1091,1145,1122,1148]" pageId="15" pageNumber="16">type</typeStatus>
|
||
A1) more concentrated posteriorly toward spinnerets, point distad, but toward proximal region, hairs stand erect, medially curved (toward posterior). Ventral hair
|
||
<typeStatus id="54DE88057652FFA79010FB57FDC1D091" box="[532,595,1197,1223]" pageId="15" pageNumber="16">types</typeStatus>
|
||
almost same as dorsal, although
|
||
<typeStatus id="54DE88057652FFA791E6FB57FB87D091" box="[994,1045,1197,1223]" pageId="15" pageNumber="16">type</typeStatus>
|
||
A2 and A3 more dense entirely.
|
||
<typeStatus id="54DE88057652FFA79293FB28FF43D0BA" box="[151,209,1234,1260]" pageId="15" pageNumber="16">Type</typeStatus>
|
||
A4 hair in very dense patch, trapezoid, posterior of pedicel between median regions of cox. IV. Hairs of medium length, wavy and pallid. Between anterior pair of booklungs, setal mat uniform with rest of venter (
|
||
<figureCitation id="135E2A227652FFA7929BFAE7FF64D161" box="[159,246,1309,1335]" captionStart="FIGURES 15 – 19" captionStartId="12.[151,269,1580,1604]" captionTargetBox="[151,1436,194,1556]" captionTargetId="figure@12.[151,1436,194,1559]" captionTargetPageId="12" captionText="FIGURES 15 – 19. Lyrognathus achilles sp. nov., holotype female: 15. Carapace, dorsal view. 16. Eyes, dorsal view. 17. Sternum, labium, maxillae and coxa, ventral view. 18. Labium, ventral view. 19. Abdomen, legs IV, ventral view. Scale bar = 10 mm for 15, 17, 19; 2 mm for 16, 18. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275732/files/figure.png" pageId="15" pageNumber="16">Fig. 19</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0FAB8FD97D241" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Genitalia (
|
||
<figureCitation id="135E2A227652FFA79344FAB8FE0BD10A" box="[320,409,1346,1372]" captionStart="FIGURES 20 – 23" captionStartId="13.[151,270,1796,1820]" captionTargetBox="[151,1436,343,1773]" captionTargetId="figure@13.[151,1436,343,1775]" captionTargetPageId="13" captionText="FIGURES 20 – 23. Lyrognathus achilles sp. nov., holotype female: 20. Leg IV, dorsal view. 21. Leg IV, prolateral view. 22. Tar. IV, ventral view. 23. Spermathecae, dorsal view. Scale bar = 10 mm for 20, 21; 5 mm for 22; 2 mm for 23. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275733/files/figure.png" pageId="15" pageNumber="16">Fig. 23</figureCitation>
|
||
): epigastric fold 6.51 wide. Spermathecae paired but not fused, separated by 0.8 (width including spermathecae 2.4), bilobular apically, each lobe with strong apical rounded appendages, lateral/ outer lobes distally swollen, ovular: lateral lobe distal widths 0.45 (left)–0.41 (right): medial lobe distal widths 0.26 (left)–0.27 (right), heavily sclerotized gradually transition to weakly sclerotized shafts proximally. Lobes apically swollen, lateral lobes (including shaft length) at 4/5 length to medials. Epigastic fold extends to
|
||
<emphasis id="B911EAB57652FFA7975FFA22FAECD1A7" box="[1371,1406,1496,1521]" italics="true" pageId="15" pageNumber="16">ca.</emphasis>
|
||
2/ 3 length of spermathecal lobes.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0F9D8FCF6D2FA" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Spinnerets: PMS: length 2.1, width (medially) 0.95. PLS: section lengths; basal 3.45, medial 1.45, apical 2.55. PLS section widths (medially); basal 1.78, medial 1.32, apical 0.85. Spinnerets with 2 discernible hair
|
||
<typeStatus id="54DE88057652FFA79293F997FF47D2D1" box="[151,213,1645,1671]" pageId="15" pageNumber="16">types</typeStatus>
|
||
(SP1, SP2): SP1 very short and distally blunt pallid, covers both pairs ventrally, second form (SP2), very similar but up to three times longer, dorsally on all segments.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0F942FC63D284" blockId="15.[151,1437,152,1970]" box="[196,1009,1720,1746]" pageId="15" pageNumber="16">
|
||
Male
|
||
<typeStatus id="54DE88057652FFA79303F942FEFED284" box="[263,364,1720,1746]" pageId="15" pageNumber="16" type="paratype">paratype</typeStatus>
|
||
(
|
||
<figureCitation id="135E2A227652FFA79378F942FE41D284" box="[380,467,1720,1746]" captionStart="FIGURES 13 – 14. 13" captionStartId="11.[151,269,1971,1995]" captionTargetBox="[151,1436,306,1949]" captionTargetId="figure@11.[151,1436,306,1950]" captionTargetPageId="11" captionText="FIGURES 13 – 14. 13. Lyrognathus achilles sp. nov. 13. Holotype female in life. 14. Paratype male in life. Photos: R. C. West." httpUri="https://zenodo.org/record/275731/files/figure.png" pageId="15" pageNumber="16">Fig. 14</figureCitation>
|
||
, as in female except) with body length: 34.09.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0F927FB19D34A" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Color (in life,
|
||
<figureCitation id="135E2A227652FFA79368F927FE51D2A1" box="[364,451,1757,1783]" captionStart="FIGURES 13 – 14. 13" captionStartId="11.[151,269,1971,1995]" captionTargetBox="[151,1436,306,1949]" captionTargetId="figure@11.[151,1436,306,1950]" captionTargetPageId="11" captionText="FIGURES 13 – 14. 13. Lyrognathus achilles sp. nov. 13. Holotype female in life. 14. Paratype male in life. Photos: R. C. West." httpUri="https://zenodo.org/record/275731/files/figure.png" pageId="15" pageNumber="16">Fig. 14</figureCitation>
|
||
): With brass sheen to dorsal carapace, coxae and trochantera hairs, also less distinct in color anteriorly to posteriorly, clearly paler distal leg segments (pat., tib., met., tar.).
|
||
</paragraph>
|
||
<paragraph id="8BDA36A77652FFA792C0F8D2FECCD3E4" blockId="15.[151,1437,152,1970]" pageId="15" pageNumber="16">
|
||
Carapace (
|
||
<figureCitation id="135E2A227652FFA79342F8D2FE30D314" box="[326,418,1832,1858]" captionStart="FIGURES 29 – 34" captionStartId="16.[151,269,1915,1939]" captionTargetBox="[151,1434,746,1894]" captionTargetId="figure@16.[151,1436,746,1894]" captionTargetPageId="16" captionText="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="15" pageNumber="16">Fig. 31</figureCitation>
|
||
): length 13.85, and width 13.1 (width across anterior edge 7.35). Fovea 1.68 wide, procurved, deep, narrower than OT. Distance from anterior carapace to fovea, 9.89. Carapace hair
|
||
<typeStatus id="54DE88057652FFA7971BF8B7FACFD331" box="[1311,1373,1869,1895]" pageId="15" pageNumber="16">types</typeStatus>
|
||
as in female except C1, C2 and C4 are brass color, iridescent in alcohol. Eyes (
|
||
<figureCitation id="135E2A227652FFA7961DF888FBE7D3DA" box="[1049,1141,1906,1932]" captionStart="FIGURES 29 – 34" captionStartId="16.[151,269,1915,1939]" captionTargetBox="[151,1434,746,1894]" captionTargetId="figure@16.[151,1436,746,1894]" captionTargetPageId="16" captionText="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="15" pageNumber="16">Fig. 32</figureCitation>
|
||
): ocular tubercle; length 1.75, width 2.43.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A7764DFFB892C0FF62FEF5D574" blockId="16.[151,1436,152,700]" pageId="16" pageNumber="17">
|
||
Chelicerae (right,
|
||
<figureCitation id="135E2A22764DFFB8939FFF62FE66D4E4" box="[411,500,152,178]" captionStart="FIGURES 35 – 40" captionStartId="17.[151,270,1248,1272]" captionTargetBox="[151,1435,194,1226]" captionTargetId="figure@17.[151,1436,194,1226]" captionTargetPageId="17" captionText="FIGURES 35 – 40. Lyrognathus achilles sp. nov., paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10 mm for 35; 2 mm for 38, 39; 1 mm for 36, 37, 40. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275736/files/figure.png" pageId="16" pageNumber="17">Fig. 36</figureCitation>
|
||
): length 7.89, width at base of each lobe 3.35 (at bases widest point, dorsally). Ectal lyrate region a series of strikers (>95) in 8 horizontal rows. Cheliceral teeth, 12,>65 basomesal teeth. Intercheliceral pegs (>9 larger,>17 smaller) in tight cluster on basodorsal surface, pegs more elongate than in females (
|
||
<figureCitation id="135E2A22764DFFB89306FEF2FEC8D574" box="[258,346,264,290]" captionStart="FIGURES 35 – 40" captionStartId="17.[151,270,1248,1272]" captionTargetBox="[151,1435,194,1226]" captionTargetId="figure@17.[151,1436,194,1226]" captionTargetPageId="17" captionText="FIGURES 35 – 40. Lyrognathus achilles sp. nov., paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10 mm for 35; 2 mm for 38, 39; 1 mm for 36, 37, 40. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275736/files/figure.png" pageId="16" pageNumber="17">Fig. 37</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36A7764DFFB892C0FED7FE11D671" blockId="16.[151,1436,152,700]" pageId="16" pageNumber="17">
|
||
Maxillae.
|
||
<taxonomicName id="4C654D24764DFFB89339FED7FEE1D511" box="[317,371,301,327]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
|
||
(
|
||
<figureCitation id="135E2A22764DFFB89380FED7FE4CD511" box="[388,478,301,327]" captionStart="FIGURES 29 – 34" captionStartId="16.[151,269,1915,1939]" captionTargetBox="[151,1434,746,1894]" captionTargetId="figure@16.[151,1436,746,1894]" captionTargetPageId="16" captionText="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="16" pageNumber="17">Fig. 30</figureCitation>
|
||
): large number of bacilliform rods (>170) form dense, ovoid patch on prolateral surface (2.45 long, 1.35 high), with lowest row composing of>14 bacillae, longest rods in center of lowest row. Rods distally paddle-shaped (paddles to 0.24 long), total length including paddles to 0.41, largest with no distal blades. At widest point, lyrate patch 12 rows deep, smallest rods dorsally. Posterior end of patch truncate but rounded, anterior end widely rounded. Immediately above suture>35 spiniform setae on anterior margin in disordered rows. Labium (
|
||
<figureCitation id="135E2A22764DFFB89052FE12FD21D654" box="[598,691,488,514]" captionStart="FIGURES 29 – 34" captionStartId="16.[151,269,1915,1939]" captionTargetBox="[151,1434,746,1894]" captionTargetId="figure@16.[151,1436,746,1894]" captionTargetPageId="16" captionText="FIGURES 29 – 34. Lyrognathus achilles sp. nov., paratype female QM S 88047 for 29, L. achilles sp. nov. paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10 mm for 34; 5 mm for 31; 2 mm for 29, 30, 32, 33. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="16" pageNumber="17">Fig. 33</figureCitation>
|
||
): length 1.73, width 2.39. Many small cuspules (>420) along anterior 1/4 surface.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A7764DFFB892C0FDC8FD05D6EA" blockId="16.[151,1436,152,700]" pageId="16" pageNumber="17">
|
||
Sternum (
|
||
<figureCitation id="135E2A22764DFFB89333FDC8FE1DD61A" box="[311,399,562,588]" captionStart="FIGURES 35 – 40" captionStartId="17.[151,270,1248,1272]" captionTargetBox="[151,1435,194,1226]" captionTargetId="figure@17.[151,1436,194,1226]" captionTargetPageId="17" captionText="FIGURES 35 – 40. Lyrognathus achilles sp. nov., paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10 mm for 35; 2 mm for 38, 39; 1 mm for 36, 37, 40. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275736/files/figure.png" pageId="16" pageNumber="17">Fig. 35</figureCitation>
|
||
): length 6.34, width 5.53, saddle-form. Sigilla: 3 pairs, posterior large; right with length 0.89, width 0.43, left with length 0.88, width 0.41. Ovular, 1.67 apart, 1.12–1.18 from margin. Median pair 1/ 2 size of posterior, similar shape, 0.5–0.8 of their length from margin. Anterior pair large, as in female. Labiosternal sigilla largest, 1.6 times size of PSS.
|
||
</paragraph>
|
||
<caption id="DF1A662F764DFFB89293F881FE71D3A5" httpUri="https://zenodo.org/record/275735/files/figure.png" pageId="16" pageNumber="17" targetBox="[151,1434,746,1894]" targetPageId="16">
|
||
<paragraph id="8BDA36A7764DFFB89293F881FE71D3A5" blockId="16.[151,1436,1915,2035]" pageId="16" pageNumber="17">
|
||
<emphasis id="B911EAB5764DFFB89293F881FECFD3C5" bold="true" box="[151,349,1915,1939]" pageId="16" pageNumber="17">FIGURES 29–34.</emphasis>
|
||
<taxonomicName id="4C654D24764DFFB89360F886FDD0D3C5" box="[356,578,1916,1939]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB5764DFFB89360F886FDD0D3C5" box="[356,578,1916,1939]" italics="true" pageId="16" pageNumber="17">Lyrognathus achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE764DFFB8904DF881FD0CD3C5" box="[585,670,1915,1939]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B911EAB5764DFFB8904DF881FD0CD3C5" bold="true" box="[585,670,1915,1939]" pageId="16" pageNumber="17">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype female QM
|
||
<accessionNumber id="9436AB44764DFFB8918BF886FC72D3C5" box="[911,992,1916,1939]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88047" pageId="16" pageNumber="17" type="EnaNcbi">S88047</accessionNumber>
|
||
for 29,
|
||
<taxonomicName id="4C654D24764DFFB89630F886FB31D3C5" box="[1076,1187,1916,1939]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB5764DFFB89630F886FB31D3C5" box="[1076,1187,1916,1939]" italics="true" pageId="16" pageNumber="17">L. achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE764DFFB896AEF881FB6DD3C5" box="[1194,1279,1915,1939]" pageId="16" pageNumber="17" rank="species">
|
||
<emphasis id="B911EAB5764DFFB896AEF881FB6DD3C5" bold="true" box="[1194,1279,1915,1939]" pageId="16" pageNumber="17">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
paratype male all others: 29. Left maxillae, prolateral view. 30. Right maxillae, prolateral view. 31. Carapace, dorsal view. 32. Eyes, dorsal view. 33. Labium, ventral view. 34. Leg IV, prolateral view. Scale bar = 10mm for 34; 5mm for 31; 2mm for 29, 30, 32, 33. Photos: S. C. Nunn.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF1A662F764CFFB99293FB1AFEF9D121" httpUri="https://zenodo.org/record/275736/files/figure.png" pageId="17" pageNumber="18" targetBox="[151,1435,194,1226]" targetPageId="17">
|
||
<paragraph id="8BDA36A7764CFFB99293FB1AFEF9D121" blockId="17.[151,1437,1248,1399]" pageId="17" pageNumber="18">
|
||
<emphasis id="B911EAB5764CFFB99293FB1AFECCD0AE" bold="true" box="[151,350,1248,1272]" pageId="17" pageNumber="18">FIGURES 35–40.</emphasis>
|
||
<taxonomicName id="4C654D24764CFFB99361FB1AFDD7D0A1" box="[357,581,1248,1271]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
|
||
<emphasis id="B911EAB5764CFFB99361FB1AFDD7D0A1" box="[357,581,1248,1271]" italics="true" pageId="17" pageNumber="18">Lyrognathus achilles</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A22257CE764CFFB99048FB1AFD30D0AE" box="[588,674,1248,1272]" pageId="17" pageNumber="18" rank="species">
|
||
<emphasis id="B911EAB5764CFFB99048FB1AFD30D0AE" bold="true" box="[588,674,1248,1272]" pageId="17" pageNumber="18">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10mm for 35; 2mm for 38, 39; 1mm for 36, 37, 40. Photos: S. C. Nunn.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BDA36A7764CFFB992C0FA5FFE86D2BC" blockId="17.[151,1437,1445,1994]" pageId="17" pageNumber="18">
|
||
Legs: formula (length); IV, I, II, III: (width); III, IV, I, II. Leg
|
||
<collectingCountry id="F3727637764CFFB99196FA5FFC2AD1E9" box="[914,952,1445,1471]" name="Russia" pageId="17" pageNumber="18">RF</collectingCountry>
|
||
~93.14. Leg lengths (fem., pat., tib., met., tar., total): palp: 8.83, 5.0, 6.77, 0.00, 2.13, 22.73. I: 12.25, 6.83, 10.93, 9.78, 5.26, 45.05. II: 10.66, 6.38, 8.54, 8.17, 4.88, 38.63. III: 8.51, 5.26, 6.72, 7.8, 4.27, 32.56. IV: 13.28, 6.84, 10.38, 12.68, 5.19, 48.37. Leg widths (fem., pat., tib., met., tar., total.): palp: 2.22, 1.92, 1.90, 0.00, 1.97, 8.01. I: 2.9, 2.53, 2.13, 1.52, 1.6, 10.68. II: 2.8, 2.56, 1.91, 1.35, 1.43, 10.05. III: 3.73, 2.7, 2.36, 1.55, 1.56, 11.9. IV: 2.75, 2.58, 2.39, 1.72, 1.77, 11.21. Tib. IV with retrolateral villous setal fringe, proximally to distally uniform, not as elongate or dense as in females. Ventral measurements for coxae: palp–length 5.41, width 2.57; I–5.89, 2.67; II–5.45, 2.67; III–3.89, 2.78; IV–5.1, 3.78. Trochantera: palp–length 2.57, width 2.1; I–3.34, 3.1; II–2.78, 2.54; III–2.43, 2.45; IV– 3.35, 2.93.
|
||
</paragraph>
|
||
<paragraph id="8BDA36A7764CFFB992C0F90FFADCD329" blockId="17.[151,1437,1445,1994]" pageId="17" pageNumber="18">
|
||
Genitalia: Palp (
|
||
<figureCitation id="135E2A22764CFFB99380F90FFD9DD359" box="[388,527,1781,1807]" captionStart="FIGURES 35 – 40" captionStartId="17.[151,270,1248,1272]" captionTargetBox="[151,1435,194,1226]" captionTargetId="figure@17.[151,1436,194,1226]" captionTargetPageId="17" captionText="FIGURES 35 – 40. Lyrognathus achilles sp. nov., paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10 mm for 35; 2 mm for 38, 39; 1 mm for 36, 37, 40. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275736/files/figure.png" pageId="17" pageNumber="18">Figs 38–40</figureCitation>
|
||
): tibia mildly incrassate, tapering distally. Cymbium bipartite, tegulum large. Lanceolate embolus slightly tapers distally, very little flaring, proximally emerges anterior to tegulum. Relative length of palpal tibia about three times length of embolus. Longitudinal keel present, distinct (embolus not terete), along entire length of embolus, twisted through 90° prolaterally distally (
|
||
<figureCitation id="135E2A22764CFFB996EDF89FFAD0D329" box="[1257,1346,1893,1919]" captionStart="FIGURES 35 – 40" captionStartId="17.[151,270,1248,1272]" captionTargetBox="[151,1435,194,1226]" captionTargetId="figure@17.[151,1436,194,1226]" captionTargetPageId="17" captionText="FIGURES 35 – 40. Lyrognathus achilles sp. nov., paratype male: 35. Sternum, labium, maxillae and coxa, ventral view. 36. Right maxillae, retrotrolateral view showing strikers. 37. Right chelicerae, intercheliceral pegs, prolateral view. 38. Right distal palp showing bulb, prolateral view. 39. Right distal palp showing bulb, retrolateral view. 40. Right bulb and embolus, ventro-anterior view showing keeled embolus. Scale bar = 10 mm for 35; 2 mm for 38, 39; 1 mm for 36, 37, 40. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275736/files/figure.png" pageId="17" pageNumber="18">Fig. 40</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BDA36A7764CFFB992C0F871FC61D39C" blockId="17.[151,1437,1445,1994]" pageId="17" pageNumber="18">Spinnerets: PMS: length 1.23, width (medially) 0.51. PLS: section lengths; basal 1.95, medial 1.25, apical 1.85. PLS section widths (medially); basal 1.12, medial 0.75, apical 0.55.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37F652C764FFFBA92C0FF6DFD38D511" pageId="18" pageNumber="19" type="distribution">
|
||
<paragraph id="8BDA36A7764FFFBA92C0FF6DFD38D511" blockId="18.[151,1437,151,327]" pageId="18" pageNumber="19">
|
||
<emphasis id="B911EAB5764FFFBA92C0FF6DFDC7D4E7" bold="true" box="[196,597,151,177]" pageId="18" pageNumber="19">Distribution and natural history</emphasis>
|
||
(
|
||
<figureCitation id="135E2A22764FFFBA9067FF62FD78D4E4" box="[611,746,152,178]" captionStart="FIGURES 41 – 44. 41" captionStartId="18.[151,270,1327,1351]" captionTargetBox="[152,1434,384,1304]" captionTargetId="figure@18.[152,1435,384,1305]" captionTargetPageId="18" captionText="FIGURES 41 – 44. 41. Lyrognathus achilles sp. nov. Upper slopes of Gunung Nibung, Borneo, Indonesia. 42. Biotope, Gunung Nibung. 43. Biotope with arrow pointing to burrow entrance. 44. Detail of burrow entrance among foliage. Photos: 41 – 44: H. Yanto." httpUri="https://zenodo.org/record/275737/files/figure.png" pageId="18" pageNumber="19">Figs 41–44</figureCitation>
|
||
,
|
||
<figureCitation id="135E2A22764FFFBA90F1FF62FCB7D4E4" box="[757,805,152,178]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="18" pageNumber="19">114</figureCitation>
|
||
): Known only from the
|
||
<typeStatus id="54DE8805764FFFBA963EFF62FBFFD4E4" box="[1082,1133,152,178]" pageId="18" pageNumber="19">type</typeStatus>
|
||
locality, Gunung Nibung, Kotawaringin Barat, Borneo,
|
||
<collectingCountry id="F3727637764FFFBA93F7FF47FDFBD481" box="[499,617,189,215]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
|
||
. Little is known; however,
|
||
<taxonomicName id="4C654D24764FFFBA91A3FF47FBB3D480" box="[935,1057,189,214]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="achilles">
|
||
<emphasis id="B911EAB5764FFFBA91A3FF47FBB3D480" box="[935,1057,189,214]" italics="true" pageId="18" pageNumber="19">L. achilles</emphasis>
|
||
</taxonomicName>
|
||
is a fossorial spider that lives in higher shaded tropical forests away from the riparian flood plains. Burrows were found only on sloped embankments amongst leaf litter and low vegetation and were formed in clay-like soil. A mature male was found in early April (H. Yanto, pers. comm.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |