166 lines
21 KiB
XML
166 lines
21 KiB
XML
<document id="3D311D1FB84E1414B6E2C1F9626B8AC6" ID-CLB-Dataset="112023" ID-DOI="10.11646/zootaxa.5103.1.1" ID-GBIF-Dataset="1bf32ba1-8e0d-4435-ba04-6d7dbd0d7238" ID-ISSN="1175-5326" ID-Zenodo-Dep="6245709" ID-ZooBank="9E1A72E7-3862-44F7-B69F-ECE64B239FF9" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1645601651449" checkinUser="plazi" docAuthor="Kirejtshuk, Alexander G. & Kovalev, Alexey V." docDate="2022" docId="03AC7326767FD67475E0FC21FA83FEFA" docLanguage="en" docName="zootaxa.5103.1.1.pdf" docOrigin="Zootaxa 5103 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Conotelus Erichson 1843" docType="treatment" docVersion="8" lastPageNumber="12" masterDocId="FF950B5E7674D6787577FFF3FFF4FFE7" masterDocTitle="Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily" masterLastPageNumber="133" masterPageNumber="1" pageNumber="12" updateTime="1699299454408" updateUser="plazi">
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<mods:title id="33A48536FC54CEEABF2AD94F812AE8E5">Monograph on the Cillaeinae (Coleoptera: Nitidulidae) from the Australian Region with comments on the taxonomy of the subfamily</mods:title>
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<mods:namePart id="33F76236C779AD591498DDF16DC7C32E">Kirejtshuk, Alexander G.</mods:namePart>
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<mods:namePart id="EED2FE0B754FFF1A8F13F87BBBFC904E">Kovalev, Alexey V.</mods:namePart>
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<mods:title id="3C2AE5F0AA8FBBC98A2266BA72A7A00E">Zootaxa</mods:title>
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<mods:date id="F1097A595D92AA09BA0B5FD6558D084A">2022</mods:date>
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<mods:number id="C2567DC798F708E896803377EAC3EC3B">2022-02-23</mods:number>
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<mods:number id="AE49E2A40E63161C681440AA9905825C">5103</mods:number>
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<treatment id="03AC7326767FD67475E0FC21FA83FEFA" ID-DOI="http://doi.org/10.5281/zenodo.6828576" ID-GBIF-Taxon="198574625" ID-Zenodo-Dep="6828576" LSID="urn:lsid:plazi:treatment:03AC7326767FD67475E0FC21FA83FEFA" httpUri="http://treatment.plazi.org/id/03AC7326767FD67475E0FC21FA83FEFA" lastPageId="12" lastPageNumber="12" pageId="11" pageNumber="12">
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<subSubSection id="C31F91BB767FD67375E0FC21FC51FC0A" box="[151,933,978,1005]" pageId="11" pageNumber="12" type="nomenclature">
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<paragraph id="8BBAC230767FD67375E0FC21FC51FC0A" blockId="11.[151,1437,978,2013]" box="[151,933,978,1005]" pageId="11" pageNumber="12">
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<emphasis id="B9711E22767FD67375E0FC21FF51FC0B" bold="true" box="[151,165,978,1004]" pageId="11" pageNumber="12">1</emphasis>
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.
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<taxonomicName id="4C05B9B3767FD67375C4FC20FDF9FC0A" ID-CoL="3TG5" ID-ENA="225177" authority="Erichson, 1843: 249" authorityName="Erichson" authorityPageNumber="249" authorityYear="1843" box="[179,525,979,1005]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD67375C4FC20FED1FC0A" box="[179,293,979,1005]" italics="true" pageId="11" pageNumber="12">Conotelus</emphasis>
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<bibRefCitation id="EF94BFC1767FD673745BFC20FDF9FC0A" author="Erichson, W. F." box="[300,525,979,1005]" pageId="11" pageNumber="12" pagination="225 - 363" refId="ref104407" refString="Erichson, W. F. (1843) Versuch einer systematischen Eitheilung der Nitidularien. Germar Zeitung fur Entomologie, 34, 225 - 363." type="journal article" year="1843">Erichson, 1843: 249</bibRefCitation>
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</taxonomicName>
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—Nearctic and Neotropical regions.
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</paragraph>
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</subSubSection>
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<subSubSection id="C31F91BB767FD67475B0FC05FA83FEFA" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12" type="diagnosis">
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<paragraph id="8BBAC230767FD67375B0FC05FAD3FB9A" blockId="11.[151,1437,978,2013]" pageId="11" pageNumber="12">
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<emphasis id="B9711E22767FD67375B0FC05FECEFBF7" bold="true" box="[199,314,1014,1040]" pageId="11" pageNumber="12">Diagnosis</emphasis>
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. Body subcylindrical and very narrow with convex both upper and below surface. Labrum transverse and without trace of median suture. Abdominal segments telescopically conjoined and following segment more or less retracted into previous one; articulation between ventrites and laterosternites forming arc in cross-section (but not forming sharp fold). Male anal sclerite very long and with proximal parts of its ventral edges closed.
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</paragraph>
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<paragraph id="8BBAC230767FD67375B0FB75FB0BF95A" blockId="11.[151,1437,978,2013]" pageId="11" pageNumber="12">
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<emphasis id="B9711E22767FD67375B0FB75FEAFFB47" bold="true" box="[199,347,1158,1184]" pageId="11" pageNumber="12">Comparison</emphasis>
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. The new tribe has clear differences from other genera of
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<taxonomicName id="4C05B9B3767FD67376A6FB74FBB7FB46" authorityName="Kirejtshuk et Audisio" authorityYear="1986" box="[977,1091,1159,1185]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Cillaeinae">Cillaeinae</taxonomicName>
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defined in the above diagnosis. The most prominent features of the conotelins are in the structure of their abdomen,
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<typeStatus id="54BE7C92767FD6737136FB58FB85FB22" box="[1089,1137,1195,1221]" pageId="11" pageNumber="12">type</typeStatus>
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of intersegmental junction and peculiar male anal sclerite, which are very different from those not only in all other cillaeines, about also in all other nitidulids. The body of the genus
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<taxonomicName id="4C05B9B3767FD6737739FB00FD34FAEA" authorityName="Erichson" authorityYear="1843" box="[590,704,1267,1293]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD6737739FB00FD34FAEA" box="[590,704,1267,1293]" italics="true" pageId="11" pageNumber="12">Conotelus</emphasis>
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</taxonomicName>
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is rather convex from below and more or less convex from above (up to at most only slightly subflattened), but, in contrast to other convex nitidulids, the conotelin abdomen has not any sharp compressed fold along ventrite and laterosternite.
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<bibRefCitation id="EF94BFC1767FD6737603FAC8FBA2FAB2" author="Murray, A." box="[884,1110,1338,1365]" pageId="11" pageNumber="12" pagination="211 - 414" refId="ref107402" refString="Murray, A. (1864) Monograph of the family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414." type="journal article" year="1864">Murray (1864: 256)</bibRefCitation>
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put
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<taxonomicName id="4C05B9B3767FD67371FDFAC8FB08FAB2" authorityName="Erichson" authorityYear="1843" box="[1162,1276,1339,1365]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD67371FDFAC8FB08FAB2" box="[1162,1276,1339,1365]" italics="true" pageId="11" pageNumber="12">Conotelus</emphasis>
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</taxonomicName>
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with
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<taxonomicName id="4C05B9B3767FD6737048FAC8FEDBFA9E" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" class="Insecta" family="Nitidulidae" genus="Ctilodes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD6737048FAC8FA68FAB2" box="[1343,1436,1339,1365]" italics="true" pageId="11" pageNumber="12">Ctilodes</emphasis>
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<bibRefCitation id="EF94BFC1767FD67375E0FAACFEDBFA9E" author="Murray, A." box="[151,303,1375,1401]" pageId="11" pageNumber="12" pagination="211 - 414" refId="ref107402" refString="Murray, A. (1864) Monograph of the family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414." type="journal article" year="1864">Murray, 1864</bibRefCitation>
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</taxonomicName>
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and
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<taxonomicName id="4C05B9B3767FD6737412FAACFD57FA9E" authority="Stephens, 1829" authorityName="Stephens" authorityYear="1829" box="[357,675,1375,1401]" class="Insecta" family="Nitidulidae" genus="Carpophilus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD6737412FAACFE05FA9E" box="[357,497,1375,1401]" italics="true" pageId="11" pageNumber="12">Carpophilus</emphasis>
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Stephens, 1829
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</taxonomicName>
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(including
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<taxonomicName id="4C05B9B3767FD6737653FAACFBB4FA9E" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[804,1088,1375,1401]" class="Insecta" family="Nitidulidae" genus="Urophorus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD6737653FAACFC6AFA9E" box="[804,926,1375,1401]" italics="true" pageId="11" pageNumber="12">Urophorus</emphasis>
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<bibRefCitation id="EF94BFC1767FD67376D2FAACFBB4FA9E" author="Murray, A." box="[933,1088,1375,1401]" pageId="11" pageNumber="12" pagination="211 - 414" refId="ref107402" refString="Murray, A. (1864) Monograph of the family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414." type="journal article" year="1864">Murray, 1864</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C05B9B3767FD673713CFAACFA6CFA9E" authority="Kirejtshuk, 1990 a" authorityName="Kirejtshuk" authorityYear="1990" box="[1099,1432,1375,1401]" class="Insecta" family="Nitidulidae" genus="Anophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD673713CFAACFB32FA9E" box="[1099,1222,1375,1401]" italics="true" pageId="11" pageNumber="12">Anophorus</emphasis>
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<bibRefCitation id="EF94BFC1767FD67371BAFAACFA6CFA9E" author="Kirejtshuk, A. G." box="[1229,1432,1375,1401]" pageId="11" pageNumber="12" pagination="92 - 127" refId="ref105805" refString="Kirejtshuk, A. G. (1990 a) [New taxa of the Nitidulidae) of the Eastern Hemisphere. Part 4.] Tрudы ZooЛogiciсKogo iнсtituta РAН [Proceedings of Zoological Institute of Russian Academy of Sciences], 257, 92 - 127. [in Russian]" type="journal article" year="1990">Kirejtshuk, 1990a</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C05B9B3767FD67375E0FA70FE73FA7A" authority="Murray, 1864" authorityName="Murray" authorityYear="1864" box="[151,391,1411,1437]" class="Insecta" family="Nitidulidae" genus="Nitops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD67375E0FA70FF14FA7B" box="[151,224,1411,1436]" italics="true" pageId="11" pageNumber="12">Nitops</emphasis>
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<bibRefCitation id="EF94BFC1767FD673759DFA70FE73FA7A" author="Murray, A." box="[234,391,1411,1437]" pageId="11" pageNumber="12" pagination="211 - 414" refId="ref107402" refString="Murray, A. (1864) Monograph of the family of Nitidulariae. Part I. Transactions of the Linnean Society of London, 24 (3), 211 - 414." type="journal article" year="1864">Murray, 1864</bibRefCitation>
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</taxonomicName>
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,
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<taxonomicName id="4C05B9B3767FD67374E2FA70FCE0FA7A" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[405,788,1410,1437]" class="Insecta" family="Nitidulidae" genus="Ecnomorphus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD67374E2FA70FDC5FA7A" box="[405,561,1411,1437]" italics="true" pageId="11" pageNumber="12">Ecnomorphus</emphasis>
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Motschulsky, 1858
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</taxonomicName>
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, etc.) in the group “
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<emphasis id="B9711E22767FD6737689FA70FB3EFA7A" box="[1022,1226,1411,1437]" italics="true" pageId="11" pageNumber="12">Anguste-fimbriata</emphasis>
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”, considering that
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<taxonomicName id="4C05B9B3767FD67375E0FA54FEFDFA26" authorityName="Erichson" authorityYear="1843" box="[151,265,1447,1473]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
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<emphasis id="B9711E22767FD67375E0FA54FEFDFA26" box="[151,265,1447,1473]" italics="true" pageId="11" pageNumber="12">Conotelus</emphasis>
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</taxonomicName>
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is intermediate between the genera “
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<emphasis id="B9711E22767FD67377C2FA54FCACFA26" box="[693,856,1447,1473]" italics="true" pageId="11" pageNumber="12">Late-fimbriata</emphasis>
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” (mostly
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<taxonomicName id="4C05B9B3767FD67376BAFA54FBD9FA26" box="[973,1069,1447,1473]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="30" phylum="Arthropoda" rank="tribe" tribe="Cillaeini">Cillaeini</taxonomicName>
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<emphasis id="B9711E22767FD6737140FA54FB80FA27" box="[1079,1140,1447,1472]" italics="true" pageId="11" pageNumber="12">s. str.</emphasis>
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) and “
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<emphasis id="B9711E22767FD67371B3FA54FA64FA26" box="[1220,1424,1447,1473]" italics="true" pageId="11" pageNumber="12">Anguste-fimbriata</emphasis>
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” (mostly
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<taxonomicName id="4C05B9B3767FD6737580FA38FE6FFA01" authorityName="sensu Kirejtshuk" authorityYear="1986" box="[247,411,1483,1510]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Carpophilinae">Carpophilinae</taxonomicName>
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). Indeed, the carpophiline abdomen has the somewhat convex upper sclerotized surface, however, the sides of its “ventrites” turn on the upper surface without visible laterosternites (this peculiarity in the structure of the carpophiline abdomen also represents a rather modified condition and, therefore, the similarity of abdomens of the
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<taxonomicName id="4C05B9B3767FD6737409F9C4FE00F9B6" authority="Kirejtshuk & Kovalev, 2022" authorityName="Kirejtshuk & Kovalev" authorityYear="2022" box="[382,500,1591,1617]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Conotelini">Conotelini</taxonomicName>
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<taxonomicNameLabel id="A242A359767FD673748AF9C5FD99F9B6" box="[509,621,1590,1617]" pageId="11" pageNumber="12" rank="tribe">
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<emphasis id="B9711E22767FD673748AF9C5FD92F9B6" bold="true" box="[509,614,1590,1617]" pageId="11" pageNumber="12">trib. nov</emphasis>
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.
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</taxonomicNameLabel>
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and
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<taxonomicName id="4C05B9B3767FD67377D0F9C4FCB2F9B6" authorityName="sensu Kirejtshuk" authorityYear="1986" box="[679,838,1591,1617]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Carpophilinae">Carpophilinae</taxonomicName>
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are certainly convergent). It is difficult to say if the carpophiline laterosternites are completely reduced or the initial suture between the ventrites and laterosternites are completely fused (become not traced). And as a result, it is impossible to be sure in a strict homology of the upper sclerotized sides externally from abdominal pleurites of the
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<taxonomicName id="4C05B9B3767FD6737640F950FC59F95A" authority="Kirejtshuk & Kovalev, 2022" authorityName="Kirejtshuk & Kovalev" authorityYear="2022" box="[823,941,1699,1725]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="tribe" status="trib. nov." tribe="Conotelini">Conotelini</taxonomicName>
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<taxonomicNameLabel id="A242A359767FD67376C3F951FBD6F95A" box="[948,1058,1698,1725]" pageId="11" pageNumber="12" rank="tribe">
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<emphasis id="B9711E22767FD67376C3F951FBEFF95A" bold="true" box="[948,1051,1698,1725]" pageId="11" pageNumber="12">trib. nov</emphasis>
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.
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</taxonomicNameLabel>
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and
|
||
<taxonomicName id="4C05B9B3767FD673712FF950FB0EF95A" authorityName="sensu Kirejtshuk" authorityYear="1986" box="[1112,1274,1699,1725]" class="Insecta" family="Nitidulidae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Carpophilinae">Carpophilinae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBAC230767FD67375B0F934FB74F8AA" blockId="11.[151,1437,978,2013]" pageId="11" pageNumber="12">
|
||
The labrum of
|
||
<taxonomicName id="4C05B9B3767FD673741AF934FE2BF906" authorityName="Erichson" authorityYear="1843" box="[365,479,1735,1761]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9711E22767FD673741AF934FE2BF906" box="[365,479,1735,1761]" italics="true" pageId="11" pageNumber="12">Conotelus</emphasis>
|
||
</taxonomicName>
|
||
looks like transverse or slightly curved monolyte stripe without any trace of a median suture. The generalized condition of this structure represents two lobes and it has in many groups a trend to close these lobes and in these cases the labrum obtains monolyte outline with a more or less traced suture or remnants of it. In very rare cases among cillaeines the suture between labral lobes became not visible.
|
||
</paragraph>
|
||
<paragraph id="8BBAC230767FD67475B0F8A4FA83FEFA" blockId="11.[151,1437,978,2013]" lastBlockId="12.[151,1436,151,285]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
|
||
Many cillaeines have abdomen with long intersegmental membrane between segments and this circumstance conditions a comparatively great variability in length of abdomen of many species (but not so great as in staphylinids with telescopically folding abdominal segments). Nevertheless, the sclerotized base of abdominal segments of most cillaeines are located at most closely to apex of previous segment, while the abdominal segments of species of
|
||
<taxonomicName id="4C05B9B37678D67475E0FF64FEFDFF56" authorityName="Erichson" authorityYear="1843" box="[151,265,151,177]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9711E227678D67475E0FF64FEFDFF56" box="[151,265,151,177]" italics="true" pageId="12" pageNumber="13">Conotelus</emphasis>
|
||
</taxonomicName>
|
||
are frequently joined telescopically (at least in museum specimens). Among nitidulids somewhat similar condition can be observed in the last abdominal segment of carpophilines. Finally, the ultimate abdominal segment in the new tribe is extremely long and the male anal sclerite of
|
||
<taxonomicName id="4C05B9B37678D6747614FF2CFC21FF1E" authorityName="Erichson" authorityYear="1843" box="[867,981,223,249]" class="Insecta" family="Nitidulidae" genus="Conotelus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="13" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9711E227678D6747614FF2CFC21FF1E" box="[867,981,223,249]" italics="true" pageId="12" pageNumber="13">Conotelus</emphasis>
|
||
</taxonomicName>
|
||
, in conrast to all other nitidulids, is also abnormally long and its ventral lobes are strongly enlarged and forming a long tube for movement of aedeagus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |