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<document id="F029FE47F188E31B38996D1E79310A30" ID-CLB-Dataset="20367" ID-DOI="10.11646/zootaxa.5105.1.3" ID-GBIF-Dataset="16f0d04d-e6ef-4eba-8bb3-9c5ebd3f778f" ID-ISSN="1175-5326" ID-Zenodo-Dep="6332331" ID-ZooBank="8B30E589-CEFC-415B-9505-EAE71C204E61" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1646596273552" checkinUser="plazi" docAuthor="Lencioni, F. A. A." docDate="2022" docId="03CC87902779AA236CB1FE6A6C97F9A9" docLanguage="en" docName="zootaxa.5105.1.3.pdf" docOrigin="Zootaxa 5105 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Fredyagrion elongatum Lencioni 2022, comb. nov." docType="treatment" docVersion="8" lastPageNumber="78" masterDocId="FFF5FFE82776AA2C6C26FFC26F0CFFAF" masterDocTitle="Analysis of male Leptagrion Selys, 1876 sensu lato (Odonata: Coenagrionidae) with description of four new genera and a new species" masterLastPageNumber="104" masterPageNumber="63" pageNumber="78" updateTime="1699317828930" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="19E1C08D2C712BAECAF50ECF553070C9">Analysis of male Leptagrion Selys, 1876 sensu lato (Odonata: Coenagrionidae) with description of four new genera and a new species</mods:title>
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<mods:namePart id="018E0697421CA0083A559216088977AD">Lencioni, F. A. A.</mods:namePart>
<mods:affiliation id="F7B4FEABE73FAE1D35BF21CA5B4C9412">Private researcher, Rua Anibal, 216 - Jd. Coleginho, Vila Zezé, Jacareí, CEP (ZIP) 12310 - 780, São Paulo. Brazil.</mods:affiliation>
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<mods:title id="FF8D9F2EC36B0C9F8EE1EF49537C4D8E">Zootaxa</mods:title>
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<mods:date id="E2F8F50386016C164A800A616F474A97">2022</mods:date>
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<mods:number id="AF5A05BB0FDDB93FFAE31DB84C77A2E3">2022-03-02</mods:number>
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<mods:number id="00BE22A4FCE67280DD636D9468BD5543">5105</mods:number>
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<mods:identifier id="6B51145CA5FB8D3CDF9060396CC46616" type="DOI">10.11646/zootaxa.5105.1.3</mods:identifier>
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<treatment id="03CC87902779AA236CB1FE6A6C97F9A9" ID-DOI="http://doi.org/10.5281/zenodo.6335801" ID-GBIF-Taxon="193878576" ID-Zenodo-Dep="6335801" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9" httpUri="http://treatment.plazi.org/id/03CC87902779AA236CB1FE6A6C97F9A9" lastPageNumber="78" pageId="15" pageNumber="78">
<subSubSection id="C37F650D2779AA236CB1FE6A6DE4FE6C" box="[151,744,424,451]" pageId="15" pageNumber="78" type="nomenclature">
<paragraph id="8BDA36862779AA236CB1FE6A6DE4FE6C" blockId="15.[151,744,424,488]" box="[151,744,424,451]" pageId="15" pageNumber="78">
<heading id="D09281EA2779AA236CB1FE6A6DE4FE6C" bold="true" box="[151,744,424,451]" fontSize="11" level="1" pageId="15" pageNumber="78" reason="1">
<emphasis id="B911EA942779AA236CB1FE6A6DE4FE6C" bold="true" box="[151,744,424,451]" pageId="15" pageNumber="78">
<taxonomicName id="4C654D052779AA236CB1FE6A6D56FE6D" ID-CoL="9X5KB" authority="(Selys, 1876)" authorityName="Lencioni" authorityYear="2022" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[151,602,424,451]" class="Insecta" family="Coenagrionidae" genus="Fredyagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="elongatum" status="comb. nov.">
<emphasis id="B911EA942779AA236CB1FE6A6EBBFE6C" bold="true" box="[151,439,424,451]" italics="true" pageId="15" pageNumber="78">Fredyagrion elongatum</emphasis>
(Selys, 1876)
</taxonomicName>
<taxonomicNameLabel id="A22257EF2779AA236E44FE6B6DE4FE6C" box="[610,744,425,451]" pageId="15" pageNumber="78" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
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</paragraph>
</subSubSection>
<subSubSection id="C37F650D2779AA236CB1FE0F6DD0FE47" box="[151,732,461,488]" pageId="15" pageNumber="78" type="description">
<paragraph id="8BDA36862779AA236CB1FE0F6DD0FE47" blockId="15.[151,744,424,488]" box="[151,732,461,488]" pageId="15" pageNumber="78">
<figureCitation id="135E2A032779AA236CB1FE0F6FEBFE48" box="[151,231,461,487]" captionStart="FIGURE 21" captionStartId="15.[151,250,2015,2039]" captionTargetBox="[151,1436,1570,1986]" captionTargetId="figure-592@15.[151,1436,1565,1991]" captionTargetPageId="15" captionText="FIGURE 21. Fredyagrion elongatum: A—Appendages, lateral view, B—Appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6332377" httpUri="https://zenodo.org/record/6332377/files/figure.png" pageId="15" pageNumber="78">Fig. 21</figureCitation>
(app),
<figureCitation id="135E2A032779AA236D11FE0F6E8AFE48" box="[311,390,461,487]" captionStart="FIGURE 31" captionStartId="20.[151,250,1127,1151]" captionTargetBox="[151,1436,681,1103]" captionTargetId="figure-41@20.[151,1436,679,1104]" captionTargetPageId="20" captionText="FIGURE 31. Fredyagrion dispar: A—Prothorax, lateral view, B— Prothorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6332397" httpUri="https://zenodo.org/record/6332397/files/figure.png" pageId="15" pageNumber="78">Fig. 31</figureCitation>
(prothorax),
<figureCitation id="135E2A032779AA236E3CFE0F6D66FE48" box="[538,618,461,487]" captionStart="FIGURE 40" captionStartId="23.[151,250,1547,1571]" captionTargetBox="[159,1426,1210,1511]" captionTargetId="figure-38@23.[151,1436,1201,1523]" captionTargetPageId="23" captionText="FIGURE 40. Fredyagrion capixabae: Habitus." figureDoi="http://doi.org/10.5281/zenodo.6332415" httpUri="https://zenodo.org/record/6332415/files/figure.png" pageId="15" pageNumber="78">Fig. 40</figureCitation>
(habitus).
</paragraph>
</subSubSection>
<subSubSection id="C37F650D2779AA236CB1FDD26D93FB0C" pageId="15" pageNumber="78" type="etymology">
<paragraph id="8BDA36862779AA236CB1FDD26DACFD86" blockId="15.[151,1437,527,1542]" box="[151,672,527,554]" pageId="15" pageNumber="78">
<emphasis id="B911EA942779AA236CB1FDD26EA8FD85" bold="true" box="[151,420,527,554]" pageId="15" pageNumber="78">
Etymology:
<taxonomicName id="4C654D052779AA236D0FFDCD6EA8FD85" authority="Latin" authorityName="Lencioni" authorityYear="2022" baseAuthorityName="Selys" baseAuthorityYear="1876" box="[297,420,527,554]" class="Insecta" family="Coenagrionidae" genus="Fredyagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="elongatum">
<emphasis id="B911EA942779AA236D0FFDCD6EA8FD85" bold="true" box="[297,420,527,554]" italics="true" pageId="15" pageNumber="78">elongatum</emphasis>
</taxonomicName>
</emphasis>
= Latin for elongated.
</paragraph>
<paragraph id="8BDA36862779AA236CE1FDF16C92FBF1" blockId="15.[151,1437,527,1542]" pageId="15" pageNumber="78">
Selys 1876: 979 (description of male and female);
<bibRefCitation id="EFF44B772779AA236F3DFDF16CD7FDE2" author="Kirby, W. F." box="[795,987,562,589]" pageId="15" pageNumber="78" refId="ref15730" refString="Kirby, W. F. (1890) A synonymic catalogue of Neuroptera Odonata, or Dragonflies. With an appendix of fossil species. Gurney &amp; Jackson, London, ix + 202 pp." type="book" year="1890">Kirby 1890: 155</bibRefCitation>
(data on publication and distribution);
<bibRefCitation id="EFF44B772779AA236CB1FD946E6FFDDE" author="Calvert, P. P." box="[151,355,598,625]" pageId="15" pageNumber="78" pagination="73 - 264" refId="ref14891" refString="Calvert, P. P. (1909) Contributions to a knowledge of the Odonata of the Neotropical region, exclusive of Mexico and Central America. Annals of the Carnegie Museum, 6, 73 - 264." type="journal article" year="1909">Calvert 1909: 158</bibRefCitation>
(data on distribution);
<bibRefCitation id="EFF44B772779AA236E4FFD946C26FDDF" author="Williamson, E. B." box="[617,810,598,624]" pageId="15" pageNumber="78" pagination="241 - 254" refId="ref16997" refString="Williamson, E. B. (1917) Some species of Leptagrion with descriptions of a new genus and a new species. Entomological News, 28, 241 - 254." type="journal article" year="1917">Williamson 1917</bibRefCitation>
: Plate XVII, 9–10 (illustration of appendages in lateral and dorsal views);
<bibRefCitation id="EFF44B772779AA236D49FDBB6D18FD3C" author="Kennedy, C. H." box="[367,532,633,659]" pageId="15" pageNumber="78" refId="ref15683" refString="Kennedy, C. H. (1919) A study of the phylogeny of the Zygoptera from evidence given by the genitalia. Ph. D. thesis, Cornell University, Ithaca, New York, 293 pp., 1396 + 21 figs." type="book" year="1919">Kennedy 1919</bibRefCitation>
: Figs. 1076–1077 (illustration of penis in lateral and ectal views);
<bibRefCitation id="EFF44B772779AA23692CFDBB6FCDFD19" author="Santos, N. D." pageId="15" pageNumber="78" pagination="359 - 362" refId="ref16210" refString="Santos, N. D. (1961) Duas novas especies do genero &quot; Leptagrion &quot; Selys, 1876. (Coenagriidae, Odonata). Revista Brasileira de Biologia, 21 (4), 359 - 362." type="journal article" year="1961">Santos 1961: 360</bibRefCitation>
(compared with
<taxonomicName id="4C654D052779AA236DA7FD5F6EFDFD19" authorityName="Santos" authorityYear="1961" box="[385,497,669,694]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="78" phylum="Arthropoda" rank="species" species="acutum">
<emphasis id="B911EA942779AA236DA7FD5F6EFDFD19" box="[385,497,669,694]" italics="true" pageId="15" pageNumber="78">L. acutum</emphasis>
</taxonomicName>
);
<bibRefCitation id="EFF44B772779AA236E2CFD5E6DBFFD18" author="Santos, N. D." box="[522,691,668,695]" pageId="15" pageNumber="78" pagination="1 - 8" refId="ref16251" refString="Santos, N. D. (1962) Fauna do estado da Guanabara. - descricao de Leptagrion perlongum Calvert, 1909, femea, e notas sobre outras especies do genero. Boletim do Museu Nacional, Rio de Janeiro, 233, 1 - 8." type="journal article" year="1962">Santos 1962: 5</bibRefCitation>
,
<figureCitation id="135E2A032779AA236E99FD5E6C01FD19" box="[703,781,668,694]" captionStart="FIGURE 8" captionStartId="7.[151,250,1279,1303]" captionTargetBox="[151,1430,828,1255]" captionTargetId="figure-30@7.[151,1436,828,1255]" captionTargetPageId="7" captionText="FIGURE 8. Leptagrion auriceps: A—Prothorax, lateral view (holotype); B—Detail." figureDoi="http://doi.org/10.5281/zenodo.6332347" httpUri="https://zenodo.org/record/6332347/files/figure.png" pageId="15" pageNumber="78">Figs. 8</figureCitation>
,
<figureCitation id="135E2A032779AA236F3CFD5E6C3AFD19" box="[794,822,668,694]" captionStart="FIGURE 13" captionStartId="9.[151,250,1653,1677]" captionTargetBox="[181,1416,1231,1621]" captionTargetId="figure-29@9.[151,1436,1218,1629]" captionTargetPageId="9" captionText="FIGURE 13. Leptagrion bocainense: Habitus." figureDoi="http://doi.org/10.5281/zenodo.6332359" httpUri="https://zenodo.org/record/6332359/files/figure.png" pageId="15" pageNumber="78">13</figureCitation>
(data on distribution, female prothorax in dorsal view, female hindwing pterostigma);
<bibRefCitation id="EFF44B772779AA236DD0FD026DA6FD75" author="Santos, N. D." box="[502,682,703,730]" pageId="15" pageNumber="78" pagination="83 - 85" refId="ref16386" refString="Santos, N. D. (1966) Contribuicao ao conhecimento da fauna do Estado da Guanabara. 56. Notas sobre Coenagriideos (Odonata) que se criam em bromelias. Atas da Sociedade de Biologia do Rio de Janeiro, 10 (3), 83 - 85." type="journal article" year="1966">Santos 1966: 83</bibRefCitation>
(data on larvae);
<bibRefCitation id="EFF44B772779AA236F48FD7D6B84FD75" author="Davies, D. A. L. &amp; Tobin, P." box="[878,1160,703,730]" pageId="15" pageNumber="78" pagination="1 - 127" refId="ref15100" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The Dragonflies of the world: A systematic list of the extant species of Odonata Volume I Zygoptera, Anisozygoptera. Societas Internationalis Odonatologica Rapid Communication (Supplement), 3, i - ix + 1 - 127." type="journal article" year="1984">Davies &amp; Tobin 1984: 82</bibRefCitation>
(data on publication and distribution);
<bibRefCitation id="EFF44B772779AA236D14FD216EC1FD52" author="Bridges, C. A." box="[306,461,739,765]" pageId="15" pageNumber="78" refId="ref14771" refString="Bridges, C. A. (1994) Catalogue of the family - group, genus - group and species - group names of the Odonata of the world. 3 rd Edition. C. A. Bridges, Urbana, Illinois, xlvi + 905 pp. https: // doi. org / 10.5962 / bhl. title. 15291" type="book" year="1994">Bridges 1994</bibRefCitation>
: (VII) 77 (data on publication,
<typeStatus id="54DE88242779AA236F17FD216C6DFD52" box="[817,865,739,765]" pageId="15" pageNumber="78">type</typeStatus>
depository,
<typeStatus id="54DE88242779AA236FCBFD216B11FD52" box="[1005,1053,739,765]" pageId="15" pageNumber="78">type</typeStatus>
locality and references);
<bibRefCitation id="EFF44B772779AA236918FD216E79FC8F" author="Costa, J. M. &amp; Garrison, R. W." pageId="15" pageNumber="78" pagination="381 - 394" refId="ref14931" refString="Costa, J. M. &amp; Garrison, R. W. (2001) Description of the female of Leptagrion aculeatum Santos, 1965 with keys to the known species (Zygoptera: Coenagrionidae). Odonatologica, 30 (4), 381 - 394." type="journal article" year="2001">Costa &amp; Garrison 2001: 383</bibRefCitation>
,
<figureCitation id="135E2A032779AA236DA6FCC46D09FC8F" box="[384,517,774,800]" captionStart="FIGURE 7" captionStartId="7.[151,250,780,804]" captionTargetBox="[163,1422,192,754]" captionTargetId="figure-16@7.[154,1433,181,755]" captionTargetPageId="7" captionText="FIGURE 7. Leptagrion vilelai: Penis, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6332345" httpUri="https://zenodo.org/record/6332345/files/figure.png" pageId="15" pageNumber="78">Figs. 7a, 7c</figureCitation>
,
<figureCitation id="135E2A032779AA236E37FCC46D54FC8F" box="[529,600,774,800]" captionStart="FIGURE 16" captionStartId="12.[151,250,693,717]" captionTargetBox="[151,1430,242,666]" captionTargetId="figure-77@12.[151,1436,236,669]" captionTargetPageId="12" captionText="FIGURE 16. Fredyagrion aculeatum: A—Appendages, lateral view, B—Appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6332365" httpUri="https://zenodo.org/record/6332365/files/figure.png" pageId="15" pageNumber="78">16a–b</figureCitation>
,
<figureCitation id="135E2A032779AA236E42FCC46D8EFC8F" box="[612,642,774,800]" captionStart="FIGURE 28" captionStartId="19.[151,250,1510,1534]" captionTargetBox="[151,1436,1064,1483]" captionTargetId="figure-163@19.[151,1436,1059,1487]" captionTargetPageId="19" captionText="FIGURE 28. Fredyagrion andromache: A—Prothorax, lateral view, B— Prothorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6332391" httpUri="https://zenodo.org/record/6332391/files/figure.png" pageId="15" pageNumber="78">28</figureCitation>
,
<figureCitation id="135E2A032779AA236EA8FCC46DD9FC8F" box="[654,725,774,800]" captionStart="FIGURE 43" captionStartId="24.[151,250,1163,1187]" captionTargetBox="[202,1381,839,1134]" captionTargetId="figure-35@24.[194,1393,826,1140]" captionTargetPageId="24" captionText="FIGURE 43. Fredyagrion itabaiana: Habitus." figureDoi="http://doi.org/10.5281/zenodo.6332421" httpUri="https://zenodo.org/record/6332421/files/figure.png" pageId="15" pageNumber="78">43a–b</figureCitation>
,
<figureCitation id="135E2A032779AA236EC7FCC46C02FC8E" box="[737,782,774,801]" captionStart="FIGURE 51" captionStartId="28.[151,250,1981,2005]" captionTargetBox="[151,1435,1537,1957]" captionTargetId="figure-203@28.[151,1436,1537,1957]" captionTargetPageId="28" captionText="FIGURE 51. Machadagrion dardanoi: A—Prothorax, lateral view, B— Prothorax, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6332437" httpUri="https://zenodo.org/record/6332437/files/figure.png" pageId="15" pageNumber="78">51b</figureCitation>
,
<figureCitation id="135E2A032779AA236F3CFCC46C3AFC8E" box="[794,822,774,801]" captionStart="FIGURE 59" captionStartId="33.[151,250,1691,1715]" captionTargetBox="[151,1422,1245,1668]" captionTargetId="figure-479@33.[151,1436,1245,1668]" captionTargetPageId="33" captionText="FIGURE 59. Nathaliagrion perlongum: A—Appendages, lateral view, B—Appendages, dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6332455" httpUri="https://zenodo.org/record/6332455/files/figure.png" pageId="15" pageNumber="78">59</figureCitation>
(key to Brazilian species of
<taxonomicName id="4C654D052779AA23685FFCC56BF9FC8F" authorityName="Selys" authorityYear="1876" box="[1145,1269,775,800]" class="Insecta" family="Coenagrionidae" genus="Leptagrion" kingdom="Animalia" order="Odonata" pageId="15" pageNumber="78" phylum="Arthropoda" rank="genus">
<emphasis id="B911EA942779AA23685FFCC56BF9FC8F" box="[1145,1269,775,800]" italics="true" pageId="15" pageNumber="78">Leptagrion</emphasis>
</taxonomicName>
, illustration of appendages in lateral and dorsal views, male prothorax in dorsal and lateral views, female prothorax in dorsal and lateral views, male forewing pterostigma and female S
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lateral view, female hindwing pterostigma);
<bibRefCitation id="EFF44B772779AA23691FFC8E6E05FC25" author="Lencioni, F. A. A." pageId="15" pageNumber="78" refId="ref15767" refString="Lencioni, F. A. A. (2006) Damselflies of Brazil - An illustrated identification guide. II. - Coenagrionidae. All Print Editora. Sao Paulo, viii + 419 pp." type="book" year="2006">Lencioni 2006: 147</bibRefCitation>
, Figs. 88A–I (data on description,
<typeStatus id="54DE88242779AA236EAFFCB26DB5FC25" box="[649,697,880,906]" pageId="15" pageNumber="78">type</typeStatus>
locality,
<typeStatus id="54DE88242779AA236F38FCB26C42FC25" box="[798,846,880,906]" pageId="15" pageNumber="78">type</typeStatus>
deposition, distribution, and larval description, illustration of appendages in lateral and dorsal views, male and female prothorax in lateral and dorsal views, male fore wing pterostigma, female hind wing pterostigma, female S
<quantity id="4C9D9B632779AA236F02FC746C76FC7F" box="[804,890,950,976]" metricMagnitude="-1" metricUnit="m" metricValue="2.286" metricValueMax="2.54" metricValueMin="2.032" pageId="15" pageNumber="78" unit="in" value="9.0" valueMax="10.0" valueMin="8.0">8–10 in</quantity>
lateral view);
<bibRefCitation id="EFF44B772779AA23683DFC746A1CFC7F" author="Costa, J. M. &amp; Oldrini, B. B. &amp; Anjos-Santos, D." box="[1051,1296,950,976]" pageId="15" pageNumber="78" pagination="93 - 111" refId="ref15056" refString="Costa, J. M., Oldrini, B. B. &amp; Anjos-Santos, D. (2009) Eight new Leptagrion larvae from Brazil (Zygoptera: Coenagrionidae). Odonatologica, 38 (2), 93 - 111." type="journal article" year="2009">
Costa
<emphasis id="B911EA942779AA236847FC756B95FC7F" box="[1121,1177,950,976]" italics="true" pageId="15" pageNumber="78">et al.</emphasis>
2009: 101
</bibRefCitation>
(description and illustration of larva);
<bibRefCitation id="EFF44B772779AA236D91FC1B6DDEFC5C" author="Garrison, R. W. &amp; von Ellenrieder, N. &amp; Louton, J. A." box="[439,722,985,1011]" pageId="15" pageNumber="78" refId="ref15509" refString="Garrison, R. W., von Ellenrieder, N. &amp; Louton, J. A. (2010) Damselfly Genera of the New World: An Illustrated and Annotated Key to the Zygoptera. The Johns Hopkins University Press, Baltimore, Maryland, 490 pp." type="book" year="2010">
Garrison
<emphasis id="B911EA942779AA236E06FC186D55FC5C" box="[544,601,985,1011]" italics="true" pageId="15" pageNumber="78">et al.</emphasis>
2010: 264
</bibRefCitation>
(data on species, illustration of penis in lateral and ectal views, appendages in lateral and dorsal views);
<bibRefCitation id="EFF44B772779AA236E4FFC3E6C5BFBB8" author="Lencioni, F. A. A." box="[617,855,1020,1047]" pageId="15" pageNumber="78" refId="ref15804" refString="Lencioni, F. A. A. (2017 a) Damselflies of Brazil - An illustrated identification guide - Southeast Region. E-book, 559 pp." type="book" year="2017">Lencioni 2017a: 375</bibRefCitation>
, Figs. 219A–D, 220A–B (data on description,
<typeStatus id="54DE88242779AA23694AFC3F6A90FBB8" box="[1388,1436,1021,1047]" pageId="15" pageNumber="78">type</typeStatus>
locality,
<typeStatus id="54DE88242779AA236CDEFBE26E24FB95" box="[248,296,1056,1082]" pageId="15" pageNumber="78">type</typeStatus>
deposition, distribution, and larval description, illustration of appendages in lateral and dorsal views, male prothorax in dorsal and lateral views, male and female habitus).
</paragraph>
<paragraph id="8BDA36862779AA236CE1FBA46C6EFB2E" blockId="15.[151,1437,527,1542]" box="[199,866,1126,1153]" pageId="15" pageNumber="78">
<emphasis id="B911EA942779AA236CE1FBA46EFCFB2F" bold="true" box="[199,496,1126,1152]" pageId="15" pageNumber="78">Deposition of type series.</emphasis>
<materialsCitation id="3B0D3CDB2779AA236DD1FBA46C53FB2E" ID-GBIF-Occurrence="3697349333" box="[503,863,1126,1153]" collectionCode="IRSNB" pageId="15" pageNumber="78" specimenCount="1" typeStatus="syntype">
<typeStatus id="54DE88242779AA236DD1FBA46D52FB2F" box="[503,606,1126,1152]" pageId="15" pageNumber="78" type="syntype">Syntypes</typeStatus>
in
<collectionCode id="ED74AE432779AA236EA4FBA46DD8FB2F" box="[642,724,1126,1152]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="15" pageNumber="78">IRSNB</collectionCode>
(examined)
</materialsCitation>
.
</paragraph>
<paragraph id="8BDA36862779AA236CE1FB4B6D93FB0C" blockId="15.[151,1437,527,1542]" box="[199,671,1161,1188]" pageId="15" pageNumber="78">
<emphasis id="B911EA942779AA236CE1FB4B6E9EFB0C" bold="true" box="[199,402,1161,1187]" pageId="15" pageNumber="78">Larva described.</emphasis>
Yes (
<bibRefCitation id="EFF44B772779AA236DE9FB4B6D9FFB0C" author="Costa, J. M. &amp; Oldrini, B. B. &amp; Anjos-Santos, D." box="[463,659,1161,1187]" pageId="15" pageNumber="78" pagination="93 - 111" refId="ref15056" refString="Costa, J. M., Oldrini, B. B. &amp; Anjos-Santos, D. (2009) Eight new Leptagrion larvae from Brazil (Zygoptera: Coenagrionidae). Odonatologica, 38 (2), 93 - 111." type="journal article" year="2009">
Costa
<emphasis id="B911EA942779AA236E31FB486D43FB0C" box="[535,591,1161,1187]" italics="true" pageId="15" pageNumber="78">et al.</emphasis>
2009
</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F650D2779AA236CE1FB6E6A1EFA4E" pageId="15" pageNumber="78" type="materials_examined">
<paragraph id="8BDA36862779AA236CE1FB6E6A1EFA4E" blockId="15.[151,1437,527,1542]" pageId="15" pageNumber="78">
<emphasis id="B911EA942779AA236CE1FB6E6EB8FB68" bold="true" box="[199,436,1196,1223]" pageId="15" pageNumber="78">Material examined.</emphasis>
(
<specimenCount id="9D63FD0F2779AA236DE1FB6E6D17FB68" box="[455,539,1196,1223]" pageId="15" pageNumber="78" type="male">10 ♂♂</specimenCount>
,
<specimenCount id="9D63FD0F2779AA236E0DFB6E6D61FB68" box="[555,621,1196,1223]" pageId="15" pageNumber="78" type="female">4 ♀♀</specimenCount>
):
<materialsCitation id="3B0D3CDB2779AA236EAFFB6E6E08FAA2" ID-GBIF-Occurrence="3697349339" collectingDate="2000-01-14" collectorName="F. A. A. Lencioni" country="Brazil" location="Parque Estadual da Serra do Mar - Nucleo Caraguatatuba" municipality="Caraguatatuba" pageId="15" pageNumber="78" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
<specimenCount id="9D63FD0F2779AA236EAFFB6E6DBBFB68" box="[649,695,1196,1223]" pageId="15" pageNumber="78" type="male">1 ♂</specimenCount>
<collectingCountry id="F37276162779AA236EE7FB6E6C06FB69" box="[705,778,1196,1222]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
:
<collectingRegion id="49A1F8642779AA236F3EFB6E6C9CFB69" box="[792,912,1196,1222]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
,
<collectingMunicipality id="6BBEACFC2779AA236FB8FB6E6B4FFB69" box="[926,1091,1196,1222]" pageId="15" pageNumber="78">Caraguatatuba</collectingMunicipality>
,
<location id="8EBA605D2779AA236874FB6F6ED1FB46" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9:8EBA605D2779AA236874FB6F6ED1FB46" country="Brazil" municipality="Caraguatatuba" name="Parque Estadual da Serra do Mar - Nucleo Caraguatatuba" pageId="15" pageNumber="78" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar—Núcleo Caraguatatuba</location>
(coletas sob permissão—COTEC 608/99—SMA 42.086/99),
<date id="FFDB10462779AA2368BDFB0D6A0BFB46" box="[1179,1287,1231,1257]" pageId="15" pageNumber="78" value="2000-01-14">
<collectingDate id="EF9FE9AE2779AA2368BDFB0D6A0BFB46" box="[1179,1287,1231,1257]" pageId="15" pageNumber="78" value="2000-01-14">14.i.2000</collectingDate>
</date>
,
<collectorName id="269053502779AA236932FB126FC3FAA2" pageId="15" pageNumber="78">F.A.A. Lencioni</collectorName>
leg.
</materialsCitation>
;
<specimenCount id="9D63FD0F2779AA236D34FB316E32FAA2" box="[274,318,1267,1293]" pageId="15" pageNumber="78" type="male">1 ♂</specimenCount>
+
<materialsCitation id="3B0D3CDB2779AA236D79FB316BEDFA9F" ID-GBIF-Occurrence="3697349348" collectingDate="2000-01-15" collectorName="F. A. A. Lencioni" country="Brazil" location="Parque Estadual da Serra do Mar - Nucleo" municipality="Caraguatatuba" pageId="15" pageNumber="78" specimenCount="2" specimenCount-female="2" stateProvince="Sao Paulo">
<specimenCount id="9D63FD0F2779AA236D79FB316EACFAA2" box="[351,416,1267,1293]" pageId="15" pageNumber="78" type="female">2 ♀♀</specimenCount>
<collectingCountry id="F37276162779AA236D8FFB316EFEFAA2" box="[425,498,1267,1293]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
:
<collectingRegion id="49A1F8642779AA236DD8FB316D7FFAA2" box="[510,627,1267,1293]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
,
<collectingMunicipality id="6BBEACFC2779AA236EA6FB316C29FAA2" box="[640,805,1267,1293]" pageId="15" pageNumber="78">Caraguatatuba</collectingMunicipality>
,
<location id="8EBA605D2779AA236F17FB316A14FAA2" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9:8EBA605D2779AA236F17FB316A14FAA2" box="[817,1304,1267,1293]" country="Brazil" municipality="Caraguatatuba" name="Parque Estadual da Serra do Mar - Nucleo" pageId="15" pageNumber="78" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar—Núcleo</location>
Caraguatatuba (coletas sob permissão—COTEC 608/99—SMA 42.086/99),
<date id="FFDB10462779AA236FA6FAD46CE0FA9E" box="[896,1004,1302,1329]" pageId="15" pageNumber="78" value="2000-01-15">
<collectingDate id="EF9FE9AE2779AA236FA6FAD46CE0FA9E" box="[896,1004,1302,1329]" pageId="15" pageNumber="78" value="2000-01-15">15.i.2000</collectingDate>
</date>
,
<collectorName id="269053502779AA236FDEFAD46BA2FA9F" box="[1016,1198,1302,1328]" pageId="15" pageNumber="78">F.A.A. Lencioni</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B0D3CDB2779AA2368CBFAD46C3AFAD9" ID-GBIF-Occurrence="3697349323" collectingDate="2000-02-05" collectorName="F. A. A. Lencioni" country="Brazil" location="Parque Estadual da Serra do Mar - Nucleo Caraguatatuba" municipality="Caraguatatuba" pageId="15" pageNumber="78" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<specimenCount id="9D63FD0F2779AA2368CBFAD46A14FA9F" box="[1261,1304,1302,1328]" pageId="15" pageNumber="78" type="female">1 ♀</specimenCount>
<collectingCountry id="F37276162779AA236939FAD46A64FA9F" box="[1311,1384,1302,1328]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
:
<collectingRegion id="49A1F8642779AA236954FAD46FD6FAFC" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
,
<collectingMunicipality id="6BBEACFC2779AA236CC3FAFB6E86FAFC" box="[229,394,1337,1363]" pageId="15" pageNumber="78">Caraguatatuba</collectingMunicipality>
,
<location id="8EBA605D2779AA236DB3FAF86B13FAFC" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9:8EBA605D2779AA236DB3FAF86B13FAFC" box="[405,1055,1337,1364]" country="Brazil" municipality="Caraguatatuba" name="Parque Estadual da Serra do Mar - Nucleo Caraguatatuba" pageId="15" pageNumber="78" stateProvince="Sao Paulo">Parque Estadual da Serra do Mar—Núcleo Caraguatatuba</location>
(coletas sob permissão—COTEC 608/99—SMA 42.086/99),
<date id="FFDB10462779AA236DEAFA9E6D4DFAD8" box="[460,577,1372,1399]" pageId="15" pageNumber="78" value="2000-02-05">
<collectingDate id="EF9FE9AE2779AA236DEAFA9E6D4DFAD8" box="[460,577,1372,1399]" pageId="15" pageNumber="78" value="2000-02-05">05.ii.2000</collectingDate>
</date>
,
<collectorName id="269053502779AA236E6BFA9E6C0FFAD9" box="[589,771,1372,1398]" pageId="15" pageNumber="78">F.A.A. Lencioni</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B0D3CDB2779AA236F65FA9E6CDEFA36" ID-GBIF-Occurrence="3697349325" collectingDate="2004-01-31" collectorName="F. A. A. Lencioni" country="Brazil" latitude="-23.586945" location="Praia de Ponta Aguda" longLatPrecision="21" longitude="-45.265835" municipality="Caraguatatuba" pageId="15" pageNumber="78" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo">
<specimenCount id="9D63FD0F2779AA236F65FA9E6C61FAD8" box="[835,877,1372,1399]" pageId="15" pageNumber="78" type="male">1 ♂</specimenCount>
<collectingCountry id="F37276162779AA236F53FA9E6CB2FAD9" box="[885,958,1372,1398]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
:
<collectingRegion id="49A1F8642779AA236FEFFA9E6B31FAD9" box="[969,1085,1372,1398]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
,
<collectingMunicipality id="6BBEACFC2779AA23686FFA9E6BE2FAD9" box="[1097,1262,1372,1398]" pageId="15" pageNumber="78">Caraguatatuba</collectingMunicipality>
,
<location id="8EBA605D2779AA2368DFFA9E6FE8FA36" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9:8EBA605D2779AA2368DFFA9E6FE8FA36" country="Brazil" latitude="-23.586945" longLatPrecision="21" longitude="-45.265835" municipality="Caraguatatuba" name="Praia de Ponta Aguda" pageId="15" pageNumber="78" stateProvince="Sao Paulo">Praia de Ponta Aguda</location>
,
<geoCoordinate id="EE5150412779AA236CC9FABD6E7DFA35" box="[239,369,1407,1434]" degrees="23" direction="south" minutes="35" orientation="latitude" pageId="15" pageNumber="78" precision="15" seconds="13" value="-23.586945">23º35’13”S</geoCoordinate>
&amp;
<geoCoordinate id="EE5150412779AA236DB3FABD6D2DFA35" box="[405,545,1407,1434]" degrees="45" direction="west" minutes="15" orientation="longitude" pageId="15" pageNumber="78" precision="15" seconds="57" value="-45.265835">45º15’57”W</geoCoordinate>
, 07 m,
<date id="FFDB10462779AA236E55FABD6DD3FA36" box="[627,735,1407,1433]" pageId="15" pageNumber="78" value="2004-01-31">
<collectingDate id="EF9FE9AE2779AA236E55FABD6DD3FA36" box="[627,735,1407,1433]" pageId="15" pageNumber="78" value="2004-01-31">31.i.2004</collectingDate>
</date>
,
<collectorName id="269053502779AA236ECCFA426C93FA36" box="[746,927,1407,1434]" pageId="15" pageNumber="78">F.A.A. Lencioni</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B0D3CDB2779AA236FFBFA426B9CFA12" ID-GBIF-Occurrence="3697349326" collectingDate="2004-02-01" collectorName="F. A. A. Lencioni" country="Brazil" latitude="-23.586945" location="Praia de Ponta Aguda" longLatPrecision="21" longitude="-45.265835" municipality="Caraguatatuba" pageId="15" pageNumber="78" specimenCount="7" specimenCount-male="7" stateProvince="Sao Paulo">
<specimenCount id="9D63FD0F2779AA236FFBFA426B10FA35" box="[989,1052,1408,1434]" pageId="15" pageNumber="78" type="male">7 ♂♂</specimenCount>
<collectingCountry id="F37276162779AA236805FABD6B60FA36" box="[1059,1132,1407,1433]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
:
<collectingRegion id="49A1F8642779AA236853FA426BE5FA36" box="[1141,1257,1407,1434]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
,
<collectingMunicipality id="6BBEACFC2779AA2368D2FABD6A95FA36" box="[1268,1433,1407,1433]" pageId="15" pageNumber="78">Caraguatatuba</collectingMunicipality>
,
<location id="8EBA605D2779AA236CB1FA616E82FA12" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9:8EBA605D2779AA236CB1FA616E82FA12" box="[151,398,1443,1469]" country="Brazil" latitude="-23.586945" longLatPrecision="21" longitude="-45.265835" municipality="Caraguatatuba" name="Praia de Ponta Aguda" pageId="15" pageNumber="78" stateProvince="Sao Paulo">Praia de Ponta Aguda</location>
,
<geoCoordinate id="EE5150412779AA236DBDFA606D11FA12" box="[411,541,1442,1469]" degrees="23" direction="south" minutes="35" orientation="latitude" pageId="15" pageNumber="78" precision="15" seconds="13" value="-23.586945">23º35’13”S</geoCoordinate>
&amp;
<geoCoordinate id="EE5150412779AA236E65FA606DC3FA12" box="[579,719,1442,1469]" degrees="45" direction="west" minutes="15" orientation="longitude" pageId="15" pageNumber="78" precision="15" seconds="57" value="-45.265835">45º15’57”W</geoCoordinate>
, 07 m,
<date id="FFDB10462779AA236F03FA616C94FA12" box="[805,920,1443,1469]" pageId="15" pageNumber="78" value="2004-02-01">
<collectingDate id="EF9FE9AE2779AA236F03FA616C94FA12" box="[805,920,1443,1469]" pageId="15" pageNumber="78" value="2004-02-01">01.ii.2004</collectingDate>
</date>
,
<collectorName id="269053502779AA236F83FA616B50FA12" box="[933,1116,1443,1469]" pageId="15" pageNumber="78">F.A.A. Lencioni</collectorName>
leg.
</materialsCitation>
;
<materialsCitation id="3B0D3CDB2779AA2368BAFA616A1EFA4E" ID-GBIF-Occurrence="3697349319" collectingDate="2004-01-20" collectorName="E. A. Lencioni" country="Brazil" latitude="-23.586945" location="Praia de Ponta Aguda" longLatPrecision="21" longitude="-45.265835" municipality="Caraguatatuba" pageId="15" pageNumber="78" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<specimenCount id="9D63FD0F2779AA2368BAFA616BCBFA12" box="[1180,1223,1443,1469]" pageId="15" pageNumber="78" type="female">1 ♀</specimenCount>
<collectingCountry id="F37276162779AA2368E9FA616A14FA12" box="[1231,1304,1443,1469]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
:
<collectingRegion id="49A1F8642779AA236905FA616A94FA12" box="[1315,1432,1443,1469]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
,
<collectingMunicipality id="6BBEACFC2779AA236CB1FA056E30FA4E" box="[151,316,1479,1505]" pageId="15" pageNumber="78">Caraguatatuba</collectingMunicipality>
,
<location id="8EBA605D2779AA236D61FA056D37FA4E" LSID="urn:lsid:plazi:treatment:03CC87902779AA236CB1FE6A6C97F9A9:8EBA605D2779AA236D61FA056D37FA4E" box="[327,571,1479,1505]" country="Brazil" latitude="-23.586945" longLatPrecision="21" longitude="-45.265835" municipality="Caraguatatuba" name="Praia de Ponta Aguda" pageId="15" pageNumber="78" stateProvince="Sao Paulo">Praia de Ponta Aguda</location>
,
<geoCoordinate id="EE5150412779AA236E61FA046DC4FA4E" box="[583,712,1478,1505]" degrees="23" direction="south" minutes="35" orientation="latitude" pageId="15" pageNumber="78" precision="15" seconds="13" value="-23.586945">23º35’13”S</geoCoordinate>
&amp;
<geoCoordinate id="EE5150412779AA236ECBFA046C75FA4E" box="[749,889,1478,1505]" degrees="45" direction="west" minutes="15" orientation="longitude" pageId="15" pageNumber="78" precision="15" seconds="57" value="-45.265835">45º15’57”W</geoCoordinate>
, 07 m,
<date id="FFDB10462779AA236FEAFA056B34FA4E" box="[972,1080,1479,1505]" pageId="15" pageNumber="78" value="2004-01-20">
<collectingDate id="EF9FE9AE2779AA236FEAFA056B34FA4E" box="[972,1080,1479,1505]" pageId="15" pageNumber="78" value="2004-01-20">20.i.2004</collectingDate>
</date>
,
<collectorName id="269053502779AA236865FA056BEEFA4E" box="[1091,1250,1479,1505]" pageId="15" pageNumber="78">E.A. Lencioni</collectorName>
leg.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37F650D2779AA236CE1FA286C97F9A9" box="[199,923,1514,1542]" pageId="15" pageNumber="78" type="distribution">
<paragraph id="8BDA36862779AA236CE1FA286C97F9A9" blockId="15.[151,1437,527,1542]" box="[199,923,1514,1542]" pageId="15" pageNumber="78">
<emphasis id="B911EA942779AA236CE1FA286E6DF9AB" bold="true" box="[199,353,1514,1540]" pageId="15" pageNumber="78">Distribution.</emphasis>
<collectingCountry id="F37276162779AA236D41FA296EA7F9AA" box="[359,427,1515,1541]" name="Brazil" pageId="15" pageNumber="78">Brazil</collectingCountry>
(
<collectingRegion id="49A1F8642779AA236D9CFA296D53F9AA" box="[442,607,1515,1541]" country="Brazil" name="Espirito Santo" pageId="15" pageNumber="78">Espírito Santo</collectingRegion>
,
<collectingRegion id="49A1F8642779AA236E4CFA296C03F9AA" box="[618,783,1515,1541]" country="Brazil" name="Rio de Janeiro" pageId="15" pageNumber="78">Rio de Janeiro</collectingRegion>
,
<collectingRegion id="49A1F8642779AA236F3FFA296C83F9A9" box="[793,911,1515,1542]" country="Brazil" name="Sao Paulo" pageId="15" pageNumber="78">São Paulo</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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