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<document id="EE2B98EA176D3CB954E247FC8A7FF55A" ID-DOI="10.5281/zenodo.275727" ID-GBIF-Dataset="1e79f840-6ced-48b5-ab20-94992c6f0a88" ID-ISSN="1175-5326" ID-Zenodo-Dep="275727" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460160971924" checkinUser="plazi" docAuthor="West, Rick C. &amp; Nunn, Steven C." docDate="2010" docId="03CC87B1764FFFB09293FA1FFE1ED5E1" docLanguage="en" docName="zt02362p043.pdf" docOrigin="Zootaxa 2362" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Lyrognathus fuscus West &amp; Nunn, 2010, sp.nov." docType="treatment" docVersion="8" lastPageNumber="25" masterDocId="FFF5FFC9765DFFA89204FFFAFF92D456" masterDocTitle="A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae" masterLastPageNumber="43" masterPageNumber="1" pageNumber="19" updateTime="1698235793967" updateUser="plazi">
<mods:mods id="A580EC3BBCFA6FBEE59C871F08E210BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C6D6747FE739433FA64DBBCFDB429DC0">
<mods:title id="0E14D83C18C9C466077CAEA9D7172725">A taxonomic revision of the tarantula spider genus Lyrognathus Pocock 1895 (Araneae, Theraphosidae), with notes on the Selenocosmiinae</mods:title>
</mods:titleInfo>
<mods:name id="099A8C02FFE9F3B2659AEACBD3488454" type="personal">
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<mods:namePart id="DA0093058C87D5A5BC1B2B08687D6505">West, Rick C.</mods:namePart>
</mods:name>
<mods:name id="D2D996E0E97C2109FA8339326320EC3F" type="personal">
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<mods:namePart id="2F8F3A95AEB28EC079778E9CDFAFF198">Nunn, Steven C.</mods:namePart>
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<mods:typeOfResource id="CB5C438E65D4640FD290A8825831C4AE">text</mods:typeOfResource>
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<mods:title id="F484E532C06380B5C43F42D15623894A">Zootaxa</mods:title>
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<mods:part id="C5CB691CA9B695ABF942C947317C7CB7">
<mods:date id="C5595100AF1E2FC14FC2BD3F924C87BC">2010</mods:date>
<mods:detail id="30003A82B0F4393907EB8C5ECD18473B" type="volume">
<mods:number id="D1AAA59BEBD4044771BF0D96B5278FD8">2362</mods:number>
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<mods:start id="AD68E5F803842B8AAD4C07E4BE735830">1</mods:start>
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<mods:classification id="8FB3EFDCDDC4C82AB85FB1503BE61E92">journal article</mods:classification>
<mods:identifier id="EC855592A5A12D95C8AEED9CBFB0EBB1" type="DOI">10.5281/zenodo.275727</mods:identifier>
<mods:identifier id="342A2C70255F2CB6B317DB61D8906279" type="GBIF-Dataset">1e79f840-6ced-48b5-ab20-94992c6f0a88</mods:identifier>
<mods:identifier id="3A39BFEE62C90014536A1D8B51E758F3" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E90CCF1C5BEA181F051A6DD1D2775B9E" type="Zenodo-Dep">275727</mods:identifier>
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<treatment id="03CC87B1764FFFB09293FA1FFE1ED5E1" ID-DOI="http://doi.org/10.5281/zenodo.6210154" ID-GBIF-Taxon="127654811" ID-Zenodo-Dep="6210154" LSID="urn:lsid:plazi:treatment:03CC87B1764FFFB09293FA1FFE1ED5E1" httpUri="http://treatment.plazi.org/id/03CC87B1764FFFB09293FA1FFE1ED5E1" lastPageId="24" lastPageNumber="25" pageId="18" pageNumber="19">
<subSubSection id="C37F652C764FFFBA9293FA1FFEF5D274" pageId="18" pageNumber="19" type="nomenclature">
<paragraph id="8BDA36A7764FFFBA9293FA1FFE71D1A9" blockId="18.[151,483,1509,1570]" box="[151,483,1509,1535]" pageId="18" pageNumber="19">
<heading id="D09281CB764FFFBA9293FA1FFE71D1A9" bold="true" box="[151,483,1509,1535]" fontSize="11" level="1" pageId="18" pageNumber="19" reason="1">
<emphasis id="B911EAB5764FFFBA9293FA1FFE71D1A9" bold="true" box="[151,483,1509,1535]" pageId="18" pageNumber="19">
<taxonomicName id="4C654D24764FFFBA9293FA1FFE14D1A9" ID-CoL="3WTQK" box="[151,390,1509,1535]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB5764FFFBA9293FA1FFE14D1A9" bold="true" box="[151,390,1509,1535]" italics="true" pageId="18" pageNumber="19">Lyrognathus fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE764FFFBA9389FA1FFE71D1A9" box="[397,483,1509,1535]" pageId="18" pageNumber="19" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA36A7764FFFBA9293F9F2FEF5D274" blockId="18.[151,483,1509,1570]" box="[151,359,1544,1570]" pageId="18" pageNumber="19">
(
<figureCitation id="135E2A22764FFFBA929BF9F2FEB4D274" box="[159,294,1544,1570]" captionStart-0="FIGURES 45 – 46" captionStart-1="FIGURES 47 – 52" captionStart-2="FIGURES 53 – 57" captionStart-3="FIGURES 58 – 62" captionStartId-0="19.[151,269,1971,1995]" captionStartId-1="20.[151,271,1644,1668]" captionStartId-2="21.[151,270,1949,1973]" captionStartId-3="22.[151,270,1138,1162]" captionTargetBox-0="[151,1436,293,1948]" captionTargetBox-1="[151,1436,194,1619]" captionTargetBox-2="[151,1435,815,1935]" captionTargetBox-3="[151,1434,194,1116]" captionTargetId-0="figure@19.[151,1436,293,1950]" captionTargetId-1="figure@20.[151,1436,194,1622]" captionTargetId-2="figure@21.[151,1436,815,1938]" captionTargetId-3="figure@22.[151,1436,194,1116]" captionTargetPageId-0="19" captionTargetPageId-1="20" captionTargetPageId-2="21" captionTargetPageId-3="22" captionText-0="FIGURES 45 – 46. Lyrognathus fuscus sp. nov. 45. Holotype female in life. 46. Male in life. Photos: 45, R. C. West; 46, H. Yanto." captionText-1="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." captionText-2="FIGURES 53 – 57. Lyrognathus fuscus sp. nov. holotype female (except paratype QM S 88051; 56 – 57): 53. Left leg IV, dorsal view. 54. Left leg IV, retrolateral view. 55. Left met, tar. IV, prolateral view. 56. Left maxillae, prolateral view. 57. Maxillary lyra, prolateral view. Scale bar = 10 mm for 53 – 55; 2 mm for 56; 1 mm for 57. Photos: S. C. Nunn." captionText-3="FIGURES 58 – 62. Lyrognathus fuscus sp. nov. paratype female QM S 88051: 58. Left chelicerae, retrolateral view. 59. Left cheliceral strikers, ectal view. 60. Basomedial cheliceral spines, retrolateral view. 61. Left chelicerae, prolateral view. 62. Left intercheliceral pegs, prolateral view. Scale bar = 5 mm for 58, 61; 2 mm for 59; 1 mm for 60, 62. Photos: S. C. Nunn." httpUri-0="https://zenodo.org/record/275738/files/figure.png" httpUri-1="https://zenodo.org/record/275739/files/figure.png" httpUri-2="https://zenodo.org/record/275740/files/figure.png" httpUri-3="https://zenodo.org/record/275741/files/figure.png" pageId="18" pageNumber="19">Figs 45–62</figureCitation>
,
<figureCitation id="135E2A22764FFFBA9336F9F2FECCD274" box="[306,350,1544,1570]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="18" pageNumber="19">114</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37F652C764FFFBA9293F9A8FEF6D2E1" pageId="18" pageNumber="19" type="description">
<paragraph id="8BDA36A7764FFFBA9293F9A8FEF6D2E1" blockId="18.[151,1437,1618,2018]" pageId="18" pageNumber="19">
<emphasis id="B911EAB5764FFFBA9293F9A8FF7BD23A" bold="true" box="[151,233,1618,1644]" pageId="18" pageNumber="19">
<typeStatus id="54DE8805764FFFBA9293F9A8FF72D23A" box="[151,224,1618,1644]" pageId="18" pageNumber="19">Types</typeStatus>
:
</emphasis>
female
<typeStatus id="54DE8805764FFFBA934AF9A8FE28D23A" box="[334,442,1618,1644]" pageId="18" pageNumber="19" type="holotype">holotype</typeStatus>
,
<specimenCount id="9D63FD2E764FFFBA93CCF9A8FDA3D23A" box="[456,561,1618,1644]" pageId="18" pageNumber="19" type="female">3 female</specimenCount>
<typeStatus id="54DE8805764FFFBA903DF9A8FD3ED23A" box="[569,684,1618,1644]" pageId="18" pageNumber="19" type="paratype">paratypes</typeStatus>
<accessionNumber id="9436AB44764FFFBA90B1F9A8FC86D23A" box="[693,788,1618,1644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88054" pageId="18" pageNumber="19" type="EnaNcbi">S88054</accessionNumber>
,
<accessionNumber id="9436AB44764FFFBA9125F9A8FC13D23A" box="[801,897,1618,1644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88051" pageId="18" pageNumber="19" type="EnaNcbi">S88051</accessionNumber>
,
<accessionNumber id="9436AB44764FFFBA9189F9A8FC7BD23A" box="[909,1001,1618,1644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88052" pageId="18" pageNumber="19" type="EnaNcbi">S88052</accessionNumber>
and
<accessionNumber id="9436AB44764FFFBA9621F9A8FB17D23A" box="[1061,1157,1618,1644]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88053" pageId="18" pageNumber="19" type="EnaNcbi">S88053</accessionNumber>
, respectively, Simpang Kubur (
<geoCoordinate id="EE515060764FFFBA92F5F982FEDAD2C4" box="[241,328,1656,1682]" direction="south" orientation="latitude" pageId="18" pageNumber="19" precision="925" value="-2.4833333">2°29' S</geoCoordinate>
,
<geoCoordinate id="EE515060764FFFBA9350F982FE5AD2C4" box="[340,456,1656,1682]" direction="east" orientation="longitude" pageId="18" pageNumber="19" precision="925" value="111.416664">111°25' E</geoCoordinate>
), Katawaringin, Kotawaringin Barat, Borneo,
<collectingCountry id="F3727637764FFFBA91E7F982FBCAD2C4" box="[995,1112,1656,1682]" name="Indonesia" pageId="18" pageNumber="19">Indonesia</collectingCountry>
, col. H. Yanto,
<date id="FFDB1067764FFFBA970FF982FA0AD2C4" box="[1291,1432,1656,1682]" pageId="18" pageNumber="19" value="2009-04-02">2 Apr. 2009</date>
, QM.—examined.
</paragraph>
</subSubSection>
<subSubSection id="C37F652C764FFFBA92C2F938FB8DD371" pageId="18" pageNumber="19" type="etymology">
<paragraph id="8BDA36A7764FFFBA92C2F938FD80D354" blockId="18.[151,1437,1618,2018]" pageId="18" pageNumber="19">
<emphasis id="B911EAB5764FFFBA92C2F938FEC5D28A" bold="true" box="[198,343,1730,1756]" pageId="18" pageNumber="19">Etymology:</emphasis>
From Latin meaning dark brown/grey. The genus name is masculine, thus
<taxonomicName id="4C654D24764FFFBA9707F939FADED28A" box="[1283,1356,1731,1756]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B911EAB5764FFFBA9707F939FADED28A" box="[1283,1356,1731,1756]" italics="true" pageId="18" pageNumber="19">fuscus</emphasis>
</taxonomicName>
is the masculine form of the adjective.
</paragraph>
<paragraph id="8BDA36A7764FFFBA92C2F8F7FB8DD371" blockId="18.[151,1437,1618,2018]" box="[198,1055,1805,1831]" pageId="18" pageNumber="19">
<emphasis id="B911EAB5764FFFBA92C2F8F7FE1DD371" bold="true" box="[198,399,1805,1831]" pageId="18" pageNumber="19">Common name:</emphasis>
Hariyanto’s tarantula, after the collector of this species.
</paragraph>
</subSubSection>
<subSubSection id="C37F652C764FFFBB92C2F8C8FEF5D481" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19" type="diagnosis">
<paragraph id="8BDA36A7764FFFBB92C2F8C8FEF5D481" blockId="18.[151,1437,1618,2018]" lastBlockId="19.[151,1436,152,252]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B911EAB5764FFFBA92C2F8C8FEDAD31A" bold="true" box="[198,328,1842,1868]" pageId="18" pageNumber="19">Diagnosis:</emphasis>
Differs from both
<taxonomicName id="4C654D24764FFFBA9028F8C9FD3ED31A" box="[556,684,1843,1868]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="crotalus">
<emphasis id="B911EAB5764FFFBA9028F8C9FD3ED31A" box="[556,684,1843,1868]" italics="true" pageId="18" pageNumber="19">L. crotalus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C654D24764FFFBA90EDF8C9FCF6D31A" box="[745,868,1843,1868]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="saltator">
<emphasis id="B911EAB5764FFFBA90EDF8C9FCF6D31A" box="[745,868,1843,1868]" italics="true" pageId="18" pageNumber="19">L. saltator</emphasis>
</taxonomicName>
in having intercheliceral pegs (not spines), and retrolateral basomedial cheliceral spines. Differs from
<taxonomicName id="4C654D24764FFFBA911DF8A2FC0ED327" box="[793,924,1880,1905]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="robustus">
<emphasis id="B911EAB5764FFFBA911DF8A2FC0ED327" box="[793,924,1880,1905]" italics="true" pageId="18" pageNumber="19">L. robustus</emphasis>
</taxonomicName>
in having undivided scopula for full length of metatarsi IV, swollen distal lateral lobes of spermathecae but no third claw on tarsi IV. Differs from
<taxonomicName id="4C654D24764FFFBA9780F887FE91D3EA" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="lessunda" status="sp. nov.">
<emphasis id="B911EAB5764FFFBA9780F887FE91D3EA" italics="true" pageId="18" pageNumber="19">L. lessunda</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE764FFFBA9308F858FEFDD3EA" box="[268,367,1954,1980]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B911EAB5764FFFBA9308F858FEFDD3EA" bold="true" box="[268,367,1954,1980]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
in the absence of “mushroom cap” medial lobes of the spermathecae. Differs from
<taxonomicName id="4C654D24764FFFBA9780F859FF67D3B7" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
<emphasis id="B911EAB5764FFFBA9780F859FF67D3B7" italics="true" pageId="18" pageNumber="19">L. achilles</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE764FFFBA92F9F83DFECFD3B7" box="[253,349,1991,2017]" pageId="18" pageNumber="19" rank="species">
<emphasis id="B911EAB5764FFFBA92F9F83DFECFD3B7" bold="true" box="[253,349,1991,2017]" pageId="18" pageNumber="19">sp. nov.</emphasis>
</taxonomicNameLabel>
in less pronounced maxillary lobe, lack of distal cream/white leg segment bands, smaller size, color and proximal-distal gradation of retrolateral tibiae IV penicillate brush (uniform for length in
<taxonomicName id="4C654D24764EFFBB9780FF62FF60D480" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="achilles" status="sp. nov.">
<emphasis id="B911EAB5764EFFBB9780FF62FF60D480" italics="true" pageId="19" pageNumber="20">L. achilles</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE764EFFBB92FDFF47FEC5D481" box="[249,343,189,215]" pageId="19" pageNumber="20" rank="species">
<emphasis id="B911EAB5764EFFBB92FDFF47FEC5D481" bold="true" box="[249,343,189,215]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F652C764EFFB092C0FF18FEEED4AA" lastPageId="24" lastPageNumber="25" pageId="19" pageNumber="20" type="description">
<paragraph id="8BDA36A7764EFFBB92C0FF18FC2DD4AA" blockId="19.[151,1436,152,252]" box="[196,959,226,252]" pageId="19" pageNumber="20">
<emphasis id="B911EAB5764EFFBB92C0FF18FECCD4AA" bold="true" box="[196,350,226,252]" pageId="19" pageNumber="20">Description:</emphasis>
Female
<typeStatus id="54DE8805764EFFBB93C7FF18FDB8D4AA" box="[451,554,226,252]" pageId="19" pageNumber="20" type="holotype">holotype</typeStatus>
(
<figureCitation id="135E2A22764EFFBB903EFF18FD02D4AA" box="[570,656,226,252]" captionStart="FIGURES 45 – 46" captionStartId="19.[151,269,1971,1995]" captionTargetBox="[151,1436,293,1948]" captionTargetId="figure@19.[151,1436,293,1950]" captionTargetPageId="19" captionText="FIGURES 45 – 46. Lyrognathus fuscus sp. nov. 45. Holotype female in life. 46. Male in life. Photos: 45, R. C. West; 46, H. Yanto." httpUri="https://zenodo.org/record/275738/files/figure.png" pageId="19" pageNumber="20">Fig. 45</figureCitation>
) with body length: 45.87.
</paragraph>
<caption id="DF1A662F764EFFBB9293F849FF6FD3BC" httpUri="https://zenodo.org/record/275738/files/figure.png" pageId="19" pageNumber="20" targetBox="[151,1436,293,1948]" targetPageId="19">
<paragraph id="8BDA36A7764EFFBB9293F849FF6FD3BC" blockId="19.[151,1436,1971,2026]" pageId="19" pageNumber="20">
<emphasis id="B911EAB5764EFFBB9293F849FECCD39D" bold="true" box="[151,350,1971,1995]" pageId="19" pageNumber="20">FIGURES 45–46.</emphasis>
<taxonomicName id="4C654D24764EFFBB9360F849FDA6D39C" box="[356,564,1971,1994]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="19" pageNumber="20" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB5764EFFBB9360F849FDA6D39C" box="[356,564,1971,1994]" italics="true" pageId="19" pageNumber="20">Lyrognathus fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE764EFFBB903EF849FD02D39D" box="[570,656,1971,1995]" pageId="19" pageNumber="20" rank="species">
<emphasis id="B911EAB5764EFFBB903EF849FD02D39D" bold="true" box="[570,656,1971,1995]" pageId="19" pageNumber="20">sp. nov.</emphasis>
</taxonomicNameLabel>
45. Holotype female in life. 46. Male in life. Photos: 45, R. C. West; 46, H. Yanto.
</paragraph>
</caption>
<caption id="DF1A662F7649FFBC9293F996FA8BD295" httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="20" pageNumber="21" targetBox="[151,1436,194,1619]" targetPageId="20">
<paragraph id="8BDA36A77649FFBC9293F996FA8BD295" blockId="20.[151,1437,1644,1731]" pageId="20" pageNumber="21">
<emphasis id="B911EAB57649FFBC9293F996FEF2D2D2" bold="true" box="[151,352,1644,1668]" pageId="20" pageNumber="21">FIGURES 47–52.</emphasis>
<taxonomicName id="4C654D247649FFBC9363F996FDABD2D5" box="[359,569,1644,1667]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="20" pageNumber="21" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB57649FFBC9363F996FDABD2D5" box="[359,569,1644,1667]" italics="true" pageId="20" pageNumber="21">Lyrognathus fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7649FFBC9045F996FD0AD2D2" box="[577,664,1644,1668]" pageId="20" pageNumber="21" rank="species">
<emphasis id="B911EAB57649FFBC9045F996FD0AD2D2" bold="true" box="[577,664,1644,1668]" pageId="20" pageNumber="21">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10mm for 47, 49, 52; 2mm for 48, 50; 1mm for 51. Photos: S. C. Nunn.
</paragraph>
</caption>
<paragraph id="8BDA36A77649FFBC92C2F908FBC8D35A" blockId="20.[151,1437,1778,2028]" box="[198,1114,1778,1804]" pageId="20" pageNumber="21">
Color (in life,
<figureCitation id="135E2A227649FFBC936AF908FE57D35A" box="[366,453,1778,1804]" captionStart="FIGURES 45 – 46" captionStartId="19.[151,269,1971,1995]" captionTargetBox="[151,1436,293,1948]" captionTargetId="figure@19.[151,1436,293,1950]" captionTargetPageId="19" captionText="FIGURES 45 – 46. Lyrognathus fuscus sp. nov. 45. Holotype female in life. 46. Male in life. Photos: 45, R. C. West; 46, H. Yanto." httpUri="https://zenodo.org/record/275738/files/figure.png" pageId="20" pageNumber="21">Fig. 45</figureCitation>
): Entirely dark brown/grey, no distal leg segment bands.
</paragraph>
<paragraph id="8BDA36A77649FFBD92C0F8EDFCD0D481" blockId="20.[151,1437,1778,2028]" lastBlockId="21.[151,1437,152,775]" lastPageId="21" lastPageNumber="22" pageId="20" pageNumber="21">
Carapace (
<figureCitation id="135E2A227649FFBC934EF8EDFE39D367" box="[330,427,1815,1841]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="20" pageNumber="21">Fig. 47</figureCitation>
): length 16.96, width 12.77 (width across anterior edge 8.85). Fovea 2.20 wide, procurved, deep, smaller in width than OT. Distance from anterior carapace to fovea, 12.51; with 4 discernible hair
<typeStatus id="54DE88057649FFBC92C8F898FE98D32A" box="[204,266,1890,1916]" pageId="20" pageNumber="21">types</typeStatus>
(C1 to C4):
<typeStatus id="54DE88057649FFBC93A5F898FE41D32A" box="[417,467,1890,1916]" pageId="20" pageNumber="21">type</typeStatus>
C1, long needle-form pallids border carapace entirely;
<typeStatus id="54DE88057649FFBC9666F898FB06D32A" box="[1122,1172,1890,1916]" pageId="20" pageNumber="21">type</typeStatus>
C2, short black, blunt tipped, entirely over carapace;
<typeStatus id="54DE88057649FFBC900AF87DFDD0D3F7" box="[526,578,1927,1953]" pageId="20" pageNumber="21">type</typeStatus>
C3, short thin, wavy pallids, form basis of mat;
<typeStatus id="54DE88057649FFBC9696F87DFB54D3F7" box="[1170,1222,1927,1953]" pageId="20" pageNumber="21">type</typeStatus>
C4, emerge from bothrial-like collars, long, distally spiniform, pallid, in uniform arrangement: (&gt;5) along anterior edge of each radial groove, (&gt;4) along anterior edge of each thoracic groove, (&gt;5) along anterior edge of sulcal groove, (&gt;6) along anterior edge of each caput groove, (&gt;4) along medial caput ridge (between OT/fovea), (&gt;3) located behind each side of OT, nearest median caput row.
</paragraph>
<paragraph id="8BDA36A77648FFBD92C0FF18FDFAD654" blockId="21.[151,1437,152,775]" pageId="21" pageNumber="22">
Eyes (
<figureCitation id="135E2A227648FFBD9314FF18FEFFD4AA" box="[272,365,226,252]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="21" pageNumber="22">Fig. 48</figureCitation>
): ocular tubercle; length 1.95, width 2.91. OT highest posteriorly behind AME, sloping gradually anteriorly. Anterior row slightly procurved. Posterior row recurved. Eyes in lengths/widths: AME: 0.67/0.51, ALE: 0.71/0.27, PLE: 0.41/0.25, PME: 0.40/0.28. Inter-ocular distances: AME–AME: 0.28, AME– ALE: 0.24, AME–PME: 0.15, ALE–ALE: 1.79, ALE–PME: 0.41, PME–PME: 1.43, PME–PLE: 0.19, PLE– PLE: 2.0, ALE–PLE: 0.38. OT hairs:&gt;15 (C2) directly posterior to AME’s, two between PME and center of OT (behind AME). 16 (C2) along median anterior OT surface. OT entirely covered in small, weak, pallid setae (C3), directionally point to center of OT, posteriorly–anteriorly. Lateral small pallids very sparse with no uniform direction/random arrangement.
</paragraph>
<paragraph id="8BDA36A77648FFBD92C0FDF7FB5AD751" blockId="21.[151,1437,152,775]" pageId="21" pageNumber="22">
Chelicerae (right,
<figureCitation id="135E2A227648FFBD939CFDF7FDB3D671" box="[408,545,525,551]" captionStart="FIGURES 58 – 62" captionStartId="22.[151,270,1138,1162]" captionTargetBox="[151,1434,194,1116]" captionTargetId="figure@22.[151,1436,194,1116]" captionTargetPageId="22" captionText="FIGURES 58 – 62. Lyrognathus fuscus sp. nov. paratype female QM S 88051: 58. Left chelicerae, retrolateral view. 59. Left cheliceral strikers, ectal view. 60. Basomedial cheliceral spines, retrolateral view. 61. Left chelicerae, prolateral view. 62. Left intercheliceral pegs, prolateral view. Scale bar = 5 mm for 58, 61; 2 mm for 59; 1 mm for 60, 62. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275741/files/figure.png" pageId="21" pageNumber="22">Figs 58–62</figureCitation>
): length 8.90, width at base of lobe 4.15 (bases widest point, dorsally). Ectal lyrate region a series of strikers (&gt;75), 8 horizontal rows (
<figureCitation id="135E2A227648FFBD914BFDC8FC3BD61A" box="[847,937,562,588]" captionStart="FIGURES 58 – 62" captionStartId="22.[151,270,1138,1162]" captionTargetBox="[151,1434,194,1116]" captionTargetId="figure@22.[151,1436,194,1116]" captionTargetPageId="22" captionText="FIGURES 58 – 62. Lyrognathus fuscus sp. nov. paratype female QM S 88051: 58. Left chelicerae, retrolateral view. 59. Left cheliceral strikers, ectal view. 60. Basomedial cheliceral spines, retrolateral view. 61. Left chelicerae, prolateral view. 62. Left intercheliceral pegs, prolateral view. Scale bar = 5 mm for 58, 61; 2 mm for 59; 1 mm for 60, 62. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275741/files/figure.png" pageId="21" pageNumber="22">Fig. 59</figureCitation>
). Strongest and longest strikers on lowest rows. Each striker very stout, blade or scimitar-shaped with long filiform ends. Teeth, 13,&gt;40 basomesal teeth. Intercheliceral pegs (&gt;11 larger,&gt;18 smaller) in tight cluster on basodorsal surface, extremely stout, 1/2 as wide as long (
<figureCitation id="135E2A227648FFBD935BFD58FE2BD6EA" box="[351,441,674,700]" captionStart="FIGURES 58 – 62" captionStartId="22.[151,270,1138,1162]" captionTargetBox="[151,1434,194,1116]" captionTargetId="figure@22.[151,1436,194,1116]" captionTargetPageId="22" captionText="FIGURES 58 – 62. Lyrognathus fuscus sp. nov. paratype female QM S 88051: 58. Left chelicerae, retrolateral view. 59. Left cheliceral strikers, ectal view. 60. Basomedial cheliceral spines, retrolateral view. 61. Left chelicerae, prolateral view. 62. Left intercheliceral pegs, prolateral view. Scale bar = 5 mm for 58, 61; 2 mm for 59; 1 mm for 60, 62. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275741/files/figure.png" pageId="21" pageNumber="22">Fig. 62</figureCitation>
). Retrolaterally and basomedially, small cluster of short strong spiniform setae (&gt;7) point distad (
<figureCitation id="135E2A227648FFBD9330FD32FE1DD6B4" box="[308,399,712,738]" captionStart="FIGURES 58 – 62" captionStartId="22.[151,270,1138,1162]" captionTargetBox="[151,1434,194,1116]" captionTargetId="figure@22.[151,1436,194,1116]" captionTargetPageId="22" captionText="FIGURES 58 – 62. Lyrognathus fuscus sp. nov. paratype female QM S 88051: 58. Left chelicerae, retrolateral view. 59. Left cheliceral strikers, ectal view. 60. Basomedial cheliceral spines, retrolateral view. 61. Left chelicerae, prolateral view. 62. Left intercheliceral pegs, prolateral view. Scale bar = 5 mm for 58, 61; 2 mm for 59; 1 mm for 60, 62. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275741/files/figure.png" pageId="21" pageNumber="22">Fig. 60</figureCitation>
). Surrounding spines a patch of fine setae, dark, of medium length, blends into typical thicker pallid cheliceral setae. Setae filiform and seem to emerge from hemispherical bases.
</paragraph>
<caption id="DF1A662F7648FFBD9293F867FA84D3A2" httpUri="https://zenodo.org/record/275740/files/figure.png" pageId="21" pageNumber="22" targetBox="[151,1435,815,1935]" targetPageId="21">
<paragraph id="8BDA36A77648FFBD9293F867FA84D3A2" blockId="21.[151,1436,1949,2036]" pageId="21" pageNumber="22">
<emphasis id="B911EAB57648FFBD9293F867FECDD3E3" bold="true" box="[151,351,1949,1973]" pageId="21" pageNumber="22">FIGURES 53–57.</emphasis>
<taxonomicName id="4C654D247648FFBD9362F864FDA4D3E3" box="[358,566,1950,1973]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB57648FFBD9362F864FDA4D3E3" box="[358,566,1950,1973]" italics="true" pageId="21" pageNumber="22">Lyrognathus fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE7648FFBD9039F867FD06D3E3" box="[573,660,1949,1973]" pageId="21" pageNumber="22" rank="species">
<emphasis id="B911EAB57648FFBD9039F867FD06D3E3" bold="true" box="[573,660,1949,1973]" pageId="21" pageNumber="22">sp. nov.</emphasis>
</taxonomicNameLabel>
holotype female (except paratype QM
<accessionNumber id="9436AB447648FFBD9638F867FB03D3E2" box="[1084,1169,1949,1972]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88051" pageId="21" pageNumber="22" type="EnaNcbi">S88051</accessionNumber>
; 56–57): 53. Left leg IV, dorsal view. 54. Left leg IV, retrolateral view. 55. Left met, tar. IV, prolateral view. 56. Left maxillae, prolateral view. 57. Maxillary
<taxonomicName id="4C654D247648FFBD9303F827FEA1D3A2" box="[263,307,2013,2036]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="21" pageNumber="22" phylum="Arthropoda" rank="species" species="lyra">lyra</taxonomicName>
, prolateral view. Scale bar = 10mm for 53–55; 2mm for 56; 1mm for 57. Photos: S. C. Nunn.
</paragraph>
</caption>
<caption id="DF1A662F764BFFBE9293FB88FF6BD0BF" httpUri="https://zenodo.org/record/275741/files/figure.png" pageId="22" pageNumber="23" targetBox="[151,1434,194,1116]" targetPageId="22">
<paragraph id="8BDA36A7764BFFBE9293FB88FF6BD0BF" blockId="22.[151,1436,1138,1257]" pageId="22" pageNumber="23">
<emphasis id="B911EAB5764BFFBE9293FB88FECDD0DC" bold="true" box="[151,351,1138,1162]" pageId="22" pageNumber="23">FIGURES 58–62.</emphasis>
<taxonomicName id="4C654D24764BFFBE9362FB88FDA5D0DF" box="[358,567,1138,1161]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="fuscus" status="sp. nov.">
<emphasis id="B911EAB5764BFFBE9362FB88FDA5D0DF" box="[358,567,1138,1161]" italics="true" pageId="22" pageNumber="23">Lyrognathus fuscus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257CE764BFFBE903AFB88FD04D0DC" box="[574,662,1138,1162]" pageId="22" pageNumber="23" rank="species">
<emphasis id="B911EAB5764BFFBE903AFB88FD04D0DC" bold="true" box="[574,662,1138,1162]" pageId="22" pageNumber="23">sp. nov.</emphasis>
</taxonomicNameLabel>
paratype female QM
<accessionNumber id="9436AB44764BFFBE9187FB88FC4BD0DF" box="[899,985,1138,1161]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S88051" pageId="22" pageNumber="23" type="EnaNcbi">S88051</accessionNumber>
: 58. Left chelicerae, retrolateral view. 59. Left cheliceral strikers, ectal view. 60. Basomedial cheliceral spines, retrolateral view. 61. Left chelicerae, prolateral view. 62. Left intercheliceral pegs, prolateral view. Scale bar = 5mm for 58, 61; 2mm for 59; 1mm for 60, 62. Photos: S. C. Nunn.
</paragraph>
</caption>
<paragraph id="8BDA36A7764BFFBE92C0FAEDFD7DD2F1" blockId="22.[151,1437,1303,2001]" pageId="22" pageNumber="23">
Maxillae (
<figureCitation id="135E2A22764BFFBE9339FAEDFE0BD167" box="[317,409,1303,1329]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="22" pageNumber="23">Figs 49</figureCitation>
,
<figureCitation id="135E2A22764BFFBE93A2FAEDFE5AD167" box="[422,456,1303,1329]" captionStart="FIGURES 53 – 57" captionStartId="21.[151,270,1949,1973]" captionTargetBox="[151,1435,815,1935]" captionTargetId="figure@21.[151,1436,815,1938]" captionTargetPageId="21" captionText="FIGURES 53 – 57. Lyrognathus fuscus sp. nov. holotype female (except paratype QM S 88051; 56 – 57): 53. Left leg IV, dorsal view. 54. Left leg IV, retrolateral view. 55. Left met, tar. IV, prolateral view. 56. Left maxillae, prolateral view. 57. Maxillary lyra, prolateral view. Scale bar = 10 mm for 53 – 55; 2 mm for 56; 1 mm for 57. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275740/files/figure.png" pageId="22" pageNumber="23">56</figureCitation>
): prolaterally plano-convex, anterior lobe well pronounced, many cuspules (
<emphasis id="B911EAB5764BFFBE9756FAE2FAF4D167" box="[1362,1382,1304,1329]" italics="true" pageId="22" pageNumber="23">&gt;</emphasis>
105) on inner basoventral surface.
<taxonomicName id="4C654D24764BFFBE93F3FAC7FDBFD101" box="[503,557,1341,1367]" class="Arachnida" family="Theraphosidae" genus="Selenocosmia" kingdom="Animalia" order="Araneae" pageId="22" pageNumber="42" phylum="Arthropoda" rank="species" species="lyra">Lyra</taxonomicName>
(
<figureCitation id="135E2A22764BFFBE9039FAC7FD5BD101" box="[573,713,1341,1367]" captionStart="FIGURES 53 – 57" captionStartId="21.[151,270,1949,1973]" captionTargetBox="[151,1435,815,1935]" captionTargetId="figure@21.[151,1436,815,1938]" captionTargetPageId="21" captionText="FIGURES 53 – 57. Lyrognathus fuscus sp. nov. holotype female (except paratype QM S 88051; 56 – 57): 53. Left leg IV, dorsal view. 54. Left leg IV, retrolateral view. 55. Left met, tar. IV, prolateral view. 56. Left maxillae, prolateral view. 57. Maxillary lyra, prolateral view. Scale bar = 10 mm for 53 – 55; 2 mm for 56; 1 mm for 57. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275740/files/figure.png" pageId="22" pageNumber="23">Figs 56, 57</figureCitation>
): large number of bacilliform rods (&gt;180) form dense, ovoid patch on prolateral maxillary surface (3.23 long, 1.72 high), lowest row with up to&gt;18 bacillae, longest rods in center of lowest rows. Rods distally paddle-shaped (paddles up to 0.19 long) with short to medium length shafts (total length 0.69), largest rods with no distal blades. At widest point, lyrate patch 10 rows deep, smallest rods dorsally. Posterior end of patch truncate, anterior end not pointed distad/widely rounded. Immediately above maxillary suture&gt;45 elongated spines on anterior margin in disordered rows. Labium (
<figureCitation id="135E2A22764BFFBE92A4F9E7FF6FD261" box="[160,253,1565,1591]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="22" pageNumber="23">Fig. 50</figureCitation>
): length 2.35, width 3.0. Many small cuspules (&gt;540) along anterior 1/4 surface. Labium with 2 discernible hair
<typeStatus id="54DE8805764BFFBE9350F9B8FE00D20A" box="[340,402,1602,1628]" pageId="22" pageNumber="23">types</typeStatus>
(L1, L2);
<typeStatus id="54DE8805764BFFBE9008F9B8FDACD20A" box="[524,574,1602,1628]" pageId="22" pageNumber="23">type</typeStatus>
L1: long, reasonably dark spiniform, curve distally toward anterior, most concentrated along lateral borders. Setae longest along anterior edge (&gt;35);
<typeStatus id="54DE8805764BFFBE9651F99DFB19D2D7" box="[1109,1163,1639,1665]" pageId="22" pageNumber="23">type</typeStatus>
L2: 1/2 length of L1, interspersed with L1 but darker, more needle-form.
</paragraph>
<paragraph id="8BDA36A7764BFFBE92C0F948FA0FD387" blockId="22.[151,1437,1303,2001]" pageId="22" pageNumber="23">
Sternum (
<figureCitation id="135E2A22764BFFBE9333F948FE02D29A" box="[311,400,1714,1740]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="22" pageNumber="23">Fig. 49</figureCitation>
): length 7.65, width 6.68, saddle-form; with 5 discernible hair
<typeStatus id="54DE8805764BFFBE9674F948FB3CD29A" box="[1136,1198,1714,1740]" pageId="22" pageNumber="23">types</typeStatus>
(S1 to S5):
<typeStatus id="54DE8805764BFFBE973FF948FAFFD29A" box="[1339,1389,1714,1740]" pageId="22" pageNumber="23">type</typeStatus>
S1, longest, thick blunt tipped, darker basally, become pallid distally, entire but sparse;
<typeStatus id="54DE8805764BFFBE96C5F92DFB65D2A7" box="[1217,1271,1751,1777]" pageId="22" pageNumber="23">type</typeStatus>
S2, elongate spiniforms, entirely dark, on borders, most concentrated posteriorly;
<typeStatus id="54DE8805764BFFBE91EEF907FB8DD341" box="[1002,1055,1789,1815]" pageId="22" pageNumber="23">type</typeStatus>
S3, similar morphology to S1, although only 1/4–1/2 length in comparison, over entire surface, sparse;
<typeStatus id="54DE8805764BFFBE91E7F8D8FB84D36A" box="[995,1046,1826,1852]" pageId="22" pageNumber="23">type</typeStatus>
S4, short, thin pallids, form basis of mat;
<typeStatus id="54DE8805764BFFBE92F9F8BDFEA6D337" box="[253,308,1863,1889]" pageId="22" pageNumber="23">type</typeStatus>
S5, short, black blunt tipped, entirely cover sternum in sparse disordered arrangement. Posteriorly between left and right coxa IV, border highly acuminate, lateral points slightly acuminate. Sigilla: 3 pairs, posterior, large; right with length 1.31, width 0.65, left length 1.0, width 0.6. Ovular, 1.65 apart, 1.35– 1.38 from margin. Median pair, 1/2 size of posterior, similar shape, 1.0–0.6 of their length from margin.
</paragraph>
<paragraph id="8BDA36A7764AFFBF9293FF62FE54D481" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">Anterior pair 1/3 size of posterior, somewhat obscured, border sternal margin. Labio-sternal sigilla large, only slightly smaller than PSS.</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0FF18FA0ED5E1" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Legs: formula (length); IV, I, II, III: (width); IV, III, I, II. Leg
<collectingCountry id="F3727637764AFFBF9196FF18FC2AD4AA" box="[914,952,226,252]" name="Russia" pageId="23" pageNumber="24">RF</collectingCountry>
~86.72. Leg lengths (fem., pat., tib., met., tar., total): palp: 9.32, 5.73, 6.13, 0.00, 5.10, 26.28. I: 12.48, 7.92, 9.26, 7.68, 4.99, 42.33. II: 10.78, 6.93, 7.21, 7.31, 4.71, 36.94. III: 9.32, 5.93, 5.5, 7.22, 4.23, 32.2. IV: 13.73, 7.39, 10.43, 11.85, 5.41, 48.81. Leg widths (fem., pat., tib., met., tar., total.): palp: 2.36, 2.61, 2.17, 0.00, 2.26, 9.4. I: 3.08, 3.11, 2.99, 2.13, 2.05, 13.36. II: 3.06, 3.12, 2.57, 1.95, 1.89, 12.59. III: 3.8, 3.32, 2.91, 2.10, 1.96, 14.09. IV: 3.41, 3.26, 3.44, 2.35, 2.01, 14.47. Tib. IV with strong retrolateral penicillate setal fringe, proximally weak, gradually increases distally (
<figureCitation id="135E2A22764AFFBF9733FE67FA1CD5E1" box="[1335,1422,413,439]" captionStart="FIGURES 53 – 57" captionStartId="21.[151,270,1949,1973]" captionTargetBox="[151,1435,815,1935]" captionTargetId="figure@21.[151,1436,815,1938]" captionTargetPageId="21" captionText="FIGURES 53 – 57. Lyrognathus fuscus sp. nov. holotype female (except paratype QM S 88051; 56 – 57): 53. Left leg IV, dorsal view. 54. Left leg IV, retrolateral view. 55. Left met, tar. IV, prolateral view. 56. Left maxillae, prolateral view. 57. Maxillary lyra, prolateral view. Scale bar = 10 mm for 53 – 55; 2 mm for 56; 1 mm for 57. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275740/files/figure.png" pageId="23" pageNumber="24">Fig. 53</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A7764AFFBF92C2FE38FC1BD654" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">Scopula: Met., tar. I–III undivided, met. IV undivided, tar. IV partially divided proximally (by one row of sparse setae, nonexistent in distal 1/2). Met. I–IV for full length.</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0FDF7FD9BD704" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">Coxae: some small black thorns prolatero-dorsally, no thorns retrolaterally on I–IV. Coxae easily seen dorsally. Cox. I longest, about 1.3 times longer than II. IV widest, as long as III, basally rectangular with rounded corners. Coxae with small ventral thorns prolaterally on I–IV. I–III ventrally with many long thick blunt setae proximally, pallid. No short black setae. IV with mixture of long thick blunt setae entirely, pallid intermixed with shorter thin pallid setae. Ventral surface of coxae I–IV gently sloping anteriorly. Retrolateral setation: I–III with median narrow light brush, IV entirely covered in short thin pallid setae. I–IV retrolaterally lack ventral ledge. Ventral measurements for coxae: palp–length 6.75, width 4.12; I–7.23, 3.3; II–5.68, 2.83; III–4.44, 3.43; IV–6.3, 3.95. Trochantera: palp–length 2.65, width 1.85; I–3.05, 3.41; II–2.65, 2.78; III–2.45, 2.8; IV–3.1, 3.8.</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0FCA7FB8DD0BA" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Trichobothria: All tarsi with basal filiform field slightly wider than clavate field, merges evenly. Clavates on tar. I in distal 1/2 (very dark), long filiforms only in basal 2/3, shorter filiforms intermixed with clavates distally. Clavate extent on II–IV
<emphasis id="B911EAB5764AFFBF9010FC52FDA3D797" box="[532,561,936,961]" italics="true" pageId="23" pageNumber="24">cf.</emphasis>
I, in distal 1/2. Shorter filiforms for length. Short epitrichobothrial field on I shorter than clavates, increase in length proximally. Tarsal organ evident on legs I–III under stereo microscope (up to
<date id="FFDB1067764AFFBF9377FC08FE31D05A" box="[371,419,1010,1036]" pageId="23" pageNumber="24" value="1964-10">x64</date>
), organ on tar.
<date id="FFDB1067764AFFBF9050FC08FDFDD05A" box="[596,623,1010,1036]" pageId="23" pageNumber="24">I 1</date>
/2 size
<emphasis id="B911EAB5764AFFBF90CCFC08FD77D05D" box="[712,741,1010,1035]" italics="true" pageId="23" pageNumber="24">cf.</emphasis>
II and III, low, wide, domed without noticeable ridges. On tar. I, entire field 3.3 long, 0.98 wide distally, to 1.3 wide proximally. On tar. IV, field 3.89 long, 0.85 wide distally, increase to 1.35 proximally. Metatarsi: trichobothrial field not evident laterally or proximally. Tibia: on I prolaterally and proximally, short, possibly clavate trichobothria noted (&gt;8 per
<quantity id="4C9D9B42764AFFBF9683FB98FB7BD02A" box="[1159,1257,1122,1148]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="23" pageNumber="24" unit="mm" value="0.25">0.25mm</quantity>
squared) in Lshaped patch 0.25 long, 1.25 wide, clearly pallid:&gt;6 filiforms in semicircular pattern proximally to rest of patch; on IV prolaterally and proximally, short dark clavates (&gt;7 per
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squared) in patch 1.66 long, 0.22 wide distinct:&gt;7 filiforms in semicircular pattern proximally to rest of patch.
</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0FB02FE1DD161" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">Spines: met. I with 1 DV, met. II with 1 DV, met. III with 2 DV, 1DPV, 1 DRV, 1 DD, met. IV with 2 DV, 1DPV, 1 DRV, 1 DD.</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0FAB8FB8CD10A" blockId="23.[151,1437,152,2007]" box="[196,1054,1346,1372]" pageId="23" pageNumber="24">Claws: unarmed claws on all legs and palps. Third claw absent on leg IV.</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0FA92FAB1D2FA" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Abdomen (
<figureCitation id="135E2A22764AFFBF934FFA92FE3AD1D4" box="[331,424,1384,1410]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="23" pageNumber="24">Fig. 52</figureCitation>
): ovular, elongated, yellow brown (in alcohol); with 4 discernible hair
<typeStatus id="54DE8805764AFFBF9702FA92FAD7D1D4" box="[1286,1349,1384,1410]" pageId="23" pageNumber="24">types</typeStatus>
(A1 to A4); dorsally with 3 hair
<typeStatus id="54DE8805764AFFBF93C6FA77FD96D1F1" box="[450,516,1421,1447]" pageId="23" pageNumber="24">types</typeStatus>
;
<typeStatus id="54DE8805764AFFBF9014FA77FDD0D1F1" box="[528,578,1421,1447]" pageId="23" pageNumber="24">type</typeStatus>
A1, long dark blunt tipped, uniform in color;
<typeStatus id="54DE8805764AFFBF9667FA77FB07D1F1" box="[1123,1173,1421,1447]" pageId="23" pageNumber="24">type</typeStatus>
A2, mid-length, dark, but distally spiniform, unlike
<typeStatus id="54DE8805764AFFBF900EFA48FDADD19A" box="[522,575,1458,1484]" pageId="23" pageNumber="24">type</typeStatus>
1;
<typeStatus id="54DE8805764AFFBF906DFA48FD0CD19A" box="[617,670,1458,1484]" pageId="23" pageNumber="24">type</typeStatus>
A3, form dense mat of abdomen, mid-length, uniformly dark colored, needle-form (some stand erect, some run posteriorly along abdomen, both share similar morphology). Longest hairs (
<typeStatus id="54DE8805764AFFBF9348FA07FE10D241" box="[332,386,1533,1559]" pageId="23" pageNumber="24">type</typeStatus>
A1) more concentrated posteriorly toward spinnerets, pointing distad, but proximally, hairs stand erect and medially curved (toward posterior). Ventral hair
<typeStatus id="54DE8805764AFFBF91D7F9D8FB83D26A" box="[979,1041,1570,1596]" pageId="23" pageNumber="24">types</typeStatus>
almost same as dorsal, with
<typeStatus id="54DE8805764AFFBF976DF9D8FA0ED26A" box="[1385,1436,1570,1596]" pageId="23" pageNumber="24">type</typeStatus>
A2 and A3 denser entirely.
<typeStatus id="54DE8805764AFFBF93DFF9B2FD87D234" box="[475,533,1608,1634]" pageId="23" pageNumber="24">Type</typeStatus>
4 hair (A4) in very dense patch, trapezoid in shape, just posterior to pedicel, between median regions of cox. IV. Hairs of medium length, but wavy, pallid. Between anterior pair of booklungs, setal mat jet black, much darker than rest with more concentrated setal counts (
<figureCitation id="135E2A22764AFFBF96BAF968FA87D2FA" box="[1214,1301,1682,1708]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="23" pageNumber="24">Fig. 52</figureCitation>
).
</paragraph>
<paragraph id="8BDA36A7764AFFBF92C0F942FC05D3DA" blockId="23.[151,1437,152,2007]" pageId="23" pageNumber="24">
Genitalia (
<figureCitation id="135E2A22764AFFBF9341F942FE33D284" box="[325,417,1720,1746]" captionStart="FIGURES 47 – 52" captionStartId="20.[151,271,1644,1668]" captionTargetBox="[151,1436,194,1619]" captionTargetId="figure@20.[151,1436,194,1622]" captionTargetPageId="20" captionText="FIGURES 47 – 52. Lyrognathus fuscus sp. nov. holotype female: 47. Carapace, dorsal view. 48. Eyes, dorsal view. 49. Sternum, labium, maxillae and coxa, ventral view. 50. Labium, ventral view. 51. Spermathecae, dorsal view. 52. Abdomen, ventral view. Scale bar = 10 mm for 47, 49, 52; 2 mm for 48, 50; 1 mm for 51. Photos: S. C. Nunn." httpUri="https://zenodo.org/record/275739/files/figure.png" pageId="23" pageNumber="24">Fig. 51</figureCitation>
): epigastric fold 6.2 wide. Spermathecae paired, not fused, separated by 0.8 (width including spermathecae 4.1), bilobular apically, each lobe with strong apical rounded appendages, lateral/ outer lobes more defined distally–ovular; lateral lobe distal, widths 0.62 (left)–0.51 (right): medial lobe distal widths 0.32 (left)–0.31 (right), heavily sclerotized gradual transition to weakly sclerotized shafts proximally. Lobes apically swollen, lateral lobes (including shaft length) 4/5 length to medials (1.12 to 1.2, respectively). Epigastric fold extends to about 2/3 length of spermathecal lobes.
</paragraph>
<paragraph id="8BDA36A7764AFFB092C0F862FC0DD481" blockId="23.[151,1437,152,2007]" lastBlockId="24.[151,1437,152,439]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
Spinnerets: PMS: length 0.71, width (medially) 0.56. PLS: section lengths; basal 2.75, medial 1.46, apical 2.21. PLS section widths (medially); basal 1.35, medial 1.18, apical 0.75. Spinnerets with 2 discernible hair
<typeStatus id="54DE88057645FFB09293FF62FF44D4E4" box="[151,214,152,178]" pageId="24" pageNumber="25">types</typeStatus>
(SP1, SP2): SP1 very short and distally blunt pallid, covers both pairs ventrally; second form (SP2), very similar but up to three times longer, dorsally on all segments.
</paragraph>
<paragraph id="8BDA36A77645FFB092C0FF18FEEED4AA" blockId="24.[151,1437,152,439]" box="[196,380,226,252]" pageId="24" pageNumber="25">Male unknown.</paragraph>
</subSubSection>
<subSubSection id="C37F652C7645FFB092C0FEFDFE1ED5E1" pageId="24" pageNumber="25" type="distribution">
<paragraph id="8BDA36A77645FFB092C0FEFDFE1ED5E1" blockId="24.[151,1437,152,439]" pageId="24" pageNumber="25">
<emphasis id="B911EAB57645FFB092C0FEFDFDE6D577" bold="true" box="[196,628,263,289]" pageId="24" pageNumber="25">Distribution and natural history</emphasis>
(
<figureCitation id="135E2A227645FFB09082FEF2FD67D574" box="[646,757,264,290]" captionStart="FIGURE 114" captionStartId="38.[151,255,1297,1321]" captionTargetBox="[151,1436,194,1276]" captionTargetId="figure@38.[151,1436,194,1276]" captionTargetPageId="38" captionText="FIGURE 114. Records of Lyrognathus species in Southeast Asia. C. Senescall" httpUri="https://zenodo.org/record/275751/files/figure.png" pageId="24" pageNumber="25">Fig. 114</figureCitation>
): Known only from the
<typeStatus id="54DE88057645FFB09620FEF2FBC8D574" box="[1060,1114,264,290]" pageId="24" pageNumber="25">type</typeStatus>
locality, Simpang Kubur, Katawaringin, Kotawaringin Barat, Borneo,
<collectingCountry id="F37276377645FFB090BFFED7FCA4D511" box="[699,822,301,327]" name="Indonesia" pageId="24" pageNumber="25">Indonesia</collectingCountry>
. Little is known; however,
<taxonomicName id="4C654D247645FFB0968CFED7FB65D510" box="[1160,1271,301,326]" class="Arachnida" family="Theraphosidae" genus="Lyrognathus" kingdom="Animalia" order="Araneae" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="fuscus">
<emphasis id="B911EAB57645FFB0968CFED7FB65D510" box="[1160,1271,301,326]" italics="true" pageId="24" pageNumber="25">L. fuscus</emphasis>
</taxonomicName>
is a fossorial spider that lives in higher shaded tropical forests away from the riparian flood plains. Burrows were only found on sloped embankments amongst leaf litter and low vegetation and were formed in clay-like soil (H. Yanto, pers. comm.).
</paragraph>
</subSubSection>
</treatment>
</document>
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