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<document id="4797AA29DC6942402288E21AC97C7567" ID-DOI="10.6620/ZS.2022.61-71" ID-ISSN="1810-522X" ID-PMC="PMC9810989" ID-PubMed="36644631" ID-Zenodo-Dep="12827218" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721933062855" checkinUser="felipe" docAuthor="Zhang, Ying-Chen &amp; Shih, Hsi-Te" docDate="2022" docId="03B18F3CFFFCFFF54DC2FF6EFE4A1BB5" docLanguage="en" docName="ZoolStud.61.71.1-29.pdf" docOrigin="Zoological Studies 61 (71)" docSource="http://dx.doi.org/10.5281/zenodo.12826148" docStyle="DocumentStyle:E2601EF9E8BBDC4537EE724C10FF512F.1:ZoolStud.2019-.journal_article" docStyleId="E2601EF9E8BBDC4537EE724C10FF512F" docStyleName="ZoolStud.2019-.journal_article" docStyleVersion="1" docTitle="Tubuca arcuata" docType="treatment" docVersion="2" lastPageNumber="20" masterDocId="FF88F744FFEDFFE64D29FF93FFE61860" masterDocTitle="Fig. 3 in Historical Biogeography of the Group (Anura, Leptodactylidae): Identification of Ancestral Areas and Events that Modeled their Distribution." masterLastPageNumber="29" masterPageNumber="1" pageNumber="18" updateTime="1721937445007" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="6B4DAB8FFCACB4F8C2731EC13A907BC7">Fig. 3 in Historical Biogeography of the Group (Anura, Leptodactylidae): Identification of Ancestral Areas and Events that Modeled their Distribution.</mods:title>
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<mods:namePart id="8BD34D460EE03FB7225B2E468BDD04F3">Zhang, Ying-Chen</mods:namePart>
<mods:affiliation id="E77657E3DDF2ED3D7B8D811CF70E068E">Department of Life Science, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<mods:namePart id="586A578D7225F9C86CFD935411FC2E69">Shih, Hsi-Te</mods:namePart>
<mods:affiliation id="2EFA99564360ACDD454E4A97B666458F">Department of Life Science, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan. &amp; Research Center for Global Change Biology, National Chung Hsing University, 250 Kuo Kuang Road, Taichung 402, Taiwan.</mods:affiliation>
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<mods:title id="6728DA34F55251BDB4F5B51EF8A87A38">Zoological Studies</mods:title>
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<mods:date id="8BE1DE49C3C786462CB6748E7F06B6F1">2022</mods:date>
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<treatment id="03B18F3CFFFCFFF54DC2FF6EFE4A1BB5" ID-DOI="http://doi.org/10.5281/zenodo.12827264" ID-Zenodo-Dep="12827264" LSID="urn:lsid:plazi:treatment:03B18F3CFFFCFFF54DC2FF6EFE4A1BB5" httpUri="http://treatment.plazi.org/id/03B18F3CFFFCFFF54DC2FF6EFE4A1BB5" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18">
<subSubSection id="C3026DA1FFFCFFF74DC2FF6EFDF71956" pageId="17" pageNumber="18" type="nomenclature">
<paragraph id="8BA73E2AFFFCFFF74DC2FF6EFDF71956" blockId="17.[235,638,253,310]" pageId="17" pageNumber="18">
<taxonomicName id="4C1845A9FFFCFFF74DC2FF6EFD981976" ID-CoL="59JXX" authority="(De Haan, 1835)" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[235,638,253,278]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="B96CE238FFFCFFF74DC2FF6EFD981976" bold="true" box="[235,638,253,278]" pageId="17" pageNumber="18">
<emphasis id="B96CE238FFFCFFF74DC2FF6EFE561976" bold="true" box="[235,432,253,278]" italics="true" pageId="17" pageNumber="18">Tubuca arcuata</emphasis>
(De Haan, 1835)
</emphasis>
</taxonomicName>
(
<figureCitation id="132322AFFFFCFFF74C49FE8EFE491956" box="[352,431,285,310]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="17" pageNumber="18">Fig. 12</figureCitation>
,
<tableCitation id="C69A0B91FFFCFFF74C90FE8EFDEE1956" box="[441,520,285,310]" captionStart="Table 3" captionStartId="7.[136,198,904,928]" captionText="Table 3. The setation of different appendages in zoea stage I of 15 species of fiddler crabs from Taiwan. See figures 216" pageId="17" pageNumber="18">Table 3</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3026DA1FFFCFFF54DECFECAFE4A1BB5" lastPageId="19" lastPageNumber="20" pageId="17" pageNumber="18" type="materials_examined">
<paragraph id="8BA73E2AFFFCFFF74DECFECAFDFB19D1" blockId="17.[136,736,345,434]" pageId="17" pageNumber="18">
<emphasis id="B96CE238FFFCFFF74DECFECAFE781911" box="[197,414,345,369]" italics="true" pageId="17" pageNumber="18">Material examined</emphasis>
: zoea I (from
<geoCoordinate id="EE2C58EDFFFCFFF74F67FEC8FD421911" box="[590,676,345,371]" degrees="24.9" direction="north" orientation="latitude" pageId="17" pageNumber="18" precision="5555" value="24.9">
<emphasis id="B96CE238FFFCFFF74F67FEC8FDBB1913" bold="true" box="[590,605,347,371]" pageId="17" pageNumber="18">ñ</emphasis>
<quantity id="4CE093CFFFFCFFF74F5AFECAFD421911" box="[627,676,345,369]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="17" pageNumber="18" unit="mm" value="24.9">24.9</quantity>
</geoCoordinate>
mm, NCHUZOOL 16785, Bajhang River estuary,
<collectingRegion id="49DCF0C8FFFCFFF74FA7FEEAFD3A19F1" box="[654,732,377,401]" country="Taiwan" name="Tainan" pageId="17" pageNumber="18">Tainan</collectingRegion>
, coll. Y.-C. Zhang et al.,
<date id="FFA618EAFFFCFFF74CAEFE0AFDF719D1" box="[391,529,409,434]" pageId="17" pageNumber="18" value="2020-05-09">9 May. 2020</date>
).
</paragraph>
<paragraph id="8BA73E2AFFFCFFF74E40FF6DFAE41955" blockId="17.[813,1413,253,438]" pageId="17" pageNumber="18">
<emphasis id="B96CE238FFFCFFF74E40FF6DFC721975" box="[873,916,254,277]" italics="true" pageId="17" pageNumber="18">Size</emphasis>
: cl. 0.34 ±
<quantity id="4CE093CFFFFCFFF7493FFF6DFB611975" box="[1046,1159,254,278]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="17" pageNumber="18" unit="mm" value="0.01">0.010 mm</quantity>
, rdl. 0.53 ±
<quantity id="4CE093CFFFFCFFF74839FF6DFA671975" box="[1296,1409,254,278]" metricMagnitude="-5" metricUnit="m" metricValue="1.9" pageId="17" pageNumber="18" unit="mm" value="0.019">0.019 mm</quantity>
, dsl. 0.08 ±
<quantity id="4CE093CFFFFCFFF74E8DFE8DFBF41955" box="[932,1042,286,310]" metricMagnitude="-6" metricUnit="m" metricValue="4.0" pageId="17" pageNumber="18" unit="mm" value="0.004">0.004 mm</quantity>
, rsl. 0.11 ±
<quantity id="4CE093CFFFFCFFF749A6FE8EFB181955" box="[1167,1278,285,309]" metricMagnitude="-6" metricUnit="m" metricValue="6.0" pageId="17" pageNumber="18" unit="mm" value="0.006">0.006 mm</quantity>
.
</paragraph>
<paragraph id="8BA73E2AFFFCFFF74E40FEADFB4819D5" blockId="17.[813,1413,253,438]" pageId="17" pageNumber="18">
<emphasis id="B96CE238FFFCFFF74E40FEADFC3C1935" box="[873,986,318,341]" italics="true" pageId="17" pageNumber="18">Carapace</emphasis>
(
<figureCitation id="132322AFFFFCFFF74EC7FEAEFBB91936" box="[1006,1119,317,342]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="17" pageNumber="18">Fig. 12A</figureCitation>
): Lateral spines lacking, dorsal spine curved, relatively short, rostral spine straight, dorsal spine and rostral spine with pointed termination. Eyes round and sessile.
</paragraph>
<caption id="DF676EA2FFFCFFF74DA1F817FF241FD3" ID-DOI="http://doi.org/10.5281/zenodo.12827244" ID-Zenodo-Dep="12827244" httpUri="https://zenodo.org/record/12827244/files/figure.png" pageId="17" pageNumber="18" startId="17.[136,167,1924,1943]" targetBox="[232,1335,486,1893]" targetPageId="17" targetType="figure">
<paragraph id="8BA73E2AFFFCFFF74DA1F817FF241FD3" blockId="17.[136,1414,1924,1971]" pageId="17" pageNumber="18">
<emphasis id="B96CE238FFFCFFF74DA1F817FF2C1FF7" bold="true" box="[136,202,1924,1943]" pageId="17" pageNumber="18">Fig. 11.</emphasis>
<taxonomicName id="4C1845A9FFFCFFF74DFDF817FEAE1FF7" baseAuthorityName="Stimpson" baseAuthorityYear="1858" box="[212,328,1924,1943]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B96CE238FFFCFFF74DFDF817FEAE1FF7" box="[212,328,1924,1943]" italics="true" pageId="17" pageNumber="18">Tubuca acuta</emphasis>
</taxonomicName>
, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson.
</paragraph>
</caption>
<paragraph id="8BA73E2AFFFFFFF44DC3FF6EFE0A1935" blockId="18.[174,776,253,438]" pageId="18" pageNumber="19">
<emphasis id="B96CE238FFFFFFF44DC3FF6EFE821975" box="[234,356,253,277]" italics="true" pageId="18" pageNumber="19">Antennule</emphasis>
(
<figureCitation id="132322AFFFFFFFF44C53FF6EFE081976" box="[378,494,253,278]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="18" pageNumber="19">Fig. 12B</figureCitation>
): Uniramous, endopod absent. Exopod smooth, unarticled, with 2 aesthetascs and 2 simple setae terminally.
</paragraph>
<paragraph id="8BA73E2AFFFFFFF44DC3FECDFAF31975" blockId="18.[174,776,253,438]" lastBlockId="18.[850,1451,253,438]" pageId="18" pageNumber="19">
<emphasis id="B96CE238FFFFFFF44DC3FECDFEA81915" box="[234,334,350,373]" italics="true" pageId="18" pageNumber="19">Antenna</emphasis>
(
<figureCitation id="132322AFFFFFFFF44C4DFECEFE3F1916" box="[356,473,349,374]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="18" pageNumber="19">Fig. 12C</figureCitation>
): Protopod basal region smooth, distal end acuminate, bearing 2 rows of spines. Exopod short, with 1 terminal simple seta, setae not reaching tip of protopod. Endopod absent.
</paragraph>
<paragraph id="8BA73E2AFFFFFFF54EA6FE8EFEBF1975" blockId="18.[850,1451,253,438]" lastBlockId="19.[136,737,253,982]" lastPageId="19" lastPageNumber="20" pageId="18" pageNumber="19">
<emphasis id="B96CE238FFFFFFF44EA6FE8EFC1F1955" box="[911,1017,285,309]" italics="true" pageId="18" pageNumber="19">Maxillule</emphasis>
(
<figureCitation id="132322AFFFFFFFF44923FE8EFB921956" box="[1034,1140,285,310]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="18" pageNumber="19">Fig. 12D</figureCitation>
): Epipod seta absent; coxal endite with 4 plumodenticulate setae and 1 plumose seta; basial endite with 4 plumodenticulate setae and 1 plumose seta; endopod 2-articled, proximal article without setae, 4 sparsely plumose in distal article; exopod seta absent.
</paragraph>
<caption id="DF676EA2FFFFFFF44D87F817FEEA1FD3" ID-DOI="http://doi.org/10.5281/zenodo.12827246" ID-Zenodo-Dep="12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="18" pageNumber="19" startId="18.[174,205,1924,1943]" targetBox="[270,1370,479,1894]" targetPageId="18" targetType="figure">
<paragraph id="8BA73E2AFFFFFFF44D87F817FEEA1FD3" blockId="18.[174,1451,1924,1971]" pageId="18" pageNumber="19">
<emphasis id="B96CE238FFFFFFF44D87F817FF171FF7" bold="true" box="[174,241,1924,1943]" pageId="18" pageNumber="19">Fig. 12.</emphasis>
<taxonomicName id="4C1845A9FFFFFFF44DD4F817FE621FF7" baseAuthorityName="De Haan" baseAuthorityYear="1835" box="[253,388,1924,1943]" class="Malacostraca" family="Ocypodidae" genus="Tubuca" kingdom="Animalia" order="Decapoda" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="arcuata">
<emphasis id="B96CE238FFFFFFF44DD4F817FE621FF7" box="[253,388,1924,1943]" italics="true" pageId="18" pageNumber="19">Tubuca arcuata</emphasis>
</taxonomicName>
, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson.
</paragraph>
</caption>
<paragraph id="8BA73E2AFFFEFFF54DECFE8EFEFA19B5" blockId="19.[136,737,253,982]" pageId="19" pageNumber="20">
<emphasis id="B96CE238FFFEFFF54DECFE8EFEFE1955" box="[197,280,285,309]" italics="true" pageId="19" pageNumber="20">Maxilla</emphasis>
(
<figureCitation id="132322AFFFFEFFF54C0FFE8EFE6E1956" box="[294,392,285,310]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="19" pageNumber="20">Fig. 12E</figureCitation>
): Coxal endite bilobed with 3+3 plumose setae; basial endite bilobed with 5+4 plumose setae; endopod bilobed with 1+2 terminal sparsely plumose; scaphognathite with 4 densely plumose setae and a long acuminate posterior process with microtrichias.
</paragraph>
<paragraph id="8BA73E2AFFFEFFF54DECFE4DFEBA1A35" blockId="19.[136,737,253,982]" pageId="19" pageNumber="20">
<emphasis id="B96CE238FFFEFFF54DECFE4DFE901995" box="[197,374,477,501]" italics="true" pageId="19" pageNumber="20">First maxilliped</emphasis>
(
<figureCitation id="132322AFFFFEFFF54CAEFE4EFE0D1996" box="[391,491,477,502]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="19" pageNumber="20">Fig. 12F</figureCitation>
): Basis with 2+2+3+2 sparsely plumose setae; endopod 5-articled with 2, 2, 1, 2, 5 plumose setae; exopod 2-articled with 4 terminal long plumose setae.
</paragraph>
<paragraph id="8BA73E2AFFFEFFF54DECFDCDFE5D1AB5" blockId="19.[136,737,253,982]" pageId="19" pageNumber="20">
<emphasis id="B96CE238FFFEFFF54DECFDCDFE491A15" box="[197,431,605,629]" italics="true" pageId="19" pageNumber="20">Second maxilliped</emphasis>
(
<figureCitation id="132322AFFFFEFFF54CE1FDCDFDDE1A16" box="[456,568,605,630]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[136,167,1954,1973]" captionStartId-1="4.[174,205,1924,1943]" captionTargetBox-1="[276,1352,491,1896]" captionTargetId-1="figure-111@4.[251,1373,478,1914]" captionTargetPageId-1="4" captionText-0="Fig. 1. The measurements of the first zoeal stage used in this study." captionText-1="Fig. 2. Austruca lactea, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi-1="http://doi.org/10.5281/zenodo.12827220" httpUri-1="https://zenodo.org/record/12827220/files/figure.png" pageId="19" pageNumber="20">Fig. 12G</figureCitation>
): Basis with 1+1+1+1 sparsely plumose setae; endopod 3-articled with 0, 0, 5 plumose setae; exopod 2-articled with 4 terminal long plumose setae.
</paragraph>
<paragraph id="8BA73E2AFFFEFFF54DECFD4EFE4C1BF5" blockId="19.[136,737,253,982]" pageId="19" pageNumber="20">
<emphasis id="B96CE238FFFEFFF54DECFD4EFEEC1A95" box="[197,266,733,757]" italics="true" pageId="19" pageNumber="20">Pleon</emphasis>
(
<figureCitation id="132322AFFFFEFFF54C36FD4EFE721A96" box="[287,404,733,758]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="19" pageNumber="20">Fig. 12H</figureCitation>
): With 5 pleonites, second pleonite with a pair of relatively acute dorsolateral spines pointing anterodorsally; third pleonite with a pair of relatively obtuse dorsolateral spines pointing posterodorsally; fourth pleonite wider than other pleonites. Pleopods absent.
</paragraph>
<paragraph id="8BA73E2AFFFEFFF54DECFC0EFE4A1BB5" blockId="19.[136,737,253,982]" pageId="19" pageNumber="20">
<taxonomicName id="4C1845A9FFFEFFF54DECFC0EFEEC1BD5" authorityName="Pearse" authorityYear="1952" box="[197,266,925,949]" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="19" pageNumber="20" phylum="Arthropoda" rank="genus">
<emphasis id="B96CE238FFFEFFF54DECFC0EFEEC1BD5" box="[197,266,925,949]" italics="true" pageId="19" pageNumber="20">Telson</emphasis>
</taxonomicName>
(
<figureCitation id="132322AFFFFEFFF54C31FC0EFE9B1BD6" box="[280,381,925,950]" captionStart="Fig" captionStartId="18.[174,205,1924,1943]" captionTargetBox="[270,1370,479,1894]" captionTargetId="figure-6@18.[244,1381,471,1927]" captionTargetPageId="18" captionText="Fig. 12. Tubuca arcuata, zoea I. A, carapace; B, antennule; C, antenna; D, maxillule; E, maxilla; F, first maxilliped; G, second maxilliped; H, pleon and telson." figureDoi="http://doi.org/10.5281/zenodo.12827246" httpUri="https://zenodo.org/record/12827246/files/figure.png" pageId="19" pageNumber="20">Fig. 12H</figureCitation>
):
<taxonomicName id="4C1845A9FFFEFFF54CB9FC0EFE311BD5" authorityName="Pearse" authorityYear="1952" box="[400,471,925,949]" class="Copepoda" family="Telsidae" genus="Telson" kingdom="Animalia" order="Cyclopoida" pageId="19" pageNumber="24" phylum="Arthropoda" rank="genus">Telson</taxonomicName>
bifurcated, inner margin with 6 (3+3) serrate spines.
</paragraph>
</subSubSection>
</treatment>
</document>