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<document ID-DOI="10.12782/specdiv.27.319" ID-ISSN="2189-7301" ID-Zenodo-Dep="7522710" ID-ZooBank="4CC3122E-2159-49B5-B1FA-43058C529CC8" approvalRequired="4" approvalRequired_for_textStreams="4" checkinTime="1673382120546" checkinUser="felipe" docAuthor="Suzuki, Yuya &amp; Tatsuta, Haruki" docDate="2022" docId="03CC87B0FF84281EFC18DAC2FBF3FB7C" docLanguage="en" docName="SpeciesDiversity.27.2.319-328.pdf" docOrigin="Species Diversity 27 (2)" docSource="http://dx.doi.org/10.12782/specdiv.27.319" docStyle="DocumentStyle:27FECD2C3CD153DA405029F2D33C9514.5:SpeciesDiversity.2012-.journal_article" docStyleId="27FECD2C3CD153DA405029F2D33C9514" docStyleName="SpeciesDiversity.2012-.journal_article" docStyleVersion="5" docTitle="Cladothela bicolor Suzuki &amp; Tatsuta 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="323" masterDocId="FFF5FFC8FF85281AFFADDB4BFFA3FFF1" masterDocTitle="Two New Species of Ground Spiders (Araneae: Gnaphosidae) from Okinawa Islands, Japan" masterLastPageNumber="328" masterPageNumber="319" pageNumber="320" updateTime="1673382738774" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Two New Species of Ground Spiders (Araneae: Gnaphosidae) from Okinawa Islands, Japan</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Suzuki, Yuya</mods:namePart>
<mods:affiliation>Graduate School of Systems Life Sciences, Kyushu University, 744 Motooka, Fukuoka 819 - 0395, Japan E-mail: sasaganiya 1206 @ gmail. com &amp; The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24 Korimoto, Kagoshima 890 - 0065, Japan &amp; Corresponding author</mods:affiliation>
<mods:nameIdentifier type="email">sasaganiya1206@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Tatsuta, Haruki</mods:namePart>
<mods:affiliation>Graduate School of Systems Life Sciences, Kyushu University, 744 Motooka, Fukuoka 819 - 0395, Japan E-mail: sasaganiya 1206 @ gmail. com</mods:affiliation>
<mods:nameIdentifier type="email">sasaganiya1206@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Species Diversity</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2022</mods:date>
<mods:detail type="pubDate">
<mods:number>2022-10-13</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>27</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>2</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>319</mods:start>
<mods:end>328</mods:end>
</mods:extent>
</mods:part>
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<mods:location>
<mods:url>http://dx.doi.org/10.12782/specdiv.27.319</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.12782/specdiv.27.319</mods:identifier>
<mods:identifier type="ISSN">2189-7301</mods:identifier>
<mods:identifier type="Zenodo-Dep">7522710</mods:identifier>
<mods:identifier type="ZooBank">4CC3122E-2159-49B5-B1FA-43058C529CC8</mods:identifier>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7522676" ID-Zenodo-Dep="7522676" LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C" httpUri="http://treatment.plazi.org/id/03CC87B0FF84281EFC18DAC2FBF3FB7C" lastPageId="4" lastPageNumber="323" pageId="1" pageNumber="320">
<subSubSection pageId="1" pageNumber="320" type="nomenclature">
<paragraph blockId="1.[949,1303,393,417]" box="[949,1303,393,417]" pageId="1" pageNumber="320">
<heading box="[949,1303,393,417]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="320" reason="2">
<taxonomicName authority="Suzuki" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[949,1218,393,417]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="320" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis bold="true" box="[949,1141,393,417]" italics="true" pageId="1" pageNumber="320">Cladothela bicolor</emphasis>
Suzuki
</taxonomicName>
,
<taxonomicNameLabel box="[1227,1303,394,417]" pageId="1" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
<paragraph blockId="1.[904,1349,426,483]" pageId="1" pageNumber="320">
[New Japanese Name: Himune-washigumo] (
<figureCitation box="[1082,1169,459,483]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="1.[156,187,2000,2022]" captionStartId-1="2.[156,187,1921,1943]" captionStartId-2="3.[157,188,1469,1491]" captionTargetBox-0="[174,1417,966,1960]" captionTargetBox-1="[170,1418,207,1881]" captionTargetBox-2="[170,1418,207,1430]" captionTargetId-0="figure-278@1.[168,1418,950,1962]" captionTargetId-1="figure-7@2.[169,1419,205,1882]" captionTargetId-2="figure-256@3.[169,1419,205,1431]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionText-0="Fig. 1. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, D) and paratype male from Okinawa Island (NSMT-Ar 22040: B, E, F). A, B, Habitus of living individuals; C, E, habitus, dorsal view; D, F, ditto, ventral view. Scale bars: 1.0mm." captionText-1="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." captionText-2="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi-0="http://doi.org/10.5281/zenodo.7522712" figureDoi-1="http://doi.org/10.5281/zenodo.7522714" figureDoi-2="http://doi.org/10.5281/zenodo.7522718" httpUri-0="https://zenodo.org/record/7522712/files/figure.png" httpUri-1="https://zenodo.org/record/7522714/files/figure.png" httpUri-2="https://zenodo.org/record/7522718/files/figure.png" pageId="1" pageNumber="320">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection lastPageId="3" lastPageNumber="322" pageId="1" pageNumber="320" type="materials_examined">
<paragraph blockId="1.[821,1432,523,906]" pageId="1" pageNumber="320">
<emphasis bold="true" box="[846,1010,523,548]" pageId="1" pageNumber="320">Type material.</emphasis>
<materialsCitation collectingDate="2021-06-02" collectorName="K. Okazaki" country="Japan" county="Kunigami District" latitude="26.643978" location="Sesoko Island" longLatPrecision="1" longitude="127.867065" municipality="Motobu Town" pageId="1" pageNumber="320" specimenCount="1" stateProvince="Okinawa" typeStatus="holotype">
<typeStatus box="[1035,1132,524,548]" pageId="1" pageNumber="320">Holotype</typeStatus>
: female, NSMT-Ar 22039,
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFC98D967FC64FDB5" box="[821,967,556,580]" country="Japan" county="Kunigami District" latitude="26.643978" longLatPrecision="1" longitude="127.867065" municipality="Motobu Town" name="Sesoko Island" pageId="1" pageNumber="320" stateProvince="Okinawa">Sesoko Island</location>
,
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFC7BD967FBBDFDB5" box="[982,1054,556,580]" country="Japan" county="Kunigami District" latitude="26.643978" longLatPrecision="1" longitude="127.867065" municipality="Motobu Town" name="Sesoko" pageId="1" pageNumber="320" stateProvince="Okinawa">Sesoko</location>
,
<collectingMunicipality box="[1069,1221,556,580]" pageId="1" pageNumber="320">Motobu Town</collectingMunicipality>
,
<collectingCounty box="[1236,1427,556,580]" pageId="1" pageNumber="320">Kunigami District</collectingCounty>
,
<collectingRegion box="[821,914,589,613]" country="Japan" name="Okinawa" pageId="1" pageNumber="320">Okinawa</collectingRegion>
,
<collectingCountry box="[925,981,589,612]" name="Japan" pageId="1" pageNumber="320">Japan</collectingCountry>
(
<geoCoordinate box="[996,1131,589,613]" degrees="26.643978" direction="north" orientation="latitude" pageId="1" pageNumber="320" precision="1" value="26.643978">26.643978°N</geoCoordinate>
,
<geoCoordinate box="[1143,1285,589,613]" degrees="127.867065" direction="east" orientation="longitude" pageId="1" pageNumber="320" precision="1" value="127.867065">127.867065°E</geoCoordinate>
),
<date box="[1304,1428,589,613]" pageId="1" pageNumber="320" value="2021-06-02">
<collectingDate box="[1304,1428,589,613]" pageId="1" pageNumber="320" value="2021-06-02">2 June 2021</collectingDate>
</date>
,
<collectorName box="[821,939,621,645]" pageId="1" pageNumber="320">K. Okazaki</collectorName>
leg.
</materialsCitation>
<materialsCitation collectingDate="2021-04-15" country="Japan" county="Kunigami District" latitude="26.694485" location="Kijyoka" longLatPrecision="1" longitude="128.14673" municipality="Ogimi Village" pageId="1" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
<typeStatus box="[998,1098,622,646]" pageId="1" pageNumber="320">Paratypes</typeStatus>
:
<specimenCount box="[1114,1189,621,645]" pageId="1" pageNumber="320" type="male">1 male</specimenCount>
, NSMT-Ar 22040,
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFADAD926FCCDFD57" country="Japan" county="Kunigami District" latitude="26.694485" longLatPrecision="1" longitude="128.14673" municipality="Ogimi Village" name="Kijyoka" pageId="1" pageNumber="320" stateProvince="Okinawa">Kijyoka</location>
,
<collectingMunicipality box="[893,1042,654,681]" pageId="1" pageNumber="320">Ōgimi Village</collectingMunicipality>
,
<collectingCounty box="[1058,1250,654,678]" pageId="1" pageNumber="320">Kunigami District</collectingCounty>
,
<collectingRegion box="[1266,1359,654,678]" country="Japan" name="Okinawa" pageId="1" pageNumber="320">Okinawa</collectingRegion>
,
<collectingCountry box="[1375,1431,654,677]" name="Japan" pageId="1" pageNumber="320">Japan</collectingCountry>
(
<geoCoordinate box="[827,962,686,710]" degrees="26.694485" direction="north" orientation="latitude" pageId="1" pageNumber="320" precision="1" value="26.694485">26.694485°N</geoCoordinate>
,
<geoCoordinate box="[973,1116,686,710]" degrees="128.146732" direction="east" orientation="longitude" pageId="1" pageNumber="320" precision="1" value="128.14673">128.146732°E</geoCoordinate>
),
<date box="[1134,1277,686,710]" pageId="1" pageNumber="320" value="2021-04-15">
<collectingDate box="[1134,1277,686,710]" pageId="1" pageNumber="320" value="2021-04-15">15 April 2021</collectingDate>
</date>
, Y
</materialsCitation>
<materialsCitation box="[1304,1424,686,710]" collectingDate="2021-04-15" collectorName="Suzuki" country="Japan" county="Kunigami District" isTailEnumeration="true" latitude="26.694485" location="Kijyoka" longLatPrecision="1" longitude="128.14673" municipality="Ogimi Village" pageId="1" pageNumber="320" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
.
<collectorName box="[1314,1383,686,710]" pageId="1" pageNumber="320">Suzuki</collectorName>
leg
</materialsCitation>
<materialsCitation collectingDate="2022-05-18" country="Japan" county="Shimajiri District" latitude="26.348345" location="Kume Island" longLatPrecision="1" longitude="126.75505" municipality="Kumejima Town" pageId="1" pageNumber="320" specimenCount="1" specimenCount-male="1" stateProvince="Okinawa" typeStatus="paratype">
.;
<specimenCount box="[821,897,719,743]" pageId="1" pageNumber="320" type="male">1 male</specimenCount>
, NSMT-Ar 22041,
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFBF5D984FB40FD16" box="[1112,1251,719,743]" country="Japan" county="Shimajiri District" latitude="26.348345" longLatPrecision="1" longitude="126.75505" municipality="Kumejima Town" name="Kume Island" pageId="1" pageNumber="320" stateProvince="Okinawa">Kume Island</location>
,
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFB59D99BFABFFD1B" box="[1268,1308,720,746]" country="Japan" county="Shimajiri District" latitude="26.348345" longLatPrecision="1" longitude="126.75505" municipality="Kumejima Town" name="Ota" pageId="1" pageNumber="320" stateProvince="Okinawa">Ōta</location>
,
<collectingMunicipality pageId="1" pageNumber="320">Kumejima Town</collectingMunicipality>
,
<collectingCounty box="[898,1085,751,775]" pageId="1" pageNumber="320">Shimajiri District</collectingCounty>
,
<collectingRegion box="[1103,1196,751,775]" country="Japan" name="Okinawa" pageId="1" pageNumber="320">Okinawa</collectingRegion>
,
<collectingCountry box="[1215,1271,752,775]" name="Japan" pageId="1" pageNumber="320">Japan</collectingCountry>
(
<geoCoordinate box="[1293,1428,751,775]" degrees="26.348345" direction="north" orientation="latitude" pageId="1" pageNumber="320" precision="1" value="26.348345">26.348345°N</geoCoordinate>
,
<geoCoordinate box="[821,963,784,808]" degrees="126.755051" direction="east" orientation="longitude" pageId="1" pageNumber="320" precision="1" value="126.75505">126.755051°E</geoCoordinate>
),
<date box="[979,1110,784,808]" pageId="1" pageNumber="320" value="2022-05-18">
<collectingDate box="[979,1110,784,808]" pageId="1" pageNumber="320" value="2022-05-18">18 May 2022</collectingDate>
</date>
, N
</materialsCitation>
<materialsCitation box="[1141,1272,784,808]" collectingDate="2022-05-18" collectorName="Nakama" country="Japan" county="Shimajiri District" isTailEnumeration="true" latitude="26.348345" location="Kume Island" longLatPrecision="1" longitude="126.75505" municipality="Kumejima Town" pageId="1" pageNumber="320" specimenCount="1" stateProvince="Okinawa" typeStatus="paratype">
.
<collectorName box="[1150,1235,784,808]" pageId="1" pageNumber="320">Nakama</collectorName>
leg
</materialsCitation>
.
</paragraph>
<paragraph blockId="1.[821,1432,523,906]" lastBlockId="3.[156,767,1662,2077]" lastPageId="3" lastPageNumber="322" pageId="1" pageNumber="320">
<emphasis bold="true" box="[846,1158,816,840]" pageId="1" pageNumber="320">Other specimens examined.</emphasis>
<materialsCitation box="[1184,1275,816,840]" location="One" pageId="1" pageNumber="320" specimenCount="1">
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFB0DD87BFB68FCB9" box="[1184,1227,816,840]" name="One" pageId="1" pageNumber="320">One</location>
juv
</materialsCitation>
.
<materialsCitation collectingDate="2022-05-16" country="Japan" county="Shimajiri District" latitude="26.365307" location="Kuroishimori Park" longLatPrecision="1" longitude="126.805786" municipality="Kumejima Town" pageId="1" pageNumber="320" specimenCount="1" stateProvince="Okinawa">
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFAA2D87BFCC5FC98" country="Japan" county="Shimajiri District" latitude="26.365307" longLatPrecision="1" longitude="126.805786" municipality="Kumejima Town" name="Kuroishimori Park" pageId="1" pageNumber="320" stateProvince="Okinawa">Kuroishimori Park</location>
,
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF84281BFCDCD81AFC12FC98" box="[881,945,849,873]" country="Japan" county="Shimajiri District" latitude="26.365307" longLatPrecision="1" longitude="126.805786" municipality="Kumejima Town" name="Majya" pageId="1" pageNumber="320" stateProvince="Okinawa">Majya</location>
,
<collectingMunicipality box="[957,1130,849,873]" pageId="1" pageNumber="320">Kumejima Town</collectingMunicipality>
,
<collectingCounty box="[1142,1322,849,873]" pageId="1" pageNumber="320">Shimajiri District</collectingCounty>
,
<collectingRegion box="[1334,1427,849,873]" country="Japan" name="Okinawa" pageId="1" pageNumber="320">Okinawa</collectingRegion>
,
<collectingCountry box="[821,877,882,905]" name="Japan" pageId="1" pageNumber="320">Japan</collectingCountry>
(
<geoCoordinate box="[890,1025,881,905]" degrees="26.365306" direction="north" orientation="latitude" pageId="1" pageNumber="320" precision="1" value="26.365307">26.365306°N</geoCoordinate>
,
<geoCoordinate box="[1035,1177,881,905]" degrees="126.805788" direction="east" orientation="longitude" pageId="1" pageNumber="320" precision="1" value="126.805786">126.805788°E</geoCoordinate>
),
<date box="[1195,1328,881,906]" pageId="1" pageNumber="320" value="2022-05-16">
<collectingDate box="[1195,1328,881,906]" pageId="1" pageNumber="320" value="2022-05-16">16 May 2022</collectingDate>
</date>
, K
</materialsCitation>
<materialsCitation collectingDate="2022-05-16" collectorName="Yoshida" country="Japan" county="Shimajiri District" isTailEnumeration="true" lastPageId="3" lastPageNumber="322" latitude="26.365307" location="Kuroishimori Park" longLatPrecision="1" longitude="126.805786" municipality="Kumejima Town" pageId="1" pageNumber="320" specimenCount="1" stateProvince="Okinawa">
.
<collectorName lastPageId="3" lastPageNumber="322" pageId="1" pageNumber="320">Yoshida</collectorName>
leg
</materialsCitation>
.;
<specimenCount box="[235,291,1662,1686]" pageId="3" pageNumber="322" type="generic">
1 juv
<materialsCitation box="[287,291,1662,1686]" collectingDate="2022-05-18" country="Japan" county="Shimajiri District" latitude="26.36314" location="Nishimei" longLatPrecision="1" longitude="126.76479" municipality="Kumejima Town" pageId="3" pageNumber="322" specimenCount="1" stateProvince="Okinawa">.</materialsCitation>
</specimenCount>
<location LSID="urn:lsid:plazi:treatment:03CC87B0FF84281EFC18DAC2FBF3FB7C:8EBA607DFF862819FE85DD35FE24F967" box="[296,391,1662,1686]" country="Japan" county="Shimajiri District" latitude="26.36314" longLatPrecision="1" longitude="126.76479" municipality="Kumejima Town" name="Nishimei" pageId="3" pageNumber="322" stateProvince="Okinawa">Nishimei</location>
,
<collectingMunicipality box="[401,573,1662,1686]" pageId="3" pageNumber="322">Kumejima Town</collectingMunicipality>
,
<collectingCounty box="[584,763,1662,1686]" pageId="3" pageNumber="322">Shimajiri District</collectingCounty>
,
<collectingRegion box="[156,249,1695,1719]" country="Japan" name="Okinawa" pageId="3" pageNumber="322">Okinawa</collectingRegion>
,
<collectingCountry box="[259,315,1695,1718]" name="Japan" pageId="3" pageNumber="322">Japan</collectingCountry>
(
<geoCoordinate box="[328,463,1695,1719]" degrees="26.363140" direction="north" orientation="latitude" pageId="3" pageNumber="322" precision="1" value="26.36314">26.363140°N</geoCoordinate>
,
<geoCoordinate box="[473,615,1695,1719]" degrees="126.764791" direction="east" orientation="longitude" pageId="3" pageNumber="322" precision="1" value="126.76479">126.764791°E</geoCoordinate>
),
<date box="[632,763,1695,1719]" pageId="3" pageNumber="322" value="2022-05-18">
<collectingDate box="[632,763,1695,1719]" pageId="3" pageNumber="322" value="2022-05-18">18 May 2022</collectingDate>
</date>
, N
<materialsCitation box="[177,309,1727,1751]" collectingDate="2022-05-18" collectorName="Nakama" country="Japan" county="Shimajiri District" isTailEnumeration="true" latitude="26.36314" location="Nishimei" longLatPrecision="1" longitude="126.76479" municipality="Kumejima Town" pageId="3" pageNumber="322" specimenCount="1" stateProvince="Okinawa">
.
<collectorName box="[186,271,1727,1751]" pageId="3" pageNumber="322">Nakama</collectorName>
leg
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7522712" ID-Zenodo-Dep="7522712" httpUri="https://zenodo.org/record/7522712/files/figure.png" pageId="1" pageNumber="320" startId="1.[156,187,2000,2022]" targetBox="[174,1417,966,1960]" targetPageId="1">
<paragraph blockId="1.[156,1433,2000,2077]" pageId="1" pageNumber="320">
Fig. 1.
<taxonomicName authority="Suzuki" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[240,478,2000,2022]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="320" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[240,406,2000,2021]" italics="true" pageId="1" pageNumber="320">Cladothela bicolor</emphasis>
Suzuki
</taxonomicName>
,
<taxonomicNameLabel box="[491,566,2000,2021]" pageId="1" pageNumber="320" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female from Sesoko Island (NSMT-Ar 22039: A, C, D) and paratype male from Okinawa Island (NSMT-Ar 22040: B, E, F). A, B, Habitus of living individuals; C, E, habitus, dorsal view; D, F, ditto, ventral view. Scale bars: 1.0 mm.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7522714" ID-Zenodo-Dep="7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="2" pageNumber="321" startId="2.[156,187,1921,1943]" targetBox="[170,1418,207,1881]" targetPageId="2">
<paragraph blockId="2.[156,1433,1921,2083]" pageId="2" pageNumber="321">
Fig. 2.
<taxonomicName authority="Suzuki" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[247,481,1921,1943]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="321" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[247,411,1921,1942]" italics="true" pageId="2" pageNumber="321">Cladothela bicolor</emphasis>
Suzuki
</taxonomicName>
,
<taxonomicNameLabel box="[492,565,1921,1942]" pageId="2" pageNumber="321" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0 mm (O); 0.5 mm (AD, N); 0.2 mm (EM, PR).
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.7522718" ID-Zenodo-Dep="7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="3" pageNumber="322" startId="3.[157,188,1469,1491]" targetBox="[170,1418,207,1430]" targetPageId="3">
<paragraph blockId="3.[156,1432,1469,1631]" pageId="3" pageNumber="322">
Fig. 3. Genitalia of
<taxonomicName authority="Suzuki" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[357,591,1469,1491]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[357,521,1469,1490]" italics="true" pageId="3" pageNumber="322">Cladothela bicolor</emphasis>
Suzuki
</taxonomicName>
,
<taxonomicNameLabel box="[603,675,1470,1491]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
, holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars: 0.2 mm.
</paragraph>
</caption>
<subSubSection pageId="3" pageNumber="322" type="etymology">
<paragraph blockId="3.[156,767,1662,2077]" pageId="3" pageNumber="322">
<emphasis bold="true" box="[181,301,1760,1784]" pageId="3" pageNumber="322">Etymology.</emphasis>
The specific name refers to the bicolored body of the
<typeStatus box="[277,369,1792,1816]" pageId="3" pageNumber="322">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="4" lastPageNumber="323" pageId="3" pageNumber="322" type="diagnosis">
<paragraph blockId="3.[156,767,1662,2077]" lastBlockId="4.[156,768,198,2077]" lastPageId="4" lastPageNumber="323" pageId="3" pageNumber="322">
<emphasis bold="true" box="[181,293,1825,1849]" pageId="3" pageNumber="322">Diagnosis.</emphasis>
<taxonomicName authority="Suzuki &amp; Tatsuta, 2022" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[319,506,1825,1849]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[319,506,1825,1849]" italics="true" pageId="3" pageNumber="322">Cladothela bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[519,601,1826,1849]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
bears flattened cheliceral fangs and endites, with dense short stiff bristles (
<figureCitation box="[163,264,1890,1914]" captionStart="Fig" captionStartId="1.[156,187,2000,2022]" captionTargetBox="[174,1417,966,1960]" captionTargetId="figure-278@1.[168,1418,950,1962]" captionTargetPageId="1" captionText="Fig. 1. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, D) and paratype male from Okinawa Island (NSMT-Ar 22040: B, E, F). A, B, Habitus of living individuals; C, E, habitus, dorsal view; D, F, ditto, ventral view. Scale bars: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7522712" httpUri="https://zenodo.org/record/7522712/files/figure.png" pageId="3" pageNumber="322">Fig. 1CI</figureCitation>
), a character shared with
<taxonomicName authorityName="Kamura" authorityYear="1997" box="[535,626,1890,1914]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="auster">
<emphasis box="[535,626,1890,1914]" italics="true" pageId="3" pageNumber="322">C. auster</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis italics="true" pageId="3" pageNumber="322">C. oculinotata</emphasis>
</taxonomicName>
(
<bibRefCitation author="Kamura, T." box="[235,378,1923,1947]" pageId="3" pageNumber="322" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura 1991</bibRefCitation>
,
<bibRefCitation author="Kamura, T." box="[390,443,1923,1947]" pageId="3" pageNumber="322" pagination="77 - 81" refId="ref5555" refString="Kamura, T. 1997. Two species of the genera Cladothela and Trachyzelotes (Araneae: Gnaphosidae) from Yaeyama Islands, Southwest Japan. Acta Arachnologica 46: 77 - 81." type="journal article" year="1997">1997</bibRefCitation>
). However, female of
<taxonomicName authority="Suzuki &amp; Tatsuta, 2022" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[669,767,1923,1947]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[669,767,1923,1947]" italics="true" pageId="3" pageNumber="322">C. bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[156,233,1956,1979]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from both species based on the following unique characteristics: an elliptical, transversely wider copulatory openings surrounded with distinctive sclerotized ridges [vs. ridges located on mesial and anterior sides of copulatory openings in
<taxonomicName authorityName="Kamura" authorityYear="1997" box="[1210,1303,1663,1686]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="auster">
<emphasis box="[1210,1303,1663,1686]" italics="true" pageId="3" pageNumber="322">C. auster</emphasis>
</taxonomicName>
; mesial and posterior sides in
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" box="[1006,1148,1695,1719]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis box="[1006,1148,1695,1719]" italics="true" pageId="3" pageNumber="322">C. oculinotata</emphasis>
</taxonomicName>
;
<figureCitation box="[1161,1269,1695,1719]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="3" pageNumber="322">Figs 2K, L</figureCitation>
,
<figureCitation box="[1280,1310,1695,1719]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="3" pageNumber="322">3A</figureCitation>
vs. fig.
<quantity box="[1391,1432,1695,1719]" metricMagnitude="-2" metricUnit="m" metricValue="7.62" pageId="3" pageNumber="322" unit="in" value="3.0">3 in</quantity>
<bibRefCitation author="Kamura, T." box="[821,977,1727,1751]" pageId="3" pageNumber="322" pagination="77 - 81" refId="ref5555" refString="Kamura, T. 1997. Two species of the genera Cladothela and Trachyzelotes (Araneae: Gnaphosidae) from Yaeyama Islands, Southwest Japan. Acta Arachnologica 46: 77 - 81." type="journal article" year="1997">Kamura (1997)</bibRefCitation>
and figs 30,
<quantity box="[1114,1167,1727,1751]" metricMagnitude="-1" metricUnit="m" metricValue="7.874" pageId="3" pageNumber="322" unit="in" value="31.0">31 in</quantity>
<bibRefCitation author="Kamura, T." box="[1175,1336,1727,1751]" pageId="3" pageNumber="322" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura (1991)</bibRefCitation>
]; copulatory ducts as wide as and 2.5 times as tall as spermathecae, and curved, and close to each other before inserting to spermathecae [vs. copulatory ducts straight and narrower than spermatheca width in
<taxonomicName authorityName="Kamura" authorityYear="1997" box="[1048,1138,1858,1881]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="auster">
<emphasis box="[1048,1138,1858,1881]" italics="true" pageId="3" pageNumber="322">C. auster</emphasis>
</taxonomicName>
and 1.5 times as tall as spermathecae and bent before inserting to spermathecae in
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis italics="true" pageId="3" pageNumber="322">C. oculinotata</emphasis>
</taxonomicName>
;
<figureCitation box="[947,1034,1922,1947]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="3" pageNumber="322">Figs 2M</figureCitation>
,
<figureCitation box="[1048,1074,1923,1947]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="3" pageNumber="322">3B</figureCitation>
(CD) vs. fig.
<quantity box="[1223,1265,1922,1947]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="322" unit="in" value="4.0">4 in</quantity>
<bibRefCitation author="Kamura, T." box="[1274,1431,1923,1947]" pageId="3" pageNumber="322" pagination="77 - 81" refId="ref5555" refString="Kamura, T. 1997. Two species of the genera Cladothela and Trachyzelotes (Araneae: Gnaphosidae) from Yaeyama Islands, Southwest Japan. Acta Arachnologica 46: 77 - 81." type="journal article" year="1997">Kamura (1997)</bibRefCitation>
and fig.
<quantity box="[907,961,1955,1979]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="3" pageNumber="322" unit="in" value="32.0">32 in</quantity>
<bibRefCitation author="Kamura, T." box="[970,1130,1955,1979]" pageId="3" pageNumber="322" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura (1991)</bibRefCitation>
]; spermathecae close to each other [vs. further apart from each other in
<taxonomicName authorityName="Kamura" authorityYear="1997" box="[1289,1383,1988,2011]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="auster">
<emphasis box="[1289,1383,1988,2011]" italics="true" pageId="3" pageNumber="322">C. auster</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" box="[821,964,2020,2044]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis box="[821,964,2020,2044]" italics="true" pageId="3" pageNumber="322">C. oculinotata</emphasis>
</taxonomicName>
;
<figureCitation box="[978,1065,2020,2044]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="3" pageNumber="322">Figs 2M</figureCitation>
,
<figureCitation box="[1078,1090,2020,2044]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="3" pageNumber="322">3</figureCitation>
vs. fig.
<quantity box="[1176,1218,2020,2044]" metricMagnitude="-1" metricUnit="m" metricValue="1.016" pageId="3" pageNumber="322" unit="in" value="4.0">4 in</quantity>
<bibRefCitation author="Kamura, T." box="[1227,1384,2020,2044]" pageId="3" pageNumber="322" pagination="77 - 81" refId="ref5555" refString="Kamura, T. 1997. Two species of the genera Cladothela and Trachyzelotes (Araneae: Gnaphosidae) from Yaeyama Islands, Southwest Japan. Acta Arachnologica 46: 77 - 81." type="journal article" year="1997">Kamura (1997)</bibRefCitation>
and fig.
<quantity box="[859,911,2053,2077]" metricMagnitude="-1" metricUnit="m" metricValue="8.128" pageId="3" pageNumber="322" unit="in" value="32.0">32 in</quantity>
<bibRefCitation author="Kamura, T." box="[918,1077,2053,2077]" pageId="3" pageNumber="322" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura (1991)</bibRefCitation>
]. Male of
<taxonomicName authority="Suzuki &amp; Tatsuta, 2022" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[1179,1275,2053,2077]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="322" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[1179,1275,2053,2077]" italics="true" pageId="3" pageNumber="322">C. bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[1282,1359,2053,2076]" pageId="3" pageNumber="322" rank="species">sp. nov.</taxonomicNameLabel>
can be distinguished from
<taxonomicName authorityName="Kamura" authorityYear="1997" box="[360,452,199,222]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="auster">
<emphasis box="[360,452,199,222]" italics="true" pageId="4" pageNumber="323">C. auster</emphasis>
</taxonomicName>
and
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" box="[507,650,198,222]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis box="[507,650,198,222]" italics="true" pageId="4" pageNumber="323">C. oculinotata</emphasis>
</taxonomicName>
by the following characteristics of male palps: embolus relatively thin (embolus width/length 0.056) [vs. relatively thicker in
<taxonomicName authorityName="Kamura" authorityYear="1997" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="auster">
<emphasis italics="true" pageId="4" pageNumber="323">C. auster</emphasis>
</taxonomicName>
: embolus width/length 0.106; and thinner in
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis italics="true" pageId="4" pageNumber="323">C. oculinotata</emphasis>
</taxonomicName>
: embolus width/length 0.032;
<figureCitation box="[548,629,328,353]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Figs 2Q</figureCitation>
,
<figureCitation box="[642,672,329,353]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="4" pageNumber="323">3D</figureCitation>
vs. fig.
<quantity metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="323" unit="in" value="1.0">1 in</quantity>
<bibRefCitation author="Kamura, T." box="[183,338,361,385]" pageId="4" pageNumber="323" pagination="77 - 81" refId="ref5555" refString="Kamura, T. 1997. Two species of the genera Cladothela and Trachyzelotes (Araneae: Gnaphosidae) from Yaeyama Islands, Southwest Japan. Acta Arachnologica 46: 77 - 81." type="journal article" year="1997">Kamura (1997)</bibRefCitation>
and fig.
<quantity box="[428,480,361,385]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="4" pageNumber="323" unit="in" value="26.0">26 in</quantity>
<bibRefCitation author="Kamura, T." box="[487,646,361,385]" pageId="4" pageNumber="323" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura (1991)</bibRefCitation>
], conductor with a dark-colored distinct angular ridge on the retrolateral margin, and base of the conductor having a wide, whitish, non-sclerotized membranous region [vs. conductor having a sclerotized lobe-like structure in
<taxonomicName authorityName="Kamura" authorityYear="1997" box="[512,603,492,515]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="auster">
<emphasis box="[512,603,492,515]" italics="true" pageId="4" pageNumber="323">C. auster</emphasis>
</taxonomicName>
, ridge on retrolateral margin of conductor rounded in
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" box="[570,712,524,548]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis box="[570,712,524,548]" italics="true" pageId="4" pageNumber="323">C. oculinotata</emphasis>
</taxonomicName>
;
<figureCitation captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Figs 2PR</figureCitation>
,
<figureCitation box="[225,282,556,580]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="4" pageNumber="323">3CE</figureCitation>
vs. fig.
<quantity box="[366,407,556,580]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="323" unit="in" value="1.0">1 in</quantity>
<bibRefCitation author="Kamura, T." box="[416,572,556,580]" pageId="4" pageNumber="323" pagination="77 - 81" refId="ref5555" refString="Kamura, T. 1997. Two species of the genera Cladothela and Trachyzelotes (Araneae: Gnaphosidae) from Yaeyama Islands, Southwest Japan. Acta Arachnologica 46: 77 - 81." type="journal article" year="1997">Kamura (1997)</bibRefCitation>
and fig.
<quantity box="[668,722,556,580]" metricMagnitude="-1" metricUnit="m" metricValue="6.604" pageId="4" pageNumber="323" unit="in" value="26.0">26 in</quantity>
<bibRefCitation author="Kamura, T." pageId="4" pageNumber="323" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura (1991)</bibRefCitation>
]; tegulum very weakly protruded [vs. strongly protruded in
<taxonomicName baseAuthorityName="Bosenberg and Strand" baseAuthorityYear="1906" box="[293,434,621,645]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="oculinotata">
<emphasis box="[293,434,621,645]" italics="true" pageId="4" pageNumber="323">C. oculinotata</emphasis>
</taxonomicName>
;
<figureCitation box="[446,524,621,645]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Figs 2R</figureCitation>
,
<figureCitation box="[534,561,621,645]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="4" pageNumber="323">3E</figureCitation>
vs. fig.
<quantity box="[638,690,621,645]" metricMagnitude="-1" metricUnit="m" metricValue="6.858" pageId="4" pageNumber="323" unit="in" value="27.0">27 in</quantity>
<bibRefCitation author="Kamura, T." pageId="4" pageNumber="323" pagination="47 - 60" refId="ref5494" refString="Kamura, T. 1991. A revision of the genus Cladothela (Araneae: Gnaphosidae) from Japan. Acta Arachnologica 40: 47 - 60." type="journal article" year="1991">Kamura (1991)</bibRefCitation>
].
</paragraph>
<paragraph blockId="4.[156,768,198,2077]" pageId="4" pageNumber="323">
Live individuals of
<taxonomicName authority="Suzuki &amp; Tatsuta, 2022" authorityName="Suzuki &amp; Tatsuta" authorityYear="2022" box="[384,482,686,710]" class="Arachnida" family="Gnaphosidae" genus="Cladothela" kingdom="Animalia" order="Araneae" pageId="4" pageNumber="323" phylum="Arthropoda" rank="species" species="bicolor" status="sp. nov.">
<emphasis box="[384,482,686,710]" italics="true" pageId="4" pageNumber="323">C. bicolor</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[491,570,687,710]" pageId="4" pageNumber="323" rank="species">sp. nov.</taxonomicNameLabel>
possess a clear bicolor body: vivid red prosoma and black abdomen (
<figureCitation captionStart="Fig" captionStartId="1.[156,187,2000,2022]" captionTargetBox="[174,1417,966,1960]" captionTargetId="figure-278@1.[168,1418,950,1962]" captionTargetPageId="1" captionText="Fig. 1. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, D) and paratype male from Okinawa Island (NSMT-Ar 22040: B, E, F). A, B, Habitus of living individuals; C, E, habitus, dorsal view; D, F, ditto, ventral view. Scale bars: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7522712" httpUri="https://zenodo.org/record/7522712/files/figure.png" pageId="4" pageNumber="323">Fig. 1A, B</figureCitation>
), readily to distinguish the new species from its congeners, which possess uniformly dark reddish brown or dark brown body color.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="323" type="description">
<paragraph blockId="4.[156,768,198,2077]" pageId="4" pageNumber="323">
<emphasis bold="true" box="[181,313,849,873]" pageId="4" pageNumber="323">Description.</emphasis>
<emphasis box="[339,410,849,873]" italics="true" pageId="4" pageNumber="323">Female</emphasis>
(
<typeStatus box="[442,535,849,873]" pageId="4" pageNumber="323">holotype</typeStatus>
: NSMT-Ar 22039). Body 5.66 long. Carapace 2.44 long, 2.07 wide, 0.74 high. Eye size and interdistances: AME 0.184, ALE 0.211, PME 0.200, PLE 0.157, AMEAME 0.091, AMEALE 0.038, PMEPME 0.084, PMEPLE 0.069, ALEPLE 0.059. Leg length: I: 1.87+0.99+1.41+0.91+0.69=5.87; II: 1.60+ 0.93+1.28+0.85+0.68=5.34; III: 1.39+0.82+1.06+0.92+ 0.70=4.89; IV: 1.79+0.93+1.45+1.62+0.95=6.74. Abdomen 3.06 long, 2.33 wide, 1.71 high.
</paragraph>
<paragraph blockId="4.[156,768,198,2077]" pageId="4" pageNumber="323">
Chelicerae as long as wide (length/width 1.21) with 3 small promarginal teeth almost in form of small denticles and bearing dense short stiff bristles, retromarginal teeth absent. Fang wide, flat and curved. Epigyne (
<figureCitation box="[607,713,1239,1263]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Figs 2K, L</figureCitation>
,
<figureCitation box="[723,756,1239,1263]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="4" pageNumber="323">3A</figureCitation>
): epigynal plate trapezoidal, wider than long (plate length/ width 0.78); pair of window-like, weakly pigmented regions on anterolateral sides of epigynal plate (arrows in
<figureCitation box="[677,756,1337,1361]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Fig. 2K</figureCitation>
); posteromedian part weakly sclerotized with two pigmented, pit-like structures on posterior margin (arrows in
<figureCitation box="[680,756,1402,1426]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Fig. 2L</figureCitation>
); interdistace of copulatory openings longer than copulatory opening width: interdistance/width 2.15, elliptical, transversely wider with clear sclerotized ridges surrounding the openings; anterior side of epigynal plate sclerotized and pigmented, posterior side membranous and less pigmented. Internal genitalia (
<figureCitation box="[348,437,1597,1621]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Figs 2M</figureCitation>
,
<figureCitation box="[450,480,1597,1621]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="4" pageNumber="323">3B</figureCitation>
): copulatory ducts massive, elongated, curved, contiguous before inserting to spermathecae; lateral glands attenuating transversely; spermathecae globular, juxtaposed; fertilization ducts arising from mesial sides of spermathecae.
</paragraph>
<paragraph blockId="4.[156,768,198,2077]" pageId="4" pageNumber="323">
Coloration (
<figureCitation box="[306,364,1760,1784]" captionStart="Fig" captionStartId="1.[156,187,2000,2022]" captionTargetBox="[174,1417,966,1960]" captionTargetId="figure-278@1.[168,1418,950,1962]" captionTargetPageId="1" captionText="Fig. 1. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, D) and paratype male from Okinawa Island (NSMT-Ar 22040: B, E, F). A, B, Habitus of living individuals; C, E, habitus, dorsal view; D, F, ditto, ventral view. Scale bars: 1.0mm." figureDoi="http://doi.org/10.5281/zenodo.7522712" httpUri="https://zenodo.org/record/7522712/files/figure.png" pageId="4" pageNumber="323">Fig. 1</figureCitation>
). Carapace vividly red in live individuals, and reddish brown in specimens preserved in alcohol. Mouthparts and sternum reddish brown. Legs dark reddish brown with coxae, trochanters, metatarsi and tarsi pale. Dorsum of abdomen black in live individuals, dark grey in preserved specimens, with 2 pairs of sigilla; venter of abdomen pale whitish brown.
</paragraph>
<paragraph blockId="4.[156,768,198,2077]" lastBlockId="4.[820,1438,198,1165]" pageId="4" pageNumber="323">
<emphasis box="[181,231,1987,2011]" italics="true" pageId="4" pageNumber="323">Male</emphasis>
(
<typeStatus box="[246,339,1988,2012]" pageId="4" pageNumber="323">paratype</typeStatus>
: NSMT-Ar 22040). Body 5.58 long. Carapace 2.55 long, 2.11 wide, 0.76 high. Eye size and interdistances: AME 0.153, ALE 0.181, PME 0.185, PLE 0.164, AMEAME 0.098, AMEALE 0.024, PMEPME 0.062, PMEPLE 0.060, ALEPLE 0.057. Length of legs: I: 1.96+ 0.85+1.51+0.88+0.86=6.06; II: 1.63+0.73+1.27+0.78+ 0.82=5.23; III: 1.30+0.69+0.91+0.90+0.66=4.46; IV: 2.26+0.85+1.51+1.67+0.83=7.12. Abdomen 2.91 long, 1.79 wide, 1.33 high.
</paragraph>
<paragraph blockId="4.[820,1438,198,1165]" pageId="4" pageNumber="323">
Carapace, mouthparts, and legs as in females. Chelicerae as long as wide (length/width 1.28) with 3 promarginal teeth, retromarginal teeth absent. Palp (
<figureCitation box="[1234,1347,458,482]" captionStart="Fig" captionStartId="2.[156,187,1921,1943]" captionTargetBox="[170,1418,207,1881]" captionTargetId="figure-7@2.[169,1419,205,1882]" captionTargetPageId="2" captionText="Fig. 2. Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, C, EG, KM) and paratype male from Okinawa Island (NSMT-Ar 22040: B, D, HJ, NR). A, B, Eye region, dorsal view; C, D, mouthparts, ventral view; E, H, left chelicerae, posterior view; F, I, ditto, anterior view; G, J, ditto, ventral view; K, epigyne, ventral view; L, ditto, clarified view; M, internal genitalia, dorsal view; N, right palp, dorsal view; O, ditto, retrolateral view; P, left palp, retrolateral view; Q, ditto, ventral view; R, ditto, prolateral view. Arrows in K and L indicate pit-like structures on posterior margin and window-like regions on epigynal plate, respectively. Scale bars: 1.0mm (O); 0.5mm (AD, N); 0.2mm (EM, PR)." figureDoi="http://doi.org/10.5281/zenodo.7522714" httpUri="https://zenodo.org/record/7522714/files/figure.png" pageId="4" pageNumber="323">Figs 2NR</figureCitation>
,
<figureCitation box="[1360,1421,458,482]" captionStart="Fig" captionStartId="3.[157,188,1469,1491]" captionTargetBox="[170,1418,207,1430]" captionTargetId="figure-256@3.[169,1419,205,1431]" captionTargetPageId="3" captionText="Fig. 3. Genitalia of Cladothela bicolor Suzuki, sp. nov., holotype female from Sesoko Island (NSMT-Ar 22039: A, B) and paratype male from Okinawa Island (NSMT-Ar 22040: CE). A, Epigyne, ventral view; B, internal genitalia, dorsal view; C, left palp, prolateral view; D, ditto, ventral view; E, ditto, retrolateral view. Abbreviations: CD, copulatory ducts; CDH, copulatory duct height; CDW, copulatory duct width; CO, copulatory opening; Co, conductor; COI, copulatory openings interdistance; COW, copulatosy opening width; Em, embolus; EmL, embolus length; EmW embolus width; FD, fertilization duct; G, lateral gland; Sp, spermatheca; SpH, spermatheca height; spermatheca width; T, tegulum; To, triangular outgrowth. Arrow indicates a hook-like apophysis on retrolateral side of palpal femur. Scale bars:0.2mm." figureDoi="http://doi.org/10.5281/zenodo.7522718" httpUri="https://zenodo.org/record/7522718/files/figure.png" pageId="4" pageNumber="323">3CE</figureCitation>
): femur slightly longer than cymbium (femur length/cymbial length 1.40); base of femoral spine positioned slightly upper than half of femur (distance between basal margin of palpal femur and base of femoral spine/femoral length 0.60); patella longer than tibia (patella length/tibia length 1.50); cymbium longer than wide (length/width 1.68); base of conductor with a wide, whitish, non-sclerotized membranous region; conductor with hooked tip, dark-colored, distinct angular ridge on retrolateral margin (triangular outgrowth); embolus moderately long, weakly twisted, emerging from prolateral base of tegulum. Abdomen with 2 pairs of sigilla and a trapezoid scutum on anterior part of dorsum.
</paragraph>
<paragraph blockId="4.[820,1438,198,1165]" box="[846,1100,881,905]" pageId="4" pageNumber="323">Coloration. As in female.</paragraph>
<paragraph blockId="4.[820,1438,198,1165]" pageId="4" pageNumber="323">
<emphasis bold="true" box="[846,932,913,937]" pageId="4" pageNumber="323">Habitat.</emphasis>
Specimens were collected from litter layers and under stones in primary and secondary broad-leaf forests.
</paragraph>
</subSubSection>
<subSubSection pageId="4" pageNumber="323" type="distribution">
<paragraph blockId="4.[820,1438,198,1165]" pageId="4" pageNumber="323">
<emphasis bold="true" box="[846,985,1011,1035]" pageId="4" pageNumber="323">Distribution.</emphasis>
<collectingRegion box="[1011,1102,1011,1035]" country="Japan" name="Okinawa" pageId="4" pageNumber="323">Okinawa</collectingRegion>
Islands (Sesoko,
<collectingRegion box="[1291,1382,1011,1035]" country="Japan" name="Okinawa" pageId="4" pageNumber="323">Okinawa</collectingRegion>
and Kume Islands),
<collectingCountry box="[978,1037,1044,1067]" name="Japan" pageId="4" pageNumber="323">Japan</collectingCountry>
.
</paragraph>
<paragraph blockId="4.[820,1438,198,1165]" pageId="4" pageNumber="323">
<emphasis bold="true" box="[846,902,1076,1100]" pageId="4" pageNumber="323">Note.</emphasis>
Males and females are regarded conspecific because of the similarity in body color and morphology of cheliceral fangs and endites.
</paragraph>
</subSubSection>
</treatment>
</document>