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<document id="2B4C041E07D61C54B4D146FA172DB85A" ID-CLB-Dataset="31145" ID-DOI="10.11646/zootaxa.4365.2.5" ID-GBIF-Dataset="f910478a-92a3-4de6-8d96-d8d37ce28163" ID-ISSN="1175-5326" ID-Zenodo-Dep="1117608" ID-ZooBank="D9EA4014-FE5B-4F23-86F8-9E2853A312C2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1513583912014" checkinUser="plazi" docAuthor="Dolibaina, Diego Rodrigo, Mielke, Olaf Hermann Hendrik &amp; Casagrande, Mirna Martins" docDate="2017" docId="03CC878EFFC5FFADFF4EFA89646311B7" docLanguage="en" docName="zootaxa.4365.2.5.pdf" docOrigin="Zootaxa 4365 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rufocumbre emeryi Dolibaina &amp; Mielke &amp; Casagrande 2017, sp. nov." docType="treatment" docUuid="68418560-89E1-46EC-BA3C-BF75E39FC28B" docUuidSource="ZooBank" docVersion="8" lastPageNumber="204" masterDocId="FFF5FFF6FFC3FFA4FFD9FFCD60661333" masterDocTitle="Taxonomy of Rufocumbre gen. nov., a new Moncini skipper genus (Lepidoptera: Hesperiidae: Hesperiinae)" masterLastPageNumber="216" masterPageNumber="196" pageNumber="202" updateTime="1698510150032" updateUser="ExternalLinkService">
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<mods:title id="3BFD9285A1EF373D5B2DE8F3EA98883A">Taxonomy of Rufocumbre gen. nov., a new Moncini skipper genus (Lepidoptera: Hesperiidae: Hesperiinae)</mods:title>
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<mods:namePart id="DD2D80389BE7C06119FB93D12347A534">Dolibaina, Diego Rodrigo</mods:namePart>
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<mods:namePart id="2D5C562700611E1796807F5CEBCD092F">Mielke, Olaf Hermann Hendrik</mods:namePart>
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<mods:namePart id="011A405726A45D8C0B37EC838B28B755">Casagrande, Mirna Martins</mods:namePart>
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<mods:title id="C1A2EB9D956B272239CD641C39C7D4A5">Zootaxa</mods:title>
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<mods:date id="D661287DAA2A47BC36BF1FE5EDD9FE5F">2017</mods:date>
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<mods:number id="D0C44C4721E28AE5E3AD05AEF61D8987">2017-12-18</mods:number>
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<mods:number id="F863DBC0E67F8A65FCC01BF9CF46CB5F">4365</mods:number>
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<treatment id="03CC878EFFC5FFADFF4EFA89646311B7" ID-DOI="http://doi.org/10.5281/zenodo.6032053" ID-GBIF-Taxon="139334130" ID-Zenodo-Dep="6032053" LSID="urn:lsid:zoobank.org:act:68418560-89E1-46EC-BA3C-BF75E39FC28B" httpUri="http://treatment.plazi.org/id/03CC878EFFC5FFADFF4EFA89646311B7" lastPageId="9" lastPageNumber="204" pageId="6" pageNumber="202">
<subSubSection id="C37F6513FFC5FFA2FF4EFA8961B71692" pageId="6" pageNumber="202" type="nomenclature">
<paragraph id="8BDA3698FFC5FFA2FF4EFA89618A166D" blockId="6.[151,963,1348,1441]" box="[151,492,1348,1374]" pageId="6" pageNumber="202">
<heading id="D09281F4FFC5FFA2FF4EFA89618A166D" bold="true" box="[151,492,1348,1374]" fontSize="11" level="1" pageId="6" pageNumber="202" reason="1">
<emphasis id="B911EA8AFFC5FFA2FF4EFA89618A166D" bold="true" box="[151,492,1348,1374]" pageId="6" pageNumber="202">
<taxonomicName id="4C654D1BFFC5FFA2FF4EFA8961E1166D" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[151,391,1348,1374]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="202" phylum="Arthropoda" rank="species" species="emeryi" status="sp. nov.">
<emphasis id="B911EA8AFFC5FFA2FF4EFA8961E1166D" bold="true" box="[151,391,1348,1374]" italics="true" pageId="6" pageNumber="202">Rufocumbre emeryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC5FFA2FE57FA89618A166D" box="[398,492,1348,1374]" pageId="6" pageNumber="202" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDA3698FFC5FFA2FF4EFAAB61B71692" blockId="6.[151,963,1348,1441]" pageId="6" pageNumber="202">
urn:lsid:zoobank.org:act:68418560-89E1-46EC-BA3C-BF75E39FC28B
<figureCitation id="135E2A1DFFC5FFA2FF4EFA45609C1692" box="[151,250,1416,1441]" captionStart="FIGURES 1–20" captionStartId="3.[151,264,1502,1524]" captionTargetBox="[151,1435,196,1478]" captionTargetId="figure@3.[150,1436,195,1478]" captionTargetPageId="3" captionText="FIGURES 1–20. Species of Rufocumbre in dorsal and ventral views, male (left) and female (right). 1–4, R. celioi sp. nov., male holotype and female allotype. 5–8, R. emeryi sp. nov., male holotype and female allotype. 9–12, R. eberti stat. nov., comb. nov., male (OM 59.360) and female (OM 19.333) from Caraça, Catas Altas, Minas Gerais, Brazil. 13–16, R. schneideri sp. nov., male holotype and female allotype. 17–20, R. lucasi sp. nov., male holotype and female allotype.Scale bar = 1 cm." httpUri="https://zenodo.org/record/1117610/files/figure.png" pageId="6" pageNumber="202">Figs 5–8</figureCitation>
,
<figureCitation id="135E2A1DFFC5FFA2FEDFFA4561421693" box="[262,292,1416,1440]" captionStart="FIGURES 21–25" captionStartId="5.[151,264,1230,1252]" captionTargetBox="[156,1436,195,1206]" captionTargetId="figure@5.[151,1436,195,1206]" captionTargetPageId="5" captionText="FIGURES 21–25. Male brand in dorsal forewing of species of Rufocumbre. 21, R. celioi sp. nov., paratype from Boa Vista, Tibagi, Paraná, Brazil (OM 52.771). 22, R. emeryi sp. nov., paratype from Córrego Grotão, Planaltina, Distrito Federal, Brazil (DZ 8.676). 23, R. eberti stat. nov., comb. nov., from Ouro Preto, Minas Gerais, Brazil (OM 65.904). 24, R. schneideri sp. nov., paratype from RPPN Ninho do Corvo, Prudentópolis, Paraná, Brazil (DD 207). 25, R. lucasi sp. nov., paratype from Morro do Pai Inácio, Palmeiras, Bahia, Brazil (OM 50.016).Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117612/files/figure.png" pageId="6" pageNumber="202">22</figureCitation>
,
<figureCitation id="135E2A1DFFC5FFA2FEE9FA4561281693" box="[304,334,1416,1440]" captionStart="FIGURES 33–34" captionStartId="8.[151,264,941,963]" captionTargetBox="[151,1436,193,920]" captionTargetId="figure@8.[151,1436,193,920]" captionTargetPageId="8" captionText="FIGURES 33–34. Male genitalia of species of Rufocumbre.Left to right: dorsal view of tegumen, uncus and gnathos; ventral view of tegumen, uncus and gnathos; lateral view of tegumen, saccus, uncus and gnathos; left view of the right valva; posterior and lateral views of fultura inferior; lateral left, dorsal, ventral and lateral right views of the aedeagus. 33, R. celioi sp. nov., paratype from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 15.577). 34, R. emeryi sp. nov., holotype.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117616/files/figure.png" pageId="6" pageNumber="202">34</figureCitation>
,
<figureCitation id="135E2A1DFFC5FFA2FE83FA45611F1693" box="[346,377,1416,1440]" captionStart="FIGURES 38–39" captionStartId="10.[151,264,1936,1958]" captionTargetBox="[265,1342,205,1919]" captionTargetId="figure@10.[215,1379,195,1919]" captionTargetPageId="10" captionText="FIGURES 38–39. Female genitalia of species of Rufocumbre in ventral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix. 38, R. celioi sp. nov., paratype from Bituva, Mafra, Santa Catarina, Brazil (DZ 17.124). 39, R. emeryi sp. nov., paratype from Fazenda Água Limpa, Brasília, Distrito Federal, Brazil (DZ 20.891). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117620/files/figure.png" pageId="6" pageNumber="202">39</figureCitation>
,
<figureCitation id="135E2A1DFFC5FFA2FE5DFA4561C51693" box="[388,419,1416,1440]" captionStart="FIGURES 43–47" captionStartId="12.[151,264,1273,1295]" captionTargetBox="[163,1431,210,1244]" captionTargetId="figure@12.[151,1436,193,1252]" captionTargetPageId="12" captionText="FIGURES 43–47. Female genitalia of species of Rufocumbre in lateral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix. 43, R. celioi sp. nov., paratype from Bituva, Mafra, Santa Catarina, Brazil (DZ 17.124). 44, R. emeryi sp. nov., paratype from Fazenda Água Limpa, Brasília, Distrito Federal, Brazil (DZ 20.891). 45, R. eberti stat. nov., comb. nov., from Ouro Preto, Minas Gerais, Brazil (OM 66.016). 46, R. schneideri sp. nov., paratype from Serra Grande, Conceição dos Ouros, Minas Gerais, Brazil (OM 59.073). 47, R. lucasi sp. nov., paratype from Morro do Pai Inácio, Palmeiras, Bahia, Brazil (OM 50.179).Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117624/files/figure.png" pageId="6" pageNumber="202">44</figureCitation>
,
<figureCitation id="135E2A1DFFC5FFA2FE77FA4561AB1692" box="[430,461,1416,1441]" captionStart="FIGURE 55" captionStartId="17.[151,250,1317,1339]" captionTargetBox="[151,1436,195,1293]" captionTargetId="figure@17.[151,1436,195,1293]" captionTargetPageId="17" captionText="FIGURE 55.Geographical distribution of Rufocumbre emeryi sp. nov. and R. eberti stat. nov., comb. nov." httpUri="https://zenodo.org/record/1117630/files/figure.png" pageId="6" pageNumber="202">55</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C37F6513FFC5FFA2FF4EFA1D61891536" pageId="6" pageNumber="202" type="reference_group">
<paragraph id="8BDA3698FFC5FFA2FF4EFA1D61891536" blockId="6.[151,1437,1488,1541]" pageId="6" pageNumber="202">
<taxonomicName id="4C654D1BFFC5FFA2FF4EFA1D613D16D5" ID-CoL="5GQNV" box="[151,347,1488,1510]" class="Insecta" family="Hesperiidae" genus="Cumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="202" phylum="Arthropoda" rank="subSpecies" species="belli" subSpecies="eberti">
<emphasis id="B911EA8AFFC5FFA2FF4EFA1D613D16D5" box="[151,347,1488,1510]" italics="true" pageId="6" pageNumber="202">Cumbre belli eberti</emphasis>
</taxonomicName>
<emphasis id="B911EA8AFFC5FFA2FEB8FA1D624116D5" bold="true" box="[353,551,1488,1510]" pageId="6" pageNumber="202">[misidentification]</emphasis>
;
<bibRefCitation id="EFF44B69FFC5FFA2FDEDFA1D634B16D5" author="Pinheiro, C. E. G. &amp; Emery, E. O." box="[564,813,1488,1510]" journalOrPublisher="Biota Neotropica" pageId="6" pageNumber="202" pagination="1 - 15" part="6" publicationUrl="https://doi.org/10.1590/S1676-06032006000300012" refString="Pinheiro, C. E. G. &amp; Emery, E. O. (2006) As borboletas (Lepidoptera: Papilionoidea e Hesperioidea) da Area de Protecao Ambiental do Gama e Cabeca de Veado, Distrito Federal, Brasil. Biota Neotropica, 6 (3), 1 - 15. https: // doi. org / 10.1590 / S 1676 - 06032006000300012" title="As borboletas (Lepidoptera: Papilionoidea e Hesperioidea) da Area de Protecao Ambiental do Gama e Cabeca de Veado, Distrito Federal, Brasil" type="journal article" year="2006">Pinheiro &amp; Emery, 2006</bibRefCitation>
.
<emphasis id="B911EA8AFFC5FFA2FCECFA1D639816D5" bold="true" box="[821,1022,1488,1510]" pageId="6" pageNumber="202">
<taxonomicName id="4C654D1BFFC5FFA2FCECFA1D638A16D5" authority="Neotropica" authorityName="Neotropica" box="[821,1004,1488,1510]" class="Pinopsida" family="Cupressaceae" genus="Biota" higherTaxonomySource="GBIF" kingdom="Plantae" order="Pinales" pageId="6" pageNumber="202" phylum="Tracheophyta" rank="genus">Biota Neotropica</taxonomicName>
6
</emphasis>
(3): 6.-
<bibRefCitation id="EFF44B69FFC5FFA2FB93FA1D65FF16D5" author="Mielke, O. H. H. &amp; Emery, E. O. &amp; Pinheiro, C. E. G." box="[1098,1433,1488,1510]" journalOrPublisher="Revista Brasileira de Zoologia" pageId="6" pageNumber="202" pagination="283 - 288" part="52" refString="Mielke, O. H. H., Emery, E. O. &amp; Pinheiro, C. E. G. (2008) As borboletas Hesperiidae (Lepidoptera, Hesperioidea) do Distrito Federal, Brasil. Revista Brasileira de Zoologia, 52 (2), 283 - 288." title="As borboletas Hesperiidae (Lepidoptera, Hesperioidea) do Distrito Federal, Brasil" type="journal article" year="2008">Mielke; Emery &amp; Pinheiro, 2008</bibRefCitation>
.
<emphasis id="B911EA8AFFC5FFA2FF1EFA2261F01536" bold="true" box="[199,406,1519,1541]" pageId="6" pageNumber="202">Revta bras. Ent. 52</emphasis>
(2): 286.
</paragraph>
</subSubSection>
<subSubSection id="C37F6513FFC5FFA2FF4EF9FC65FE1587" pageId="6" pageNumber="202" type="diagnosis">
<paragraph id="8BDA3698FFC5FFA2FF4EF9FC65FE1587" blockId="6.[151,1436,1585,2036]" pageId="6" pageNumber="202">
<emphasis id="B911EA8AFFC5FFA2FF4EF9FC61681579" bold="true" box="[151,270,1585,1610]" pageId="6" pageNumber="202">Diagnosis.</emphasis>
<taxonomicName id="4C654D1BFFC5FFA2FECFF9FC6189157A" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[278,495,1585,1610]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="202" phylum="Arthropoda" rank="species" species="emeryi" status="sp. nov.">
<emphasis id="B911EA8AFFC5FFA2FECFF9FC6189157A" box="[278,495,1585,1610]" italics="true" pageId="6" pageNumber="202">Rufocumbre emeryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC5FFA2FE2EF9FC62361579" box="[503,592,1585,1610]" pageId="6" pageNumber="202" rank="species">
<emphasis id="B911EA8AFFC5FFA2FE2EF9FC62361579" bold="true" box="[503,592,1585,1610]" pageId="6" pageNumber="202">sp. nov.</emphasis>
</taxonomicNameLabel>
is distinguished from all species of
<taxonomicName id="4C654D1BFFC5FFA2FC3DF9FC640F1579" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[996,1129,1585,1610]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="213" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B911EA8AFFC5FFA2FC3DF9FC640F1579" box="[996,1129,1585,1610]" italics="true" pageId="6" pageNumber="202">Rufocumbre</emphasis>
</taxonomicName>
by the uncus slightly bifid, with pointed arms; ampulla wider than base of valva, with the proximal margin pointed; distal spine of harpe shorter, not exceeding the distal margin of the ampulla (
<figureCitation id="135E2A1DFFC5FFA2FD7FF9B5629F15A2" box="[678,761,1656,1681]" captionStart="FIGURES 33–34" captionStartId="8.[151,264,941,963]" captionTargetBox="[151,1436,193,920]" captionTargetId="figure@8.[151,1436,193,920]" captionTargetPageId="8" captionText="FIGURES 33–34. Male genitalia of species of Rufocumbre.Left to right: dorsal view of tegumen, uncus and gnathos; ventral view of tegumen, uncus and gnathos; lateral view of tegumen, saccus, uncus and gnathos; left view of the right valva; posterior and lateral views of fultura inferior; lateral left, dorsal, ventral and lateral right views of the aedeagus. 33, R. celioi sp. nov., paratype from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 15.577). 34, R. emeryi sp. nov., holotype.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117616/files/figure.png" pageId="6" pageNumber="202">Fig. 34</figureCitation>
); lamella antevaginalis absent (as in
<taxonomicName id="4C654D1BFFC5FFA2FB57F9B4648F15A2" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[1166,1257,1656,1681]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="6" pageNumber="202" phylum="Arthropoda" rank="species" species="celioi" status="sp. nov.">
<emphasis id="B911EA8AFFC5FFA2FB57F9B4648F15A2" box="[1166,1257,1656,1681]" italics="true" pageId="6" pageNumber="202">R. celioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC5FFA2FB28F9B5652F15A2" box="[1265,1353,1656,1681]" pageId="6" pageNumber="202" rank="species">
<emphasis id="B911EA8AFFC5FFA2FB28F9B5652F15A2" bold="true" box="[1265,1353,1656,1681]" pageId="6" pageNumber="202">sp. nov.</emphasis>
</taxonomicNameLabel>
); distal margin of the lamella postvaginalis with two long projections separated from each other by a short distance (
<figureCitation id="135E2A1DFFC5FFA2FAE3F95165ED1587" box="[1338,1419,1692,1717]" captionStart="FIGURES 38–39" captionStartId="10.[151,264,1936,1958]" captionTargetBox="[265,1342,205,1919]" captionTargetId="figure@10.[215,1379,195,1919]" captionTargetPageId="10" captionText="FIGURES 38–39. Female genitalia of species of Rufocumbre in ventral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix. 38, R. celioi sp. nov., paratype from Bituva, Mafra, Santa Catarina, Brazil (DZ 17.124). 39, R. emeryi sp. nov., paratype from Fazenda Água Limpa, Brasília, Distrito Federal, Brazil (DZ 20.891). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117620/files/figure.png" pageId="6" pageNumber="202">Fig. 39</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C37F6513FFC5FFACFF1EF972615F17E2" lastPageId="8" lastPageNumber="204" pageId="6" pageNumber="202" type="description">
<paragraph id="8BDA3698FFC5FFA2FF1EF972639815E4" blockId="6.[151,1436,1585,2036]" box="[199,1022,1727,1752]" pageId="6" pageNumber="202">
<emphasis id="B911EA8AFFC5FFA2FF1EF972613F15EB" bold="true" box="[199,345,1727,1752]" pageId="6" pageNumber="202">Description.</emphasis>
<emphasis id="B911EA8AFFC5FFA2FE86F90D627E15EB" box="[351,536,1727,1752]" italics="true" pageId="6" pageNumber="202">Forewing length</emphasis>
: Male 13.9-17.5 mm, female 15.2-17 mm.
</paragraph>
<paragraph id="8BDA3698FFC5FFA2FF1EF929639615CF" blockId="6.[151,1436,1585,2036]" box="[199,1008,1763,1788]" pageId="6" pageNumber="202">
<emphasis id="B911EA8AFFC5FFA2FF1EF92961E415CF" box="[199,386,1763,1788]" italics="true" pageId="6" pageNumber="202">Forewing dorsal</emphasis>
(
<figureCitation id="135E2A1DFFC5FFA2FE49F92E619215C8" box="[400,500,1763,1788]" captionStart="FIGURES 1–20" captionStartId="3.[151,264,1502,1524]" captionTargetBox="[151,1435,196,1478]" captionTargetId="figure@3.[150,1436,195,1478]" captionTargetPageId="3" captionText="FIGURES 1–20. Species of Rufocumbre in dorsal and ventral views, male (left) and female (right). 1–4, R. celioi sp. nov., male holotype and female allotype. 5–8, R. emeryi sp. nov., male holotype and female allotype. 9–12, R. eberti stat. nov., comb. nov., male (OM 59.360) and female (OM 19.333) from Caraça, Catas Altas, Minas Gerais, Brazil. 13–16, R. schneideri sp. nov., male holotype and female allotype. 17–20, R. lucasi sp. nov., male holotype and female allotype.Scale bar = 1 cm." httpUri="https://zenodo.org/record/1117610/files/figure.png" pageId="6" pageNumber="202">Figs 5, 7</figureCitation>
,
<figureCitation id="135E2A1DFFC5FFA2FDD9F92E624615C8" box="[512,544,1763,1787]" captionStart="FIGURES 21–25" captionStartId="5.[151,264,1230,1252]" captionTargetBox="[156,1436,195,1206]" captionTargetId="figure@5.[151,1436,195,1206]" captionTargetPageId="5" captionText="FIGURES 21–25. Male brand in dorsal forewing of species of Rufocumbre. 21, R. celioi sp. nov., paratype from Boa Vista, Tibagi, Paraná, Brazil (OM 52.771). 22, R. emeryi sp. nov., paratype from Córrego Grotão, Planaltina, Distrito Federal, Brazil (DZ 8.676). 23, R. eberti stat. nov., comb. nov., from Ouro Preto, Minas Gerais, Brazil (OM 65.904). 24, R. schneideri sp. nov., paratype from RPPN Ninho do Corvo, Prudentópolis, Paraná, Brazil (DD 207). 25, R. lucasi sp. nov., paratype from Morro do Pai Inácio, Palmeiras, Bahia, Brazil (OM 50.016).Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117612/files/figure.png" pageId="6" pageNumber="202">22</figureCitation>
): brand with the inferior projection short.
</paragraph>
<paragraph id="8BDA3698FFC5FFA2FF1EF8CB65221471" blockId="6.[151,1436,1585,2036]" pageId="6" pageNumber="202">
<emphasis id="B911EA8AFFC5FFA2FF1EF8CB61F1142C" box="[199,407,1798,1823]" italics="true" pageId="6" pageNumber="202">Hind wing ventral</emphasis>
(
<figureCitation id="135E2A1DFFC5FFA2FE70F8CB6274142C" box="[425,530,1798,1823]" captionStart="FIGURES 1–20" captionStartId="3.[151,264,1502,1524]" captionTargetBox="[151,1435,196,1478]" captionTargetId="figure@3.[150,1436,195,1478]" captionTargetPageId="3" captionText="FIGURES 1–20. Species of Rufocumbre in dorsal and ventral views, male (left) and female (right). 1–4, R. celioi sp. nov., male holotype and female allotype. 5–8, R. emeryi sp. nov., male holotype and female allotype. 9–12, R. eberti stat. nov., comb. nov., male (OM 59.360) and female (OM 19.333) from Caraça, Catas Altas, Minas Gerais, Brazil. 13–16, R. schneideri sp. nov., male holotype and female allotype. 17–20, R. lucasi sp. nov., male holotype and female allotype.Scale bar = 1 cm." httpUri="https://zenodo.org/record/1117610/files/figure.png" pageId="6" pageNumber="202">Figs 6, 8</figureCitation>
): basal and postdiscal bands light orange, without gray to purple scales on the basal band; discal band separated from submarginal band by a short distance, with its distal margin angled.
</paragraph>
<paragraph id="8BDA3698FFC5FFA2FF1EF880629A14C7" blockId="6.[151,1436,1585,2036]" pageId="6" pageNumber="202">
<emphasis id="B911EA8AFFC5FFA2FF1EF88061091455" box="[199,367,1869,1894]" italics="true" pageId="6" pageNumber="202">Male genitalia</emphasis>
(
<figureCitation id="135E2A1DFFC5FFA2FE58F88061BE1455" box="[385,472,1869,1894]" captionStart="FIGURES 33–34" captionStartId="8.[151,264,941,963]" captionTargetBox="[151,1436,193,920]" captionTargetId="figure@8.[151,1436,193,920]" captionTargetPageId="8" captionText="FIGURES 33–34. Male genitalia of species of Rufocumbre.Left to right: dorsal view of tegumen, uncus and gnathos; ventral view of tegumen, uncus and gnathos; lateral view of tegumen, saccus, uncus and gnathos; left view of the right valva; posterior and lateral views of fultura inferior; lateral left, dorsal, ventral and lateral right views of the aedeagus. 33, R. celioi sp. nov., paratype from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 15.577). 34, R. emeryi sp. nov., holotype.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117616/files/figure.png" pageId="6" pageNumber="202">Fig. 34</figureCitation>
): Uncus slightly bifid, with short and pointed arms; sacculus with about 1/5 of the width of valva, proximal margin close to the base of valva; ampulla wider than base of valva, proximal margin pointed; distal spine of harpe short, not exceeding the distal margin of ampulla; aedeagus about 2.3 times longer than the combined length of tegumen+uncus; aedeagus distal margin bifid, with divergent projections; cornutus as a small flattened plate with a short spine at the middle.
</paragraph>
<caption id="DF1A6610FFC4FFA3FF4EF9736137145E" httpUri="https://zenodo.org/record/1117614/files/figure.png" pageId="7" pageNumber="203" targetBox="[152,1436,196,1678]" targetPageId="7">
<paragraph id="8BDA3698FFC4FFA3FF4EF9736137145E" blockId="7.[151,1436,1726,1901]" pageId="7" pageNumber="203">
<emphasis id="B911EA8AFFC4FFA3FF4EF973613015E7" bold="true" box="[151,342,1726,1748]" pageId="7" pageNumber="203">FIGURES 26–32.</emphasis>
Variation of male brand in dorsal forewing of
<taxonomicName id="4C654D1BFFC4FFA3FCE4F973639E15E7" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[829,1016,1726,1748]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="203" phylum="Arthropoda" rank="species" species="celioi" status="sp. nov.">
<emphasis id="B911EA8AFFC4FFA3FCE4F973639E15E7" box="[829,1016,1726,1748]" italics="true" pageId="7" pageNumber="203">Rufocumbre celioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC4FFA3FBD8F973643515E7" box="[1025,1107,1726,1748]" pageId="7" pageNumber="203" rank="species">
<emphasis id="B911EA8AFFC4FFA3FBD8F973643515E7" bold="true" box="[1025,1107,1726,1748]" pageId="7" pageNumber="203">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C654D1BFFC4FFA3FB50F972648715E7" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1161,1249,1726,1748]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="203" phylum="Arthropoda" rank="species" species="eberti" status="stat. nov.">
<emphasis id="B911EA8AFFC4FFA3FB50F972648715E7" box="[1161,1249,1726,1748]" italics="true" pageId="7" pageNumber="203">R. eberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC4FFA3FB33F97360A515C1" pageId="7" pageNumber="203">
<emphasis id="B911EA8AFFC4FFA3FB33F97360A515C1" bold="true" pageId="7" pageNumber="203">stat. nov., comb. nov.</emphasis>
</taxonomicNameLabel>
26–29,
<taxonomicName id="4C654D1BFFC4FFA3FECCF913610C15C1" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[277,362,1756,1779]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="203" phylum="Arthropoda" rank="species" species="celioi" status="sp. nov.">
<emphasis id="B911EA8AFFC4FFA3FECCF913610C15C1" box="[277,362,1756,1779]" italics="true" pageId="7" pageNumber="203">R. celioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC4FFA3FEA9F91161A715C1" box="[368,449,1756,1778]" pageId="7" pageNumber="203" rank="species">
<emphasis id="B911EA8AFFC4FFA3FEA9F91161A715C1" bold="true" box="[368,449,1756,1778]" pageId="7" pageNumber="203">sp. nov.</emphasis>
</taxonomicNameLabel>
26, from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 17.077); 27, from Pico Paraná, Campina Grande do Sul, Paraná, Brazil (OM 38.041); 28, from Morro da Igreja, Urubici, Santa Catarina, Brazil (OM 48.253); 29, from Pelotas, Rio Grande do Sul, Brazil (DZ 939). 30–32,
<taxonomicName id="4C654D1BFFC4FFA3FD76F8D663601403" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[687,774,1818,1840]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="203" phylum="Arthropoda" rank="species" species="eberti" status="stat. nov.">
<emphasis id="B911EA8AFFC4FFA3FD76F8D663601403" box="[687,774,1818,1840]" italics="true" pageId="7" pageNumber="203">R. eberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFC4FFA3FCD5F8D7638D1403" box="[780,1003,1818,1840]" pageId="7" pageNumber="203">
<emphasis id="B911EA8AFFC4FFA3FCD5F8D7638D1403" bold="true" box="[780,1003,1818,1840]" pageId="7" pageNumber="203">stat. nov., comb. nov.</emphasis>
</taxonomicNameLabel>
30, from Ouro Preto, Minas Gerais, Brazil (OM 65.904); 31, from Morro do Chapéu, Bahia, Brazil (OM50.780); 32, Delfim Moreira, Minas Gerais, Brazil (DZ 15.729). Scale bar = 1 mm.
</paragraph>
</caption>
<caption id="DF1A6610FFCBFFACFF4EFC60650B1713" httpUri="https://zenodo.org/record/1117616/files/figure.png" pageId="8" pageNumber="204" targetBox="[151,1436,193,920]" targetPageId="8">
<paragraph id="8BDA3698FFCBFFACFF4EFC60650B1713" blockId="8.[151,1436,941,1056]" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFF4EFC60613210F0" bold="true" box="[151,340,941,963]" pageId="8" pageNumber="204">FIGURES 33–34.</emphasis>
Male genitalia of species of
<emphasis id="B911EA8AFFCBFFACFDA5FC60629910F0" box="[636,767,941,963]" italics="true" pageId="8" pageNumber="204">
<taxonomicName id="4C654D1BFFCBFFACFDA5FC60629C10F0" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[636,762,941,963]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="213" phylum="Arthropoda" rank="genus" status="gen. nov.">Rufocumbre</taxonomicName>
.
</emphasis>
Left to right: dorsal view of tegumen, uncus and gnathos; ventral view of tegumen, uncus and gnathos; lateral view of tegumen, saccus, uncus and gnathos; left view of the right valva; posterior and lateral views of fultura inferior; lateral left, dorsal, ventral and lateral right views of the aedeagus. 33,
<taxonomicName id="4C654D1BFFCBFFACFB3CFC21655D1732" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[1253,1339,1003,1025]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="celioi" status="sp. nov.">
<emphasis id="B911EA8AFFCBFFACFB3CFC21655D1732" box="[1253,1339,1003,1025]" italics="true" pageId="8" pageNumber="204">R. celioi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFCBFFACFA9AFC2665F31732" box="[1347,1429,1003,1025]" pageId="8" pageNumber="204" rank="species">
<emphasis id="B911EA8AFFCBFFACFA9AFC2665F31732" bold="true" box="[1347,1429,1003,1025]" pageId="8" pageNumber="204">sp. nov.</emphasis>
</taxonomicNameLabel>
, paratype from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 15.577). 34,
<taxonomicName id="4C654D1BFFCBFFACFC50FBC6638C1713" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[905,1002,1035,1056]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="emeryi" status="sp. nov.">
<emphasis id="B911EA8AFFCBFFACFC50FBC6638C1713" box="[905,1002,1035,1056]" italics="true" pageId="8" pageNumber="204">R. emeryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFCBFFACFC29FBC764271713" box="[1008,1089,1034,1056]" pageId="8" pageNumber="204" rank="species">
<emphasis id="B911EA8AFFCBFFACFC29FBC764271713" bold="true" box="[1008,1089,1034,1056]" pageId="8" pageNumber="204">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. Scale bar = 1 mm.
</paragraph>
</caption>
<paragraph id="8BDA3698FFCBFFACFF1EFB81615F17E2" blockId="8.[151,1436,1100,2025]" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFF1EFB8161E11756" box="[199,391,1100,1125]" italics="true" pageId="8" pageNumber="204">Female genitalia</emphasis>
(
<figureCitation id="135E2A1DFFCBFFACFE4EFB8161961756" box="[407,496,1100,1125]" captionStart="FIGURES 38–39" captionStartId="10.[151,264,1936,1958]" captionTargetBox="[265,1342,205,1919]" captionTargetId="figure@10.[215,1379,195,1919]" captionTargetPageId="10" captionText="FIGURES 38–39. Female genitalia of species of Rufocumbre in ventral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix. 38, R. celioi sp. nov., paratype from Bituva, Mafra, Santa Catarina, Brazil (DZ 17.124). 39, R. emeryi sp. nov., paratype from Fazenda Água Limpa, Brasília, Distrito Federal, Brazil (DZ 20.891). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117620/files/figure.png" pageId="8" pageNumber="204">Figs 39</figureCitation>
,
<figureCitation id="135E2A1DFFCBFFACFE24FB8062781756" box="[509,542,1101,1125]" captionStart="FIGURES 43–47" captionStartId="12.[151,264,1273,1295]" captionTargetBox="[163,1431,210,1244]" captionTargetId="figure@12.[151,1436,193,1252]" captionTargetPageId="12" captionText="FIGURES 43–47. Female genitalia of species of Rufocumbre in lateral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix. 43, R. celioi sp. nov., paratype from Bituva, Mafra, Santa Catarina, Brazil (DZ 17.124). 44, R. emeryi sp. nov., paratype from Fazenda Água Limpa, Brasília, Distrito Federal, Brazil (DZ 20.891). 45, R. eberti stat. nov., comb. nov., from Ouro Preto, Minas Gerais, Brazil (OM 66.016). 46, R. schneideri sp. nov., paratype from Serra Grande, Conceição dos Ouros, Minas Gerais, Brazil (OM 59.073). 47, R. lucasi sp. nov., paratype from Morro do Pai Inácio, Palmeiras, Bahia, Brazil (OM 50.179).Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117624/files/figure.png" pageId="8" pageNumber="204">44</figureCitation>
): Lamella antevaginalis absent; distal margin of the lamella postvaginalis with two long projections separated from each other by a short distance; ductus bursae with an irregular sclerotized ring near the insertion with corpus bursae; corpus bursae with two weak lateral lines of signa, shorter than half of the corpus bursae.
</paragraph>
</subSubSection>
<subSubSection id="C37F6513FFCBFFACFF1EFB116365150A" pageId="8" pageNumber="204" type="discussion">
<paragraph id="8BDA3698FFCBFFACFF1EFB1164BF16FE" blockId="8.[151,1436,1100,2025]" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFF1EFB11612917C6" bold="true" box="[199,335,1244,1269]" pageId="8" pageNumber="204">Comments.</emphasis>
<taxonomicName id="4C654D1BFFCBFFACFE85FB11625A17C7" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[348,572,1244,1269]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="emeryi" status="sp. nov.">
<emphasis id="B911EA8AFFCBFFACFE85FB11625A17C7" box="[348,572,1244,1269]" italics="true" pageId="8" pageNumber="204">Rufocumbre emeryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFCBFFACFD90FB1162CE17C6" box="[585,680,1244,1269]" pageId="8" pageNumber="204" rank="species">
<emphasis id="B911EA8AFFCBFFACFD90FB1162CE17C6" bold="true" box="[585,680,1244,1269]" pageId="8" pageNumber="204">sp. nov.</emphasis>
</taxonomicNameLabel>
was erroneously identified as
<taxonomicName id="4C654D1BFFCBFFACFBC3FB1064D917C6" box="[1050,1215,1244,1269]" class="Insecta" family="Hesperiidae" genus="Cumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="subSpecies" species="belli" subSpecies="eberti">
<emphasis id="B911EA8AFFCBFFACFBC3FB1064D917C6" box="[1050,1215,1244,1269]" italics="true" pageId="8" pageNumber="204">C. belli eberti</emphasis>
</taxonomicName>
in surveys on the butterfly fauna from the Central Brazilian Plateau (
<bibRefCitation id="EFF44B69FFCBFFACFD00FACD6388162A" author="Pinheiro, C. E. G. &amp; Emery, E. O." box="[729,1006,1280,1305]" journalOrPublisher="Biota Neotropica" pageId="8" pageNumber="204" pagination="1 - 15" part="6" publicationUrl="https://doi.org/10.1590/S1676-06032006000300012" refString="Pinheiro, C. E. G. &amp; Emery, E. O. (2006) As borboletas (Lepidoptera: Papilionoidea e Hesperioidea) da Area de Protecao Ambiental do Gama e Cabeca de Veado, Distrito Federal, Brasil. Biota Neotropica, 6 (3), 1 - 15. https: // doi. org / 10.1590 / S 1676 - 06032006000300012" title="As borboletas (Lepidoptera: Papilionoidea e Hesperioidea) da Area de Protecao Ambiental do Gama e Cabeca de Veado, Distrito Federal, Brasil" type="journal article" year="2006">Pinheiro &amp; Emery 2006</bibRefCitation>
,
<bibRefCitation id="EFF44B69FFCBFFACFC23FACD64A9162A" author="Mielke, O. H. H. &amp; Emery, E. O. &amp; Pinheiro, C. E. G." box="[1018,1231,1280,1305]" journalOrPublisher="Revista Brasileira de Zoologia" pageId="8" pageNumber="204" pagination="283 - 288" part="52" refString="Mielke, O. H. H., Emery, E. O. &amp; Pinheiro, C. E. G. (2008) As borboletas Hesperiidae (Lepidoptera, Hesperioidea) do Distrito Federal, Brasil. Revista Brasileira de Zoologia, 52 (2), 283 - 288." title="As borboletas Hesperiidae (Lepidoptera, Hesperioidea) do Distrito Federal, Brasil" type="journal article" year="2008">
Mielke
<emphasis id="B911EA8AFFCBFFACFB88FACC64EC162A" box="[1105,1162,1280,1305]" italics="true" pageId="8" pageNumber="204">et al.</emphasis>
2008
</bibRefCitation>
). Both species are easily distinguished by the presence of gray to purple scales on the ventral hind wing basal band of
<taxonomicName id="4C654D1BFFCBFFACFAD9FAEB6507160E" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1280,1377,1316,1341]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="eberti" status="stat. nov.">
<emphasis id="B911EA8AFFCBFFACFAD9FAEB6507160E" box="[1280,1377,1316,1341]" italics="true" pageId="8" pageNumber="204">R. eberti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFCBFFACFAB0FAE9613E1652" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFAB0FAE9613E1652" bold="true" pageId="8" pageNumber="204">stat. nov., comb. nov.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="135E2A1DFFCBFFACFEB1FA85618A1652" box="[360,492,1352,1377]" captionStart="FIGURES 1–20" captionStartId="3.[151,264,1502,1524]" captionTargetBox="[151,1435,196,1478]" captionTargetId="figure@3.[150,1436,195,1478]" captionTargetPageId="3" captionText="FIGURES 1–20. Species of Rufocumbre in dorsal and ventral views, male (left) and female (right). 1–4, R. celioi sp. nov., male holotype and female allotype. 5–8, R. emeryi sp. nov., male holotype and female allotype. 9–12, R. eberti stat. nov., comb. nov., male (OM 59.360) and female (OM 19.333) from Caraça, Catas Altas, Minas Gerais, Brazil. 13–16, R. schneideri sp. nov., male holotype and female allotype. 17–20, R. lucasi sp. nov., male holotype and female allotype.Scale bar = 1 cm." httpUri="https://zenodo.org/record/1117610/files/figure.png" pageId="8" pageNumber="204">Figs 10, 12</figureCitation>
). However, as this character may not be clearly seen in old or damaged specimens, the study of the genitalia is always needed. Males of
<taxonomicName id="4C654D1BFFCBFFACFD38FAA3632A16B7" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[737,844,1389,1413]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="emeryi" status="sp. nov.">
<emphasis id="B911EA8AFFCBFFACFD38FAA3632A16B7" box="[737,844,1389,1413]" italics="true" pageId="8" pageNumber="204">R. emeryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFCBFFACFC8BFAA163CD16B6" box="[850,939,1388,1413]" pageId="8" pageNumber="204" rank="species">
<emphasis id="B911EA8AFFCBFFACFC8BFAA163CD16B6" bold="true" box="[850,939,1388,1413]" pageId="8" pageNumber="204">sp. nov.</emphasis>
</taxonomicNameLabel>
are distinguished from
<taxonomicName id="4C654D1BFFCBFFACFB6BFAA3657416B6" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1202,1298,1388,1413]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="eberti">
<emphasis id="B911EA8AFFCBFFACFB6BFAA3657416B6" box="[1202,1298,1388,1413]" italics="true" pageId="8" pageNumber="204">R. eberti</emphasis>
</taxonomicName>
(
<figureCitation id="135E2A1DFFCBFFACFAF8FAA1651716B6" box="[1313,1393,1388,1413]" captionStart="FIGURES 35–37" captionStartId="9.[151,264,1837,1859]" captionTargetBox="[151,1424,700,1811]" captionTargetId="figure@9.[151,1436,700,1811]" captionTargetPageId="9" captionText="FIGURES 35–37. Male genitalia of species of Rufocumbre.Left to right: dorsal view of tegumen, uncus and gnathos; ventral view of tegumen, uncus and gnathos; lateral view of tegumen, saccus, uncus and gnathos; left view of the right valva; posterior and lateral views of fultura inferior; lateral left, dorsal, ventral and lateral right views of the aedeagus. 35, R. eberti stat. nov., comb. nov., from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 15.367). 36, R. schneideri sp. nov., paratype from Ventania, Paraná, Brazil (DZ 15.736). 37, R. lucasi sp. nov., paratype from Morro do Pai Inácio, Palmeiras, Bahia, Brazil (OM 50.016). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117618/files/figure.png" pageId="8" pageNumber="204">Fig. 35</figureCitation>
) by the shorter distal spine of harpe, never exceeding the distal margin of ampulla (
<figureCitation id="135E2A1DFFCBFFACFBC6FA5D6412169A" box="[1055,1140,1424,1449]" captionStart="FIGURES 33–34" captionStartId="8.[151,264,941,963]" captionTargetBox="[151,1436,193,920]" captionTargetId="figure@8.[151,1436,193,920]" captionTargetPageId="8" captionText="FIGURES 33–34. Male genitalia of species of Rufocumbre.Left to right: dorsal view of tegumen, uncus and gnathos; ventral view of tegumen, uncus and gnathos; lateral view of tegumen, saccus, uncus and gnathos; left view of the right valva; posterior and lateral views of fultura inferior; lateral left, dorsal, ventral and lateral right views of the aedeagus. 33, R. celioi sp. nov., paratype from Caraça, Catas Altas, Minas Gerais, Brazil (DZ 15.577). 34, R. emeryi sp. nov., holotype.Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117616/files/figure.png" pageId="8" pageNumber="204">Fig. 34</figureCitation>
), while the females of the former (
<figureCitation id="135E2A1DFFCBFFACFF2BFA79612216FE" box="[242,324,1460,1485]" captionStart="FIGURES 38–39" captionStartId="10.[151,264,1936,1958]" captionTargetBox="[265,1342,205,1919]" captionTargetId="figure@10.[215,1379,195,1919]" captionTargetPageId="10" captionText="FIGURES 38–39. Female genitalia of species of Rufocumbre in ventral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix. 38, R. celioi sp. nov., paratype from Bituva, Mafra, Santa Catarina, Brazil (DZ 17.124). 39, R. emeryi sp. nov., paratype from Fazenda Água Limpa, Brasília, Distrito Federal, Brazil (DZ 20.891). Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117620/files/figure.png" pageId="8" pageNumber="204">Fig. 39</figureCitation>
) do not have the lamella antevaginalis, whereas it is present in
<taxonomicName id="4C654D1BFFCBFFACFBD1FA7B640E16FE" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" baseAuthorityName="Evans" baseAuthorityYear="1955" box="[1032,1128,1460,1485]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="eberti">
<emphasis id="B911EA8AFFCBFFACFBD1FA7B640E16FE" box="[1032,1128,1460,1485]" italics="true" pageId="8" pageNumber="204">R. eberti</emphasis>
</taxonomicName>
(
<figureCitation id="135E2A1DFFCBFFACFBA1FA7964AA16FE" box="[1144,1228,1460,1485]" captionStart="FIGURES 40–42" captionStartId="11.[151,264,1812,1834]" captionTargetBox="[157,1409,201,1784]" captionTargetId="figure@11.[151,1436,193,1790]" captionTargetPageId="11" captionText="FIGURES 40–42. Female genitalia of species of Rufocumbre in ventral view. Papilla analis, sterigma, tergum VIII and bursa copulatrix.40, R.eberti stat. nov., comb. nov., from Ouro Preto,Minas Gerais, Brazil(OM 66.016).41, R.schneideri sp. nov., paratype from Serra Grande, Conceição dos Ouros, Minas Gerais, Brazil (OM 59.073). 42, R. lucasi sp. nov., paratype from Morro do Pai Inácio, Palmeiras, Bahia, Brazil (OM 50.179).Scale bar = 1 mm." httpUri="https://zenodo.org/record/1117622/files/figure.png" pageId="8" pageNumber="204">Fig. 40</figureCitation>
).
</paragraph>
<paragraph id="8BDA3698FFCBFFACFF1EFA156365150A" blockId="8.[151,1436,1100,2025]" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFF1EFA1562D516C2" bold="true" box="[199,691,1496,1521]" pageId="8" pageNumber="204">Geographical distribution and phenology</emphasis>
. This species was recorded only in areas of Cerrado vegetation of the states of Mato Grosso, Goiás, Distrito Federal and Minas Gerais, between 350 and 1200 m (
<figureCitation id="135E2A1DFFCBFFACFB65FA3165691526" box="[1212,1295,1532,1557]" captionStart="FIGURE 55" captionStartId="17.[151,250,1317,1339]" captionTargetBox="[151,1436,195,1293]" captionTargetId="figure@17.[151,1436,195,1293]" captionTargetPageId="17" captionText="FIGURE 55.Geographical distribution of Rufocumbre emeryi sp. nov. and R. eberti stat. nov., comb. nov." httpUri="https://zenodo.org/record/1117630/files/figure.png" pageId="8" pageNumber="204">Fig. 55</figureCitation>
). Specimens were collected in January, April to July, and December.
</paragraph>
</subSubSection>
<subSubSection id="C37F6513FFCBFFACFF1EF98963311596" pageId="8" pageNumber="204" type="etymology">
<paragraph id="8BDA3698FFCBFFACFF1EF98963311596" blockId="8.[151,1436,1100,2025]" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFF1EF9896128156E" bold="true" box="[199,334,1604,1629]" pageId="8" pageNumber="204">Etymology.</emphasis>
This new species is named in memory of our friend the late Eduardo Oliveira Emery, in gratitude to his extensive support. Eduardo always kindly brought us with relevant information about the
<taxonomicName id="4C654D1BFFCBFFACFB16F9A5653015B2" box="[1231,1366,1640,1665]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="order">Lepidoptera</taxonomicName>
fauna from Central Brazil, including specimens of
<taxonomicName id="4C654D1BFFCBFFACFD52F94362901596" authority="Dolibaina &amp; Mielke &amp; Casagrande, 2017" authorityName="Dolibaina &amp; Mielke &amp; Casagrande" authorityYear="2017" box="[651,758,1678,1701]" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="emeryi" status="sp. nov.">
<emphasis id="B911EA8AFFCBFFACFD52F94362901596" box="[651,758,1678,1701]" italics="true" pageId="8" pageNumber="204">R. emeryi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A22257F1FFCBFFACFD24F94163311596" box="[765,855,1676,1701]" pageId="8" pageNumber="204" rank="species">
<emphasis id="B911EA8AFFCBFFACFD24F94163311596" bold="true" box="[765,855,1676,1701]" pageId="8" pageNumber="204">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C37F6513FFCBFFADFF1EF97D646311B7" lastPageId="9" lastPageNumber="205" pageId="8" pageNumber="204" type="materials_examined">
<paragraph id="8BDA3698FFCBFFACFF1EF97D63491423" blockId="8.[151,1436,1100,2025]" pageId="8" pageNumber="204">
<emphasis id="B911EA8AFFCBFFACFF1EF97D61D215FA" bold="true" box="[199,436,1712,1737]" pageId="8" pageNumber="204">Examined material.</emphasis>
The holotype male has the following labels: /
<materialsCitation id="3B0D3CC5FFCBFFACFC1FF97C63491423" ID-GBIF-Occurrence="1795578602" collectingDate="1979-06-16" collectingDateMax="1979-06-30" collectingDateMin="1979-06-16" collectionCode="DZUP" collectorName="C. Elias" country="Brazil" location="Corinto - M [inas] G [erais" pageId="8" pageNumber="204" specimenCount="1" typeStatus="holotype">
<typeStatus id="54DE883AFFCBFFACFC1FF97C640A15FA" box="[966,1132,1713,1737]" pageId="8" pageNumber="204" type="holotype">HOLOTYPUS</typeStatus>
/
<collectingDate id="EF9FE9B0FFCBFFACFB5DF97D654815FA" box="[1156,1326,1712,1737]" pageId="8" pageNumber="204" value="1979-06-16" valueMax="1979-06-30" valueMin="1979-06-16">
<date id="FFDB1058FFCBFFACFB5DF97D654815FA" box="[1156,1326,1712,1737]" pageId="8" pageNumber="204" value="1979-06-16" valueMax="1979-06-30" valueMin="1979-06-16">16-30-VI-1979</date>
</collectingDate>
<location id="8EBA6043FFCBFFACFAECF97D612D15DE" LSID="urn:lsid:plazi:treatment:03CC878EFFC5FFADFF4EFA89646311B7:8EBA6043FFCBFFACFAECF97D612D15DE" country="Brazil" name="Corinto - M [inas] G [erais" pageId="8" pageNumber="204">Corinto - M[inas]G[erais</location>
,
<collectingCountry id="F3727608FFCBFFACFE8FF91961FB15DE" box="[342,413,1748,1773]" name="Brazil" pageId="8" pageNumber="204">Brazil</collectingCountry>
]
<collectorName id="2690534EFFCBFFACFE73F918626415DE" box="[426,514,1748,1773]" pageId="8" pageNumber="204">C. Elias</collectorName>
leg / Gen[italia]. Prep[ared]. Dolibaina 2010 / DZ 15.626 / Holotypus
<taxonomicName id="4C654D1BFFCBFFACFACDF919623B1422" authority="Dolibaina, Mielke &amp; Casagrande" authorityName="Dolibaina, Mielke &amp; Casagrande" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B911EA8AFFCBFFACFACDF91960851423" italics="true" pageId="8" pageNumber="204">Rufocumbre emeryi</emphasis>
Dolibaina, Mielke &amp; Casagrande
</taxonomicName>
det. 2017/.
<collectionCode id="ED74AE5DFFCBFFACFD38F934634D1423" box="[737,811,1785,1808]" collectionName="Brazil, Parana, Curitiba, Universidade Federal do Parana, Museu de Entomologia Pe. Jesus Santiago Moure" pageId="8" pageNumber="204">DZUP</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="8BDA3698FFCBFFACFF1EF8D1618A144F" blockId="8.[151,1436,1100,2025]" pageId="8" pageNumber="204">
The allotype female has the following labels: / ALLOTYPUS / Brasil D[istrito]F[ederal] Cerrado Faz[enda]. Água Limpa 20-VI-2005 E.O. Emery Coletor / DZ 17.097 / Allotypus
<taxonomicName id="4C654D1BFFCBFFACFC11F88D617C144E" authority="Dolibaina, Mielke &amp; Casagrande" authorityName="Dolibaina, Mielke &amp; Casagrande" class="Insecta" family="Hesperiidae" genus="Rufocumbre" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="204" phylum="Arthropoda" rank="species" species="emeryi">
<emphasis id="B911EA8AFFCBFFACFC11F88D64C0146B" box="[968,1190,1856,1881]" italics="true" pageId="8" pageNumber="204">Rufocumbre emeryi</emphasis>
Dolibaina, Mielke &amp; Casagrande
</taxonomicName>
det. 2017/. DZUP.
</paragraph>
<paragraph id="8BDA3698FFCBFFADFF1EF844646311B7" blockId="8.[151,1436,1100,2025]" lastBlockId="9.[151,1436,151,645]" lastPageId="9" lastPageNumber="205" pageId="8" pageNumber="204">
Paratypes: BRAZIL –
<emphasis id="B911EA8AFFCBFFACFE18F84462351493" box="[449,595,1929,1952]" italics="true" pageId="8" pageNumber="204">Mato Grosso</emphasis>
:
<emphasis id="B911EA8AFFCBFFACFDB8F845633A1492" bold="true" box="[609,860,1928,1953]" pageId="8" pageNumber="204">Ribeirão Cascalheira</emphasis>
(11-18 Km N), 350 m, 21-23.VII.1997, Mielke
<emphasis id="B911EA8AFFCBFFACFAAAF84565F31492" box="[1395,1429,1928,1953]" italics="true" pageId="8" pageNumber="204">leg</emphasis>
. 1 male (DZ 15.612*) (DZUP).
<emphasis id="B911EA8AFFCBFFACFE21F860625D14F7" box="[504,571,1965,1988]" italics="true" pageId="8" pageNumber="204">Goiás</emphasis>
:
<emphasis id="B911EA8AFFCBFFACFD93F861632914F6" bold="true" box="[586,847,1964,1989]" pageId="8" pageNumber="204">Alto Paraíso de Goiás</emphasis>
(Parque Nacional da Chapada dos Veadeiros), 1490 m, 3.VI.2011, Emery
<emphasis id="B911EA8AFFCBFFACFE4AF81D61DA14DA" box="[403,444,2000,2025]" italics="true" pageId="8" pageNumber="204">leg.</emphasis>
1 male (ex. coll. Eduardo Emery, DZ 39.080) (DZUP), (Pousada Portal da Chapada), 1160 m, 4.VI.2011, Emery
<emphasis id="B911EA8AFFCAFFADFE05FF5A61981383" box="[476,510,151,176]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. (ex. coll. Eduardo Emery, DZ 39.120) (DZUP);
<emphasis id="B911EA8AFFCAFFADFB94FF5A654E1383" bold="true" box="[1101,1320,151,176]" pageId="9" pageNumber="205">Leopoldo Bulhões</emphasis>
, 1050 m, XII.1937, Spitz
<emphasis id="B911EA8AFFCAFFADFE96FF71611713E6" box="[335,369,188,213]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (OM 11.010*) (OM);
<taxonomicName id="4C654D1BFFCAFFADFD0DFF7164E013E7" authority="(Corrego Grotao)" authorityName="Corrego Grotao" box="[724,1158,188,213]" class="Actinopterygii" family="Characidae" genus="Planaltina" higherTaxonomySource="GBIF" kingdom="Animalia" order="Characiformes" pageId="9" pageNumber="205" phylum="Chordata" rank="genus">
<emphasis id="B911EA8AFFCAFFADFD0DFF71632913E6" bold="true" box="[724,847,188,213]" pageId="9" pageNumber="205">Planaltina</emphasis>
(Córrego Grotão), 1000 m
</taxonomicName>
, 6.V.2005, Emery
<emphasis id="B911EA8AFFCAFFADFA85FF71651813E6" box="[1372,1406,188,213]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (DZ 8.676) (DZUP).
<emphasis id="B911EA8AFFCAFFADFE0DFF2C62E813CB" box="[468,654,223,248]" italics="true" pageId="9" pageNumber="205">Distrito Federal</emphasis>
:
<emphasis id="B911EA8AFFCAFFADFD7BFF12629913CB" bold="true" box="[674,767,223,248]" pageId="9" pageNumber="205">Brasília</emphasis>
(Brasília Country Club), 1200 m, 19.VI.1972, Mielke &amp; Brown
<emphasis id="B911EA8AFFCAFFADFF29FEC96174122E" box="[240,274,260,285]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (DZ 17.102) (DZUP), (Centro Olímpico), 1000 m, 4.V.1977, Gifford
<emphasis id="B911EA8AFFCAFFADFB1AFEC9648B122E" box="[1219,1261,260,285]" italics="true" pageId="9" pageNumber="205">leg.</emphasis>
1 female (DZ 15.633*) (DZUP), (Fazenda Água Limpa), 1150 m, 2.V.1980, Gifford
<emphasis id="B911EA8AFFCAFFADFC29FEEA64741273" box="[1008,1042,295,320]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (DZ 15.395*) (DZUP), 17.VII.2004, Emery
<emphasis id="B911EA8AFFCAFFADFE5BFE8161CD1256" box="[386,427,332,357]" italics="true" pageId="9" pageNumber="205">leg.</emphasis>
3 males and 1 female (ex. coll. Eduardo Emery, DZ 20.884, DZ 39.050, DZ 39.433, DZ 20.891*) (DZUP), 20.VI.2005, Emery
<emphasis id="B911EA8AFFCAFFADFD92FEA2621212BB" box="[587,628,367,392]" italics="true" pageId="9" pageNumber="205">leg.</emphasis>
1 male and 2 females (ex. coll. Eduardo Emery, DZ 39.060, DZ 20.898, DZ 39.100) (DZUP), (Jardim Botânico), 6.VI.2010, Emery
<emphasis id="B911EA8AFFCAFFADFC9FFE59630E129E" box="[838,872,404,429]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 female (ex. coll. Eduardo Emery, DZ 39.040) (DZUP) (Cabeça do Veado), 1070 m, 13.IV.1993, Mielke &amp; Becker
<emphasis id="B911EA8AFFCAFFADFC66FE7A638712E3" box="[959,993,439,464]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (OM 34.861) (OM), (Parque Nacional de Brasília), 1190 m, Emery
<emphasis id="B911EA8AFFCAFFADFD95FE11620812C6" box="[588,622,476,501]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. (ex. coll. Eduardo Emery, DZ 39.070) (DZUP), (Poço Azul), 1.I.2004, Emery
<emphasis id="B911EA8AFFCAFFADFF35FE326168112B" box="[236,270,511,536]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male and 1 female (ex. coll. Eduardo Emery, DZ 39.413, DZ 39.110) (DZUP), (Rio Maranhão), 20.V.2002, Emery
<emphasis id="B911EA8AFFCAFFADFEB1FDE961EC110E" box="[360,394,548,573]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 female (ex. coll. Eduardo Emery, DZ 39.090) (DZUP).
<emphasis id="B911EA8AFFCAFFADFBCDFDE864CA110F" box="[1044,1196,549,572]" italics="true" pageId="9" pageNumber="205">Minas Gerais</emphasis>
:
<emphasis id="B911EA8AFFCAFFADFB63FDE96558110E" bold="true" box="[1210,1342,548,573]" pageId="9" pageNumber="205">Nova Lima</emphasis>
(Parque Estadual Serra do Rola Moça), 1271 m, 25.I.2013, S. P. A. Franco
<emphasis id="B911EA8AFFCAFFADFC49FD8A63D41153" box="[912,946,583,608]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (UFMG ILE-1400058*) (UFMG);
<emphasis id="B911EA8AFFCAFFADFF4EFDA1612611B6" bold="true" box="[151,320,620,645]" pageId="9" pageNumber="205">Três Corações</emphasis>
, 900 m, 24.VII.1971, Ebert
<emphasis id="B911EA8AFFCAFFADFDA7FDA162C611B6" box="[638,672,620,645]" italics="true" pageId="9" pageNumber="205">leg</emphasis>
. 1 male (DZ 15.570*) (DZUP).
</paragraph>
</subSubSection>
</treatment>
</document>
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