287 lines
40 KiB
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287 lines
40 KiB
XML
<document id="4714B80086CDBC7B869A080F4A111449" ID-DOI="10.11646/zootaxa.5351.1.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="8391146" ID-ZooBank="B8858C76-7068-4718-83F4-3AFD00E74249" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695988530166" checkinUser="plazi" docAuthor="Fitzgerald, Scott J." docDate="2023" docId="03AC071073704C09FF7FFF58F8CEFD95" docLanguage="en" docName="zootaxa.5351.1.3.pdf" docOrigin="Zootaxa 5351 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5351.1.3" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macrorrhyncha vockerothi Fitzgerald 2023, n. sp." docType="treatment" docVersion="3" lastPageNumber="101" masterDocId="FF957F68736B4C14FFE8FFCFFB00FFF4" masterDocTitle="The Nearctic species of Asindulum Latreille and Macrorrhyncha Winnertz (Diptera: Keroplatidae)" masterLastPageNumber="106" masterPageNumber="72" pageNumber="99" updateTime="1695990118863" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="4E41BB6C10F901399FB08BF8C9B441B2">The Nearctic species of Asindulum Latreille and Macrorrhyncha Winnertz (Diptera: Keroplatidae)</mods:title>
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<mods:name id="5DA7F13C7FF3ECA3BA3750F6E052B56A" type="personal">
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<mods:namePart id="1B38655018E1E281170FAB5F6B0C0E47">Fitzgerald, Scott J.</mods:namePart>
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<mods:typeOfResource id="AE2678D7C0DE618B767F41DA4BDC8B33">text</mods:typeOfResource>
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<mods:title id="EE482802AF8E25F98C2455D45547D880">Zootaxa</mods:title>
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<mods:part id="6BCA0CA699CF39018878CBC640583191">
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<mods:date id="2CC438F797BFE7276E1D3FD451A6F492">2023</mods:date>
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<mods:number id="C4E58609A285248A0C66785A2496F345">2023-09-25</mods:number>
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<mods:number id="8390B88334EA339A25A336EDB80DC6BF">5351</mods:number>
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<mods:start id="398AE0A6C0D90C83D173B9A346856679">72</mods:start>
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<mods:end id="256643238BE8B77D194A97C128C20E0F">106</mods:end>
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<mods:url id="1951435E6E90B84914CE90460B428440">http://dx.doi.org/10.11646/zootaxa.5351.1.3</mods:url>
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<mods:classification id="E791DE9597CD634AEA951A49147581B1">journal article</mods:classification>
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<mods:identifier id="414830A1AAAB73176C94A5D9EF68567A" type="DOI">10.11646/zootaxa.5351.1.3</mods:identifier>
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<mods:identifier id="BD106174DEB3872D029375B6FE32169C" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="D675393F54F4A93EC1B377998D303D6D" type="ZooBank">B8858C76-7068-4718-83F4-3AFD00E74249</mods:identifier>
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<treatment id="03AC071073704C09FF7FFF58F8CEFD95" ID-DOI="http://doi.org/10.5281/zenodo.8391493" ID-Zenodo-Dep="8391493" LSID="urn:lsid:plazi:treatment:03AC071073704C09FF7FFF58F8CEFD95" httpUri="http://treatment.plazi.org/id/03AC071073704C09FF7FFF58F8CEFD95" lastPageId="29" lastPageNumber="101" pageId="27" pageNumber="99">
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<subSubSection id="C31FE58D73704C0FFF7FFF58F9B0FF45" box="[151,688,151,178]" pageId="27" pageNumber="99" type="nomenclature">
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<paragraph id="8BBAB60673704C0FFF7FFF58F9B0FF45" blockId="27.[151,961,151,250]" box="[151,688,151,178]" pageId="27" pageNumber="99">
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<heading id="D0F2016A73704C0FFF7FFF58F9B0FF45" bold="true" box="[151,688,151,178]" fontSize="11" level="1" pageId="27" pageNumber="99" reason="1">
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<emphasis id="B9716A1473704C0FFF7FFF58F9B0FF45" bold="true" box="[151,688,151,178]" pageId="27" pageNumber="99">
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<taxonomicName id="4C05CD8573704C0FFF7FFF58F966FF45" authority="Fitzgerald" authorityName="Fitzgerald" authorityYear="2023" box="[151,614,151,178]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="99" phylum="Arthropoda" rank="species" species="vockerothi" status="n. sp.">
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<emphasis id="B9716A1473704C0FFF7FFF58FADEFF46" bold="true" box="[151,478,151,178]" italics="true" pageId="27" pageNumber="99">Macrorrhyncha vockerothi</emphasis>
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Fitzgerald
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</taxonomicName>
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<taxonomicNameLabel id="A242D76F73704C0FFD85FF58F9B0FF45" box="[621,688,151,177]" pageId="27" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FE58D73704C0FFF7FFF74F8C1FF0E" pageId="27" pageNumber="99" type="description">
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<paragraph id="8BBAB60673704C0FFF7FFF74FA1CFF22" blockId="27.[151,961,151,250]" box="[151,284,187,214]" pageId="27" pageNumber="99">
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<figureCitation id="133EAA8373704C0FFF7FFF74FA1CFF22" box="[151,284,187,214]" captionStart-0="FIGURES 75–77" captionStart-1="FIGURES 78–81" captionStart-2="FIGURES 82–83" captionStartId-0="28.[151,264,1723,1747]" captionStartId-1="29.[151,264,1900,1924]" captionStartId-2="30.[151,264,1325,1349]" captionTargetBox-0="[336,1252,250,1687]" captionTargetBox-1="[302,1304,693,1864]" captionTargetBox-2="[220,1366,201,1288]" captionTargetId-0="figure-40@28.[323,1264,237,1700]" captionTargetId-1="figure-256@29.[268,1320,633,1877]" captionTargetId-2="figure-15@30.[207,1379,181,1301]" captionTargetPageId-0="28" captionTargetPageId-1="29" captionTargetPageId-2="30" captionText-0="FIGURES 75–77. Macrorrhyncha vockerothi n. sp., male. 75. Holotype, habitus. 76. Head and anterior thorax with box indicating area of enlarged anterior spiracle. 77.Anterior spiracle. Scale = ca. 1.0 mm (FIGS. 76–77 not to scale).Abbreviations: asp, anterior spiracle; cx1, coxa 1; CuP, posterior branch of cubital vein." captionText-1="FIGURES 78–81. Macrorrhyncha vockerothi n. sp., holotype male, terminalia. 67. Dorsal. 68. Ventral. 69. Lateral. 70. Posterior with inset showing enlarged gonostylus (inset not to scale). Scale bar = ca. 0.25 mm.Abbreviations: bl, basal lobe of gonostylus; ce, cercus; dp, dorsal process of gonocoxite; gc, gonocoxite; gs, gonostylus; ip, inner process of gonocoxite; s8, sternite 8; t8, tergite 8; t9, tergite 9." captionText-2="FIGURES 82–83. Macrorrhyncha vockerothi n. sp., holotype male, terminalia including sperm pump. 82. Lateral. 83. Ventral. Scale = ca. 0.25 mm. Abbreviations: da, dorsal apodeme; ea, ejaculatory apodeme; gc, gonocoxite; t9, tergite 9; va, ventral apodeme;?, short divergent dorsal pair of apodemes (see Discussion)." figureDoi-0="http://doi.org/10.5281/zenodo.8391209" figureDoi-1="http://doi.org/10.5281/zenodo.8391211" figureDoi-2="http://doi.org/10.5281/zenodo.8391213" httpUri-0="https://zenodo.org/record/8391209/files/figure.png" httpUri-1="https://zenodo.org/record/8391211/files/figure.png" httpUri-2="https://zenodo.org/record/8391213/files/figure.png" pageId="27" pageNumber="99">Figs. 75–83</figureCitation>
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</paragraph>
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<paragraph id="8BBAB60673704C0FFF7FFF10F8C1FF0E" blockId="27.[151,961,151,250]" box="[151,961,223,250]" pageId="27" pageNumber="99">
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<uri id="FF94BA0473704C0FFF7FFF10F8C1FF0E" box="[151,961,223,250]" pageId="27" pageNumber="99">
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urn:lsid:zoobank.org:act:
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<uuid id="FFA38CD373704C0FFE44FF10F8C1FF0E" box="[428,961,223,250]" pageId="27" pageNumber="99">98573C85-7462-42F9-B2B3-921D67A5A7CA</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31FE58D73704C0FFF7FFEE8FF60FE91" pageId="27" pageNumber="99" type="materials_examined">
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<paragraph id="8BBAB60673704C0FFF7FFEE8FF60FE91" blockId="27.[151,1437,295,2014]" pageId="27" pageNumber="99">
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<emphasis id="B9716A1473704C0FFF7FFEE8FA45FEB5" bold="true" box="[151,325,295,321]" pageId="27" pageNumber="99">Type Material</emphasis>
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.
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<materialsCitation id="3B6DBC5B73704C0FFEBFFEE8F946FE91" collectingDate="1961-08-30" collectionCode="CNCI, R" country="Canada" location="Old Chelsea" pageId="27" pageNumber="99" specimenCount="1" stateProvince="Quebec" typeStatus="holotype">
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<typeStatus id="54BE08A473704C0FFEBFFEE8FABFFEB5" box="[343,447,295,321]" pageId="27" pageNumber="99">Holotype</typeStatus>
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male (
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<collectionCode id="ED142EC373704C0FFDFCFEE8F95CFEB5" box="[532,604,295,321]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="99">CNCI</collectionCode>
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), glued to side of pin, terminalia dissected:
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<collectingCountry id="F312F69673704C0FFB8DFEE8FFE1FEB5" box="[1125,1249,295,321]" name="Canada" pageId="27" pageNumber="99">CANADA</collectingCountry>
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:
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<collectingRegion id="49C178E473704C0FFB07FEE8FE63FEB5" box="[1263,1379,295,321]" country="Canada" name="Quebec" pageId="27" pageNumber="99">QUEBEC</collectingRegion>
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:
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<location id="8EDAE0DD73704C0FFA9AFEE8FBF3FE91" LSID="urn:lsid:plazi:treatment:03AC071073704C09FF7FFF58F8CEFD95:8EDAE0DD73704C0FFA9AFEE8FBF3FE91" country="Canada" name="Old Chelsea" pageId="27" pageNumber="99" stateProvince="Quebec">Old Chelsea</location>
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,
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<date id="FFBB90C673704C0FFF16FE84FA91FE91" box="[254,401,331,358]" pageId="27" pageNumber="99" value="1961-08-30">
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<collectingDate id="EFFF692E73704C0FFF16FE84FA91FE91" box="[254,401,331,358]" pageId="27" pageNumber="99" value="1961-08-30">30 VIII 1961</collectingDate>
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</date>
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, J.
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<collectionCode id="ED142EC373704C0FFE46FE83FAC5FE92" box="[430,453,332,358]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="27" pageNumber="99">R</collectionCode>
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. Vockeroth
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</materialsCitation>
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.
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<materialsCitation id="3B6DBC5B73704C0FFDB8FE84FF5CFE91" box="[592,1116,331,358]" collectionCode="CNCI" country="Canada" pageId="27" pageNumber="99" specimenCount="2" specimenCount-male="2" stateProvince="Quebec" typeStatus="paratype">
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<emphasis id="B9716A1473704C0FFDB8FE84F9C8FE91" bold="true" box="[592,712,331,357]" pageId="27" pageNumber="99">
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||
<typeStatus id="54BE08A473704C0FFDB8FE84F9C8FE91" box="[592,712,331,357]" pageId="27" pageNumber="99">Paratypes</typeStatus>
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</emphasis>
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: same as
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<typeStatus id="54BE08A473704C0FFCDEFE84F89BFE91" box="[822,923,331,357]" pageId="27" pageNumber="99">holotype</typeStatus>
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,
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<specimenCount id="9D037D8F73704C0FFC4EFE84F8FCFE91" box="[934,1020,331,357]" pageId="27" pageNumber="99" type="male">2 males</specimenCount>
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(
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<collectionCode id="ED142EC373704C0FFBE3FE84FF54FE91" box="[1035,1108,331,357]" country="Canada" httpUri="http://grbio.org/cool/rzcb-tdp2" name="Canadian National Collection Insects" pageId="27" pageNumber="99">CNCI</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C31FE58D73704C0FFF2FFEA0FA9EF96D" pageId="27" pageNumber="99" type="description">
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<paragraph id="8BBAB60673704C0FFF2FFEA0FA14F981" blockId="27.[151,1437,295,2014]" pageId="27" pageNumber="99">
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<emphasis id="B9716A1473704C0FFF2FFEA0FA51FE7D" bold="true" box="[199,337,367,393]" pageId="27" pageNumber="99">Description</emphasis>
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. Male. Body length ca. 3.5 mm (n = 1). Entirely cream to yellowish in color except, head, antennal flagellomeres, tarsi and apex of abdomen light brown. Head brown, antennae largely brown with base of basal flagellomere and sometimes pedicel and scape cream, mouthparts light brown with palps becoming lighter (cream) distally. Antennae with 14 cylindrical, barrel-shaped flagellomeres plus a minute, apical, button-like apicule, all without distinct setae. Mouthparts as
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<figureCitation id="133EAA8373704C0FFD85FE30F9C3FDED" box="[621,707,511,537]" captionStart="FIGURES 75–77" captionStartId="28.[151,264,1723,1747]" captionTargetBox="[336,1252,250,1687]" captionTargetId="figure-40@28.[323,1264,237,1700]" captionTargetPageId="28" captionText="FIGURES 75–77. Macrorrhyncha vockerothi n. sp., male. 75. Holotype, habitus. 76. Head and anterior thorax with box indicating area of enlarged anterior spiracle. 77.Anterior spiracle. Scale = ca. 1.0 mm (FIGS. 76–77 not to scale).Abbreviations: asp, anterior spiracle; cx1, coxa 1; CuP, posterior branch of cubital vein." figureDoi="http://doi.org/10.5281/zenodo.8391209" httpUri="https://zenodo.org/record/8391209/files/figure.png" pageId="27" pageNumber="99">Fig. 76</figureCitation>
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; long, terminating approximately at level of apex of fore coxa. Palps five-segmented. Three centrally-positioned ocelli in an arc; median ocellus smaller. Mesonotum cream to yellowish with hint of three broad light brown longitudinal stripes, with short black setae in broad irregular rows medially, dorsocentrally, and laterally. Thoracic pleura bright cream to yellowish with some light brownish coloration; anepisternum, katepisternum, laterotergite, and mediotergite bare. Posterior edge of anterior spiracle with several short black setae. Coxae bright cream to yellowish, remainder of legs cream, becoming progressively more light brown distally (tarsi entirely light brown). Tibial spurs 1:2:2 with inner spurs longer. Fore tibia with apical, triangular sensory area on anterior surface. Hind tibia with trichia irregularly arranged and a small number of short black setae in regular rows on anteroventral, anterior, dorsal, posterodorsal, posterior, and posteroventral surfaces. Wing ca. 3.5 mm (n = 1), hyaline. Veins brown, anterior and posterior forks with short, closely-spaced setae on upper surface, especially apically. Anterior veins with closely-spaced setae except Sc, base Rs, and R
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<subScript id="1781B44373704C0FFA6BFCB5FE9CFC7C" attach="left" box="[1411,1436,890,904]" fontSize="6" pageId="27" pageNumber="99">2+3</subScript>
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bare. CuP with a few sparse setae, strong basally, but quickly becoming nearly transparent; in some specimens slightly traceable all the way to wing margin as a subtle crease. Halters yellowish to light brown. Abdomen bright cream to yellowish (tergites 1–2 entirely so) with brown tinge over anterior parts of tergites or sometimes over entire tergite with brown coloration becoming more pronounced and darker distally, culminating with tergites 7–9 brown. Male terminalia as
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<figureCitation id="133EAA8373704C0FFE2BFBD4F94BFBC1" box="[451,587,1051,1077]" captionStart-0="FIGURES 78–81" captionStart-1="FIGURES 82–83" captionStartId-0="29.[151,264,1900,1924]" captionStartId-1="30.[151,264,1325,1349]" captionTargetBox-0="[302,1304,693,1864]" captionTargetBox-1="[220,1366,201,1288]" captionTargetId-0="figure-256@29.[268,1320,633,1877]" captionTargetId-1="figure-15@30.[207,1379,181,1301]" captionTargetPageId-0="29" captionTargetPageId-1="30" captionText-0="FIGURES 78–81. Macrorrhyncha vockerothi n. sp., holotype male, terminalia. 67. Dorsal. 68. Ventral. 69. Lateral. 70. Posterior with inset showing enlarged gonostylus (inset not to scale). Scale bar = ca. 0.25 mm.Abbreviations: bl, basal lobe of gonostylus; ce, cercus; dp, dorsal process of gonocoxite; gc, gonocoxite; gs, gonostylus; ip, inner process of gonocoxite; s8, sternite 8; t8, tergite 8; t9, tergite 9." captionText-1="FIGURES 82–83. Macrorrhyncha vockerothi n. sp., holotype male, terminalia including sperm pump. 82. Lateral. 83. Ventral. Scale = ca. 0.25 mm. Abbreviations: da, dorsal apodeme; ea, ejaculatory apodeme; gc, gonocoxite; t9, tergite 9; va, ventral apodeme;?, short divergent dorsal pair of apodemes (see Discussion)." figureDoi-0="http://doi.org/10.5281/zenodo.8391211" figureDoi-1="http://doi.org/10.5281/zenodo.8391213" httpUri-0="https://zenodo.org/record/8391211/files/figure.png" httpUri-1="https://zenodo.org/record/8391213/files/figure.png" pageId="27" pageNumber="99">Figs. 78–83</figureCitation>
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. Male tergite 9 reduced, posterior margin with broad v-shaped emargination which nearly or entirely subdivides the sclerite into two small triangular sclerites which are closely appressed to tergite 8. In dorsal view, cerci prominent, triangular, fleshy, setose. Ventrally, gonocoxites with deep, broad u-shaped emargination which nearly subdivides sclerite; a narrow strip of sclerite anteriorly connecting the two halves. Inner process of gonocoxite, short, digitate, bearing a strong apical seta. In lateral view, gonocoxites short, broad, apically truncate, with a narrow, short, ventrally curved, asetose, apically rounded dorsal process that is longer than (i.e. reaches beyond the level of the posterior apex of) the primary lobe of the gonocoxite. Gonostylus cushion-shaped, apically (posteromedial surface) broadly truncate with four strong apically truncate setae. Gonostylus with a minute, nubbin-like basal lobe originating from medial surface; apex of basal lobe with two tiny seta (basal lobe difficult to observe; originates at about same position of origin as inner process of gonocoxite so that it lies hidden between main lobe of gonostylus and inner process of gonocoxite). Sperm pump (possibly fused with parameres?) large and anteriorly elongated, with apodemes reaching into abdominal segment 6; notable are two pairs of long apodemes (one nearly transparent pair more ventral and laterally flattened and the second pair more dorsal and rod-like), an unpaired median ejaculatory apodeme (which is the longest of all the apodemes and the most strongly laterally compressed) and a pair of short, divergent apodemes dorsally. The posterior apex of the sperm pump is difficult to discern, but includes a pair of dorsoventrally-flattened lobes (visible protruding above the thin strap of gonocoxite in
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<figureCitation id="133EAA8373704C0FFF5CF994FA06F981" box="[180,262,1627,1653]" captionStart="FIGURES 82–83" captionStartId="30.[151,264,1325,1349]" captionTargetBox="[220,1366,201,1288]" captionTargetId="figure-15@30.[207,1379,181,1301]" captionTargetPageId="30" captionText="FIGURES 82–83. Macrorrhyncha vockerothi n. sp., holotype male, terminalia including sperm pump. 82. Lateral. 83. Ventral. Scale = ca. 0.25 mm. Abbreviations: da, dorsal apodeme; ea, ejaculatory apodeme; gc, gonocoxite; t9, tergite 9; va, ventral apodeme;?, short divergent dorsal pair of apodemes (see Discussion)." figureDoi="http://doi.org/10.5281/zenodo.8391213" httpUri="https://zenodo.org/record/8391213/files/figure.png" pageId="27" pageNumber="99">Fig. 83</figureCitation>
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).
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</paragraph>
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<paragraph id="8BBAB60673704C0FFF2FF9B0FA9EF96D" blockId="27.[151,1437,295,2014]" box="[199,414,1663,1689]" pageId="27" pageNumber="99">Female. Unknown.</paragraph>
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</subSubSection>
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<subSubSection id="C31FE58D73704C08FF2FF96CFA10FF21" lastPageId="28" lastPageNumber="100" pageId="27" pageNumber="99" type="etymology">
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<paragraph id="8BBAB60673704C08FF2FF96CFA10FF21" blockId="27.[151,1437,295,2014]" lastBlockId="28.[151,1436,151,213]" lastPageId="28" lastPageNumber="100" pageId="27" pageNumber="99">
|
||
<emphasis id="B9716A1473704C0FFF2FF96CFA48F949" bold="true" box="[199,328,1699,1725]" pageId="27" pageNumber="99">Etymology</emphasis>
|
||
. This species is named in honor and memory of Richard (Dick) Vockeroth, collector of the
|
||
<typeStatus id="54BE08A473704C0FFAABF96BFE7EF94A" box="[1347,1406,1700,1726]" pageId="27" pageNumber="99">types</typeStatus>
|
||
of this species as well as specimens of
|
||
<taxonomicName id="4C05CD8573704C0FFDD8F908F9B2F915" authority="Fitzgerald, 2023" authorityName="Fitzgerald" authorityYear="2023" box="[560,690,1735,1761]" class="Insecta" family="Keroplatidae" genus="Asindulum" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="99" phylum="Arthropoda" rank="species" species="flavidum" status="n. sp.">
|
||
<emphasis id="B9716A1473704C0FFDD8F908F9B2F915" box="[560,690,1735,1761]" italics="true" pageId="27" pageNumber="99">A. flavidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9716A1473704C0FFD51F908F9F9F915" bold="true" box="[697,761,1735,1761]" pageId="27" pageNumber="99">
|
||
<taxonomicNameLabel id="A242D76F73704C0FFD51F908F9F9F915" box="[697,761,1735,1761]" pageId="27" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05CD8573704C0FFCD8F907F8B1F915" authority="Fitzgerald, 2023" authorityName="Fitzgerald" authorityYear="2023" box="[816,945,1735,1761]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="99" phylum="Arthropoda" rank="species" species="borealis" status="n. sp.">
|
||
<emphasis id="B9716A1473704C0FFCD8F907F8B1F915" box="[816,945,1735,1761]" italics="true" pageId="27" pageNumber="99">M. borealis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9716A1473704C0FFC51F908F8F8F915" bold="true" box="[953,1016,1735,1761]" pageId="27" pageNumber="99">
|
||
<taxonomicNameLabel id="A242D76F73704C0FFC51F908F8F8F915" box="[953,1016,1735,1761]" pageId="27" pageNumber="99" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
While Dick contributed significantly to the study of fungus gnats both through avid collecting and a number of important papers and book chapters, I remember him most for the personal impact he made on me. I first met Dick at the Third International
|
||
<taxonomicName id="4C05CD8573704C0FFAA0F8C0FBFFF8B9" authority="Congress" authorityName="Congress" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Diptera" pageId="27" pageNumber="99" phylum="Tracheophyta" rank="order">Diptera Congress</taxonomicName>
|
||
in Guelph,
|
||
<collectingCountry id="F312F69673704C0FFE60F8FCFADCF8B9" box="[392,476,1843,1869]" name="Canada" pageId="27" pageNumber="99">Canada</collectingCountry>
|
||
in 1994. I had just arrived at the dorm after hitchhiking from the airport, duffle bag in hand, and had not even found my room yet, when I met Dick, who had clearly already been out collecting, coming down the hall wearing a belt filled with small aspirator tubes with gnats in all of them. At the time, I was an impressionable student preparing to start a master’s degree program working on
|
||
<taxonomicName id="4C05CD8573704C0FFBB5F850FFDCF84D" box="[1117,1244,1951,1977]" class="Insecta" family="Bibionidae" kingdom="Animalia" order="Diptera" pageId="27" pageNumber="99" phylum="Arthropoda" rank="family">Bibionidae</taxonomicName>
|
||
; Dick shared his observations of bibionids and pachyneurids, and encouraged and inspired me to look at fungus gnats as well (which I had largely ignored up to that point). Fast-forward 29 years and I am happy to be able to name this fungus gnat after Dick.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF7AE68E73774C08FF7FF974F873F8E8" ID-DOI="http://doi.org/10.5281/zenodo.8391209" ID-Zenodo-Dep="8391209" httpUri="https://zenodo.org/record/8391209/files/figure.png" pageId="28" pageNumber="100" startId="28.[151,264,1723,1747]" targetBox="[336,1252,250,1687]" targetPageId="28" targetType="figure">
|
||
<paragraph id="8BBAB60673774C08FF7FF974F873F8E8" blockId="28.[151,1437,1723,1820]" pageId="28" pageNumber="100">
|
||
<emphasis id="B9716A1473774C08FF7FF974FA58F920" bold="true" box="[151,344,1723,1748]" pageId="28" pageNumber="100">FIGURES 75–77.</emphasis>
|
||
<taxonomicName id="4C05CD8573774C08FE8AF973F975F920" authority="Fitzgerald, 2023" authorityName="Fitzgerald" authorityYear="2023" box="[354,629,1724,1748]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="100" phylum="Arthropoda" rank="species" species="vockerothi" status="n. sp.">
|
||
<emphasis id="B9716A1473774C08FE8AF973F975F920" box="[354,629,1724,1748]" italics="true" pageId="28" pageNumber="100">Macrorrhyncha vockerothi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9716A1473774C08FD68F973F9BDF920" bold="true" box="[640,701,1724,1748]" pageId="28" pageNumber="100">
|
||
<taxonomicNameLabel id="A242D76F73774C08FD68F973F9BDF920" box="[640,701,1724,1748]" pageId="28" pageNumber="100" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male. 75. Holotype, habitus. 76. Head and anterior thorax with box indicating area of enlarged anterior spiracle. 77. Anterior spiracle. Scale = ca. 1.0 mm (FIGS. 76–77 not to scale). Abbreviations: asp, anterior spiracle; cx1, coxa 1; CuP, posterior branch of cubital vein.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C31FE58D73774C08FF2FF885FF00F824" pageId="28" pageNumber="100" type="diagnosis">
|
||
<paragraph id="8BBAB60673774C08FF2FF885FF00F824" blockId="28.[151,1436,1866,2036]" pageId="28" pageNumber="100">
|
||
<emphasis id="B9716A1473774C08FF2FF885FA3AF890" bold="true" box="[199,314,1866,1892]" pageId="28" pageNumber="100">Diagnosis</emphasis>
|
||
. Dorsal process of gonocoxites well-developed (longer than main lobe of gonocoxite), narrow, apically rounded; gonostylus cushion-shaped with several strong setae and a minute basal process; inner process of gonocoxite short, digitate. The female is unknown, but within the
|
||
<taxonomicName id="4C05CD8573774C08FC92F85DF8D9F858" baseAuthorityName="Evenhuis" baseAuthorityYear="2006" box="[890,985,1938,1964]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="106" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
|
||
Region it should be recognizable by the shorter mouthparts (terminating near apex of fore coxa) and yellowish thorax.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31FE58D73774C09FF2FF815FF2FFDC9" lastPageId="29" lastPageNumber="101" pageId="28" pageNumber="100" type="discussion">
|
||
<paragraph id="8BBAB60673774C09FF2FF815FE97FE25" blockId="28.[151,1436,1866,2036]" lastBlockId="29.[151,1437,151,609]" lastPageId="29" lastPageNumber="101" pageId="28" pageNumber="100">
|
||
<emphasis id="B9716A1473774C08FF2FF815FA47F800" bold="true" box="[199,327,2010,2036]" pageId="28" pageNumber="100">Comments</emphasis>
|
||
. It is possible that
|
||
<taxonomicName id="4C05CD8573774C08FDDCF814F9D2F800" authorityName="Fitzgerald" authorityYear="2023" box="[564,722,2010,2036]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="100" phylum="Arthropoda" rank="species" species="vockerothi">
|
||
<emphasis id="B9716A1473774C08FDDCF814F9D2F800" box="[564,722,2010,2036]" italics="true" pageId="28" pageNumber="100">M. vockerothi</emphasis>
|
||
</taxonomicName>
|
||
is the same species that was previously reported from
|
||
<collectingRegion id="49C178E473774C09FA80F815FA14FF45" country="United States of America" lastPageId="29" lastPageNumber="101" name="New Hampshire" pageId="28" pageNumber="100">New Hampshire</collectingRegion>
|
||
,
|
||
<collectingCountry id="F312F69673764C09FEF6FF58FA54FF45" box="[286,340,151,177]" name="United States of America" pageId="29" pageNumber="101">USA</collectingCountry>
|
||
as “
|
||
<taxonomicName id="4C05CD8573764C09FE6BFF58F9BAFF45" authority="Winnertz" authorityName="Winnertz" authorityYear="1846" box="[387,698,151,177]" class="Insecta" family="Keroplatidae" genus="Asindulum" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="flavum">
|
||
<emphasis id="B9716A1473764C09FE6BFF58F94BFF45" box="[387,587,151,177]" italics="true" pageId="29" pageNumber="101">Asindulum flavum</emphasis>
|
||
Winnertz
|
||
</taxonomicName>
|
||
” by
|
||
<bibRefCitation id="EF94CBF773764C09FD03FF58F8AEFF45" author="Johannsen, O. A." box="[747,942,151,177]" pageId="29" pageNumber="101" pagination="209 - 276" refId="ref16451" refString="Johannsen, O. A. (1910) The fungus gnats of North America. The Mycetophilidae of North America. Part I. Bulletin of the Maine Agricultural Experiment Station, 172 [1909], 209 - 276." type="journal article" year="1910">Johannsen (1910)</bibRefCitation>
|
||
as it matches Johannsen’s description. Later,
|
||
<bibRefCitation id="EF94CBF773764C09FF7FFF74FA40FF21" author="Laffoon, J. L." box="[151,320,186,213]" pageId="29" pageNumber="101" pagination="196 - 204" refId="ref16527" refString="Laffoon, J. L. (1965) Family Mycetophilidae (Fungivoridae). In: Stone, A., Sabrosky, C. W., Wirth, W. W., Foote, R. H. & Coulson, J. R. (Eds.), A catalog of the Diptera of America North of Mexico. Smithsonian Institution Press, Washington, D. C., pp. 196 - 204." type="book chapter" year="1965">Laffoon (1965)</bibRefCitation>
|
||
records presumably the same taxon (as
|
||
<taxonomicName id="4C05CD8573764C09FD1FFF74F8E7FF21" authority="(Tarwid)" baseAuthorityName="Tarwid" baseAuthorityYear="1936" box="[759,999,187,213]" class="Insecta" family="Keroplatidae" genus="Asindulum" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="winnertzi">
|
||
<emphasis id="B9716A1473764C09FD1FFF74F87CFF20" box="[759,892,187,213]" italics="true" pageId="29" pageNumber="101">A. winnertzi</emphasis>
|
||
(Tarwid)
|
||
</taxonomicName>
|
||
; now a jun. syn. of “
|
||
<taxonomicName id="4C05CD8573764C09FB20FF74FE13FF21" authorityName="Winnertz" authorityYear="1846" box="[1224,1299,187,213]" class="Insecta" family="Keroplatidae" genus="Asindulum" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="flavum">
|
||
<emphasis id="B9716A1473764C09FB20FF74FE13FF21" box="[1224,1299,187,213]" italics="true" pageId="29" pageNumber="101">flavum</emphasis>
|
||
</taxonomicName>
|
||
”) from both
|
||
<collectingRegion id="49C178E473764C09FF7FFF10FA4DFF0D" box="[151,333,223,249]" country="United States of America" name="New Hampshire" pageId="29" pageNumber="101">New Hampshire</collectingRegion>
|
||
and
|
||
<collectingRegion id="49C178E473764C09FE6DFF10F931FF0D" box="[389,561,223,249]" country="United States of America" name="North Carolina" pageId="29" pageNumber="101">North Carolina</collectingRegion>
|
||
. However, no specimens of Winnertz’s European species, originally described and currently placed in the genus
|
||
<taxonomicName id="4C05CD8573764C09FDF5FECCF9CBFEE9" box="[541,715,259,285]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9716A1473764C09FDF5FECCF9CBFEE9" box="[541,715,259,285]" italics="true" pageId="29" pageNumber="101">Macrorrhyncha</emphasis>
|
||
</taxonomicName>
|
||
, were found in
|
||
<taxonomicName id="4C05CD8573764C09FC6CFECCF8E2FEE9" baseAuthorityName="Evenhuis" baseAuthorityYear="2006" box="[900,994,259,285]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="106" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
|
||
material studied, suggesting that these previous records appear to be based on misidentifications. The only mostly-yellow
|
||
<taxonomicName id="4C05CD8573764C09FBC1FEE8FFA0FEB5" box="[1065,1184,295,321]" class="Insecta" family="Keroplatidae" genus="Asindulum" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9716A1473764C09FBC1FEE8FFA0FEB5" box="[1065,1184,295,321]" italics="true" pageId="29" pageNumber="101">Asindulum</emphasis>
|
||
</taxonomicName>
|
||
/
|
||
<taxonomicName id="4C05CD8573764C09FB4FFEE8FE55FEB5" box="[1191,1365,295,321]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9716A1473764C09FB4FFEE8FE55FEB5" box="[1191,1365,295,321]" italics="true" pageId="29" pageNumber="101">Macrorrhyncha</emphasis>
|
||
</taxonomicName>
|
||
found in
|
||
<taxonomicName id="4C05CD8573764C09FF5AFE84FA10FE91" baseAuthorityName="Evenhuis" baseAuthorityYear="2006" box="[178,272,331,357]" class="Insecta" family="Carabidae" genus="Nearctic" kingdom="Animalia" order="Coleoptera" pageId="29" pageNumber="106" phylum="Arthropoda" rank="genus">Nearctic</taxonomicName>
|
||
specimens studied are
|
||
<taxonomicName id="4C05CD8573764C09FDE4FE84F98AFE91" authority="Fitzgerald, 2023" authorityName="Fitzgerald" authorityYear="2023" box="[524,650,331,357]" class="Insecta" family="Keroplatidae" genus="Asindulum" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="flavidum" status="n. sp.">
|
||
<emphasis id="B9716A1473764C09FDE4FE84F98AFE91" box="[524,650,331,357]" italics="true" pageId="29" pageNumber="101">A. flavidum</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9716A1473764C09FD78FE84F9CDFE91" bold="true" box="[656,717,331,357]" pageId="29" pageNumber="101">
|
||
<taxonomicNameLabel id="A242D76F73764C09FD78FE84F9CDFE91" box="[656,717,331,357]" pageId="29" pageNumber="101" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C05CD8573764C09FCE8FE84F898FE91" authority="Fitzgerald, 2023" authorityName="Fitzgerald" authorityYear="2023" box="[768,920,331,357]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="vockerothi" status="n. sp.">
|
||
<emphasis id="B9716A1473764C09FCE8FE84F898FE91" box="[768,920,331,357]" italics="true" pageId="29" pageNumber="101">M. vockerothi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9716A1473764C09FC76FE84F8DAFE91" bold="true" box="[926,986,331,357]" pageId="29" pageNumber="101">
|
||
<taxonomicNameLabel id="A242D76F73764C09FC76FE84F8DAFE91" box="[926,986,331,357]" pageId="29" pageNumber="101" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, both with very different male terminalia from
|
||
<taxonomicName id="4C05CD8573764C09FF3DFEA0FA9EFE7D" authority="Winnertz" authorityName="Winnertz" authorityYear="1846" box="[213,414,367,393]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="flava">
|
||
<emphasis id="B9716A1473764C09FF3DFEA0FA31FE7D" box="[213,305,367,393]" italics="true" pageId="29" pageNumber="101">M. flava</emphasis>
|
||
Winnertz
|
||
</taxonomicName>
|
||
(see
|
||
<bibRefCitation id="EF94CBF773764C09FE32FEA0F966FE7D" author="Matile, L." box="[474,614,366,393]" pageId="29" pageNumber="101" pagination="173 - 220" refId="ref16862" refString="Matile, L. (1975) Revision des Asindulum et des Macrorrhyncha de la region Palearctique [Dipt., Mycetophilidae]. Annales de la Societe Entomologique de France (n. s.), 11 (1), 173 - 220." type="journal article" year="1975">Matile 1975</bibRefCitation>
|
||
). In agreement with
|
||
<bibRefCitation id="EF94CBF773764C09FCA3FEA0FF06FE7D" author="Evenhuis, N. L." box="[843,1030,367,393]" pageId="29" pageNumber="101" pagination="1 - 178" refId="ref16071" refString="Evenhuis, N. L. (2006) Catalog of the Keroplatidae of the World (Insecta: Diptera). Bishop Museum Bulletin of Entomology, 13, 1 - 178." type="journal article" year="2006">Evenhuis (2006)</bibRefCitation>
|
||
, it is the present authors’ conclusion that
|
||
<taxonomicName id="4C05CD8573764C09FF20FE5CFA25FE59" authorityName="Winnertz" authorityYear="1846" box="[200,293,403,429]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="flava">
|
||
<emphasis id="B9716A1473764C09FF20FE5CFA25FE59" box="[200,293,403,429]" italics="true" pageId="29" pageNumber="101">M. flava</emphasis>
|
||
</taxonomicName>
|
||
is restricted to the Palearctic region. In Palearctic keys,
|
||
<taxonomicName id="4C05CD8573764C09FC71FE5CFF33FE59" authorityName="Fitzgerald" authorityYear="2023" box="[921,1075,403,429]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="vockerothi">
|
||
<emphasis id="B9716A1473764C09FC71FE5CFF33FE59" box="[921,1075,403,429]" italics="true" pageId="29" pageNumber="101">M. vockerothi</emphasis>
|
||
</taxonomicName>
|
||
will key to couplet 15 (does not really fit either choice) in
|
||
<bibRefCitation id="EF94CBF773764C09FE50FE78F96AFE25" author="Bechev, D." box="[440,618,439,465]" pageId="29" pageNumber="101" pagination="335 - 345" refId="ref15654" refString="Bechev, D. (2010 b) Two new species of Macrorrhyncha Winnertz (Diptera: Keropolatidae) from Turkey, with redescription of Macrorrhyncha thracica Bechev, stat. n. and a key to the Western Palearctic Species of the Genus. Acta Zoologica Academiae Scientiarum Hungaricae, 56 (4), 335 - 345." type="journal article" year="2010">Bechev (2010b)</bibRefCitation>
|
||
and couplet 6 (does not really fit either choice) in
|
||
<bibRefCitation id="EF94CBF773764C09FB73FE78FE93FE25" author="Chandler, P. J. & Bechev, D. N. & Caspers, N." box="[1179,1427,438,465]" pageId="29" pageNumber="101" pagination="255 - 314" refId="ref15936" refString="Chandler, P. J., Bechev, D. N., & Caspers, N. (2005) The Fungus Gnats (Diptera: Bolitophilidae, Diadocidiidae, Ditomyiidae, Keroplatidae and Mycetophilidae) of Greece, its islands and Cyprus. Studia dipterologica, 12 (2), 255 - 314." type="journal article" year="2005">
|
||
Chandler
|
||
<emphasis id="B9716A1473764C09FAEFFE78FE3FFE25" box="[1287,1343,439,465]" italics="true" pageId="29" pageNumber="101">et al.</emphasis>
|
||
(2005)
|
||
</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BBAB60673764C09FF2FFE14FF2FFDC9" blockId="29.[151,1437,151,609]" pageId="29" pageNumber="101">
|
||
This species is most similar to the Japanese
|
||
<taxonomicName id="4C05CD8573764C09FD42FE14F88FFE01" authority="Uesugi" authorityName="Uesugi" authorityYear="2005" box="[682,911,475,501]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis id="B9716A1473764C09FD42FE14F83AFE01" box="[682,826,475,501]" italics="true" pageId="29" pageNumber="101">M. circularis</emphasis>
|
||
Uesugi
|
||
</taxonomicName>
|
||
based on the very similarly-shaped gonostylus, well developed dorsal process of the gonocoxite, and reduced tergite 9, but is easily differentiated by the short inner process of the gonocoxite (inner process very long in
|
||
<taxonomicName id="4C05CD8573764C09FD18FDECF882FDC9" authorityName="Uesugi" authorityYear="2005" box="[752,898,547,573]" class="Insecta" family="Keroplatidae" genus="Macrorrhyncha" kingdom="Animalia" order="Diptera" pageId="29" pageNumber="101" phylum="Arthropoda" rank="species" species="circularis">
|
||
<emphasis id="B9716A1473764C09FD18FDECF882FDC9" box="[752,898,547,573]" italics="true" pageId="29" pageNumber="101">M. circularis</emphasis>
|
||
</taxonomicName>
|
||
;
|
||
<bibRefCitation id="EF94CBF773764C09FC78FDECFF24FDC9" author="Uesugi, K." box="[912,1060,546,573]" pageId="29" pageNumber="101" pagination="31 - 38" refId="ref17400" refString="Uesugi, K. (2005) Three new species of the genus Macrorrhyncha Winnertz in Japan with redescription of M. ezoensis (Okada) (Diptera: Keroplatidae). Tijdschrift voor Entomologie, 148, 31 - 38." type="journal article" year="2005">Uesugi 2005</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31FE58D73764C09FF2FFD89F8CEFD95" box="[199,974,582,609]" pageId="29" pageNumber="101" type="distribution">
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||
<paragraph id="8BBAB60673764C09FF2FFD89F8CEFD95" blockId="29.[151,1437,151,609]" box="[199,974,582,609]" pageId="29" pageNumber="101">
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||
<emphasis id="B9716A1473764C09FF2FFD89FA59FD94" bold="true" box="[199,345,582,608]" pageId="29" pageNumber="101">Distribution</emphasis>
|
||
. Known only from the
|
||
<typeStatus id="54BE08A473764C09FD88FD88F990FD95" box="[608,656,583,609]" pageId="29" pageNumber="101">type</typeStatus>
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||
locality in
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||
<collectingRegion id="49C178E473764C09FCE7FD88F867FD95" box="[783,871,583,609]" country="Canada" name="Quebec" pageId="29" pageNumber="101">Quebec</collectingRegion>
|
||
,
|
||
<collectingCountry id="F312F69673764C09FC9BFD88F8CBFD95" box="[883,971,583,609]" name="Canada" pageId="29" pageNumber="101">Canada</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |