treatments-xml/data/03/9F/5A/039F5A5FFFFCFFFFFF20FA8FFB7D3D29.xml

<document id="7517B142B845F084AF9B0A89A47422C8" ID-CLB-Dataset="5285" ID-DOI="10.11646/zootaxa.3835.4.3" ID-GBIF-Dataset="dbf8de20-9c7a-4b66-9130-615b6b42e84e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921623" ID-ZooBank="AD845CF3-CB19-4924-891F-330AAE283D07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623283883149" checkinUser="felipe" docAuthor="Yoshizawa, Kazunori, Lienhard, Charles &amp; Abd, Idris" docDate="2014" docId="039F5A5FFFFCFFFFFF20FA8FFB7D3D29" docLanguage="en" docName="zt03835p500.pdf" docOrigin="Zootaxa 3835 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichadenotecnum cinnamonum Endang &amp; New 2005" docType="treatment" docVersion="7" lastPageNumber="474" masterDocId="FFA62227FFFFFFFAFFB7FF9FFF893E1E" masterDocTitle="Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)" masterLastPageNumber="500" masterPageNumber="469" pageNumber="472" updateTime="1699009618324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="567B3C370C8AC5D87C2B0819E0D6DE74">Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)</mods:title>
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<mods:namePart id="99413A9B4789ED22FEFCAB04723E176C">Yoshizawa, Kazunori</mods:namePart>
<mods:affiliation id="A10BAD6C29EECD2165CE8C8023F58E98">Systematic Entomology, School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan.</mods:affiliation>
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<mods:namePart id="26A2666E90B2713A7D801EDF0AEA6C6D">Lienhard, Charles</mods:namePart>
<mods:affiliation id="2B4A147E534298AEEC9497B7B1CB6526">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="3912202A7A3AD4F6753C5AD6795DBB03">Abd, Idris</mods:namePart>
<mods:affiliation id="4544CEB4391B25EF7A52955EB8E10184">Center for Insect Systematics, Faculty of Sciences and Technology, Universiti Kebangsaan Malaysia, 43600 Bangi, Malaysia</mods:affiliation>
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<paragraph id="8B89EB49FFFCFFF9FF20FA8FFD653B34" blockId="3.[151,748,1296,1355]" box="[151,748,1296,1322]" pageId="3" pageNumber="472">
<heading id="D0C15C25FFFCFFF9FF20FA8FFD653B34" bold="true" box="[151,748,1296,1322]" fontSize="11" level="1" pageId="3" pageNumber="472" reason="1">
<taxonomicName id="4C3690CAFFFCFFF9FF20FA8FFD653B34" ID-CoL="587CC" authority="Endang &amp; New" authorityName="Endang &amp; New" authorityYear="2005" box="[151,748,1296,1322]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="472" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFCFFF9FF20FA8FFD653B34" bold="true" box="[151,748,1296,1322]" pageId="3" pageNumber="472">
<emphasis id="B942375BFFFCFFF9FF20FA8FFDAC3B34" bold="true" box="[151,549,1296,1322]" italics="true" pageId="3" pageNumber="472">Trichadenotecnum cinnamonum</emphasis>
Endang &amp; New
</emphasis>
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<subSubSection id="C32CB8C2FFFCFFF9FF20FAADFEC53B55" box="[151,332,1330,1355]" pageId="3" pageNumber="472" type="description">
<paragraph id="8B89EB49FFFCFFF9FF20FAADFEC53B55" blockId="3.[151,748,1296,1355]" box="[151,332,1330,1355]" pageId="3" pageNumber="472">
(
<figureCitation id="130DF7CCFFFCFFF9FF28FAADFF683B55" box="[159,225,1330,1355]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="3" pageNumber="472">Figs 1</figureCitation>
BC, 2, 3)
</paragraph>
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<subSubSection id="C32CB8C2FFFCFFF9FF20FAE5FC1F3BB1" pageId="3" pageNumber="472" type="reference_group">
<paragraph id="8B89EB49FFFCFFF9FF20FAE5FCBD3B8E" blockId="3.[151,918,1402,1455]" box="[151,820,1402,1424]" pageId="3" pageNumber="472">
<treatmentCitationGroup id="AB26CC67FFFCFFF9FF20FAE5FCBD3B8E" box="[151,820,1402,1424]" pageId="3" pageNumber="472">
<treatmentCitation id="0A97CD58FFFCFFF9FF20FAE5FD6D3B8E" author="Endang, S. K. &amp; New, T. R." box="[151,740,1402,1424]" page="35" pageId="3" pageNumber="472" year="2005">
<taxonomicName id="4C3690CAFFFCFFF9FF20FAE5FD6D3B8E" ID-CoL="587CC" authority="Endang &amp; New, 2005: 35" authorityName="Endang &amp; New" authorityPageNumber="35" authorityYear="2005" box="[151,740,1402,1424]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="472" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFCFFF9FF20FAE5FE543B8E" box="[151,477,1402,1424]" italics="true" pageId="3" pageNumber="472">Trichadenotecnum cinnamonum</emphasis>
<bibRefCitation id="EFA796B8FFFCFFF9FE54FAE5FD6D3B8E" author="Endang, S. K. &amp; New, T. R." box="[483,740,1402,1424]" pageId="3" pageNumber="472" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New, 2005: 35</bibRefCitation>
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(male).
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[Not 
<taxonomicName id="4C3690CAFFFCFFF9FF7BFA05FDF93BB1" ID-CoL="587CC" authority="Endang &amp; New, 2005: 36" authorityName="Endang &amp; New" authorityPageNumber="36" authorityYear="2005" box="[204,624,1433,1455]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="472" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFCFFF9FF7BFA05FEEE3BB1" box="[204,359,1434,1455]" italics="true" pageId="3" pageNumber="472">T. cinnamonum</emphasis>
<bibRefCitation id="EFA796B8FFFCFFF9FEDAFA06FDF93BB1" author="Endang, S. K. &amp; New, T. R." box="[365,624,1433,1455]" pageId="3" pageNumber="472" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New, 2005: 36</bibRefCitation>
</taxonomicName>
(female): Misidentification.]
</paragraph>
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<subSubSection id="C32CB8C2FFFCFFF9FF20FA43FC8838D3" pageId="3" pageNumber="472" type="description">
<paragraph id="8B89EB49FFFCFFF9FF20FA43FAC53807" blockId="3.[151,1438,1500,1885]" pageId="3" pageNumber="472">
<emphasis id="B942375BFFFCFFF9FF20FA43FE193BEB" bold="true" box="[151,400,1500,1525]" pageId="3" pageNumber="472">Specimens examined</emphasis>
: [
<collectingCountry id="F321ABD9FFFCFFF9FE11FA43FD9B3BEB" box="[422,530,1500,1525]" name="Malaysia" pageId="3" pageNumber="472">Malaysia</collectingCountry>
] 
<specimenCount id="9D3020C0FFFCFFF9FD96FA43FDE53BEB" box="[545,620,1500,1525]" pageId="3" pageNumber="472" type="male">1 male</specimenCount>
(KY347), Gunung Brinchang, Cameron Highland, 
<collectingRegion id="49F225ABFFFCFFF9FB03FA43FA803BEB" box="[1204,1289,1500,1525]" country="Malaysia" name="Pahang" pageId="3" pageNumber="472">Pahang</collectingRegion>
, 
<date id="FF88CD89FFFCFFF9FAA1FA43FA113BEB" box="[1302,1432,1500,1525]" pageId="3" pageNumber="472" value="2003-07-13">13.vii.2003</date>
, H. Kojima (UKM); 
<materialsCitation id="3B5EE114FFFCFFF9FEC0F99FFAC13807" ID-GBIF-Occurrence="3326724314" box="[375,1352,1536,1561]" collectingDate="2003-07-14" collectionCode="UKM" county="Gunung Jasar" location="Cameron Highland" municipality="Path" pageId="3" pageNumber="472" specimenCount="1" specimenCount-female="1" stateProvince="Pahang">
<specimenCount id="9D3020C0FFFCFFF9FEC0F99FFE523807" box="[375,475,1536,1561]" pageId="3" pageNumber="472" type="female">1 female</specimenCount>
, 
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, 
<collectingMunicipality id="6BED7133FFFCFFF9FD3BF99FFD373807" box="[652,702,1536,1561]" pageId="3" pageNumber="472">Path</collectingMunicipality>
10, 
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, 
<collectingRegion id="49F225ABFFFCFFF9FC64F99FFBA13807" box="[979,1064,1536,1561]" country="Malaysia" name="Pahang" pageId="3" pageNumber="472">Pahang</collectingRegion>
, 
<date id="FF88CD89FFFCFFF9FB84F99FFB3C3807" box="[1075,1205,1536,1561]" pageId="3" pageNumber="472" value="2003-07-14">
<collectingDate id="EFCC3461FFFCFFF9FB84F99FFB3C3807" box="[1075,1205,1536,1561]" pageId="3" pageNumber="472" value="2003-07-14">14.vii.2003</collectingDate>
</date>
, KY (
<collectionCode id="ED27738CFFFCFFF9FB4FF99FFAB63807" box="[1272,1343,1536,1561]" country="Malaysia" httpUri="http://grbio.org/cool/ashc-3pfs" name="Universiti Kebangsaan Malaysia" pageId="3" pageNumber="472">UKM</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B89EB49FFFCFFF9FF70F9BBFC8838D3" blockId="3.[151,1438,1500,1885]" pageId="3" pageNumber="472">
<emphasis id="B942375BFFFCFFF9FF70F9BBFDB03825" box="[199,569,1572,1597]" inLineHeading="true" pageId="3" pageNumber="472" reason="1">
<emphasis id="B942375BFFFCFFF9FF70F9BBFDBB3823" bold="true" box="[199,562,1572,1597]" pageId="3" pageNumber="472">Description of female genitalia</emphasis>
.
</emphasis>
Egg guide long (
<figureCitation id="130DF7CCFFFCFFF9FD4FF9BBFCD83823" box="[760,849,1572,1597]" captionStart="FIGURE 3" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[155,1427,1171,1846]" captionTargetId="figure-74@4.[151,1435,1163,1850]" captionTargetPageId="4" captionText="FIGURE 3. Female genitalia of Trichadenotecnum cinnamonum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921632" httpUri="https://zenodo.org/record/4921632/files/figure.png" pageId="3" pageNumber="472">Fig. 3A</figureCitation>
), much longer than basal width, parallel sided, distal margin almost straight; body of subgenital plate short and wide, posteriorly with shallow membranous region next to egg guide, with V-shaped membranous region anteromedially. Gonapophyses (
<figureCitation id="130DF7CCFFFCFFF9FBFDF9F3FB2F389B" box="[1098,1190,1644,1669]" captionStart="FIGURE 3" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[155,1427,1171,1846]" captionTargetId="figure-74@4.[151,1435,1163,1850]" captionTargetPageId="4" captionText="FIGURE 3. Female genitalia of Trichadenotecnum cinnamonum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921632" httpUri="https://zenodo.org/record/4921632/files/figure.png" pageId="3" pageNumber="472">Fig. 3B</figureCitation>
). Ventral valve long; dorsal valve with short distal process; posterior lobe of external valve narrow. Spermapore plate (
<figureCitation id="130DF7CCFFFCFFF9FB53F90FFAB338B7" box="[1252,1338,1680,1705]" captionStart="FIGURE 3" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[155,1427,1171,1846]" captionTargetId="figure-74@4.[151,1435,1163,1850]" captionTargetPageId="4" captionText="FIGURE 3. Female genitalia of Trichadenotecnum cinnamonum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921632" httpUri="https://zenodo.org/record/4921632/files/figure.png" pageId="3" pageNumber="472">Fig. 3C</figureCitation>
) weakly sclerotized, anterior membranous region asymmetrical.
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFFCFFFFFF70F947FB7D3D29" lastPageId="5" lastPageNumber="474" pageId="3" pageNumber="472" type="discussion">
<paragraph id="8B89EB49FFFCFFF9FF70F947FCD33943" blockId="3.[151,1438,1500,1885]" pageId="3" pageNumber="472">
<emphasis id="B942375BFFFCFFF9FF70F947FEB238EE" box="[199,315,1752,1777]" inLineHeading="true" pageId="3" pageNumber="472" reason="1">
<emphasis id="B942375BFFFCFFF9FF70F947FEBD38EF" bold="true" box="[199,308,1752,1777]" pageId="3" pageNumber="472">Remarks</emphasis>
.
</emphasis>
The male specimen examined here is in complete agreement with the original description of 
<taxonomicName id="4C3690CAFFFCFFF9FA30F946FEA1390A" authorityName="Endang &amp; New" authorityYear="2005" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="3" pageNumber="472" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFCFFF9FA30F946FEA1390A" italics="true" pageId="3" pageNumber="472">T. cinnamonum</emphasis>
</taxonomicName>
, except that the hypandrial structures are in mirror images between them (
<figureCitation id="130DF7CCFFFCFFF9FB34F963FB54390B" box="[1155,1245,1788,1813]" captionStart="FIGURE 2" captionStartId="4.[151,250,1061,1083]" captionTargetBox="[155,1432,194,1030]" captionTargetId="figure-65@4.[150,1436,193,1039]" captionTargetPageId="4" captionText="FIGURE 2. Male terminalia of Trichadenotecnum cinnamonum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921630" httpUri="https://zenodo.org/record/4921630/files/figure.png" pageId="3" pageNumber="472">Fig. 2C</figureCitation>
). Because of the exact similarity, we concluded that the hypandrium illustrated in the original description is drawn from the reverse. This is the first record of this species from Peninsula 
<collectingCountry id="F321ABD9FFFCFFF9FD5BF8DBFCDF3943" box="[748,854,1860,1885]" name="Malaysia" pageId="3" pageNumber="472">Malaysia</collectingCountry>
.
</paragraph>
<caption id="DF49BBC1FFFBFFFEFF20FBBAFD443A44" ID-DOI="http://doi.org/10.5281/zenodo.4921630" ID-Zenodo-Dep="4921630" httpUri="https://zenodo.org/record/4921630/files/figure.png" pageId="4" pageNumber="473" startId="4.[151,250,1061,1083]" targetBox="[155,1432,194,1030]" targetPageId="4">
<paragraph id="8B89EB49FFFBFFFEFF20FBBAFD443A44" blockId="4.[151,1436,1061,1114]" pageId="4" pageNumber="473">
<emphasis id="B942375BFFFBFFFEFF20FBBAFE9E3A24" bold="true" box="[151,279,1061,1083]" pageId="4" pageNumber="473">FIGURE 2.</emphasis>
Male terminalia of 
<taxonomicName id="4C3690CAFFFBFFFEFE47FBBAFCB33A25" authorityName="Endang &amp; New" authorityYear="2005" box="[496,826,1061,1083]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="473" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFBFFFEFE47FBBAFCB33A25" box="[496,826,1061,1083]" italics="true" pageId="4" pageNumber="473">Trichadenotecnum cinnamonum</emphasis>
</taxonomicName>
. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view.
</paragraph>
</caption>
<caption id="DF49BBC1FFFBFFFEFF20F8CFFEC5399A" ID-DOI="http://doi.org/10.5281/zenodo.4921632" ID-Zenodo-Dep="4921632" httpUri="https://zenodo.org/record/4921632/files/figure.png" pageId="4" pageNumber="473" startId="4.[151,250,1872,1894]" targetBox="[155,1427,1171,1846]" targetPageId="4">
<paragraph id="8B89EB49FFFBFFFEFF20F8CFFEC5399A" blockId="4.[151,1436,1872,1924]" pageId="4" pageNumber="473">
<emphasis id="B942375BFFFBFFFEFF20F8CFFE9E397B" bold="true" box="[151,279,1872,1894]" pageId="4" pageNumber="473">FIGURE 3.</emphasis>
Female genitalia of 
<taxonomicName id="4C3690CAFFFBFFFEFE4EF8CFFCCA3978" authorityName="Endang &amp; New" authorityYear="2005" box="[505,835,1872,1894]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="4" pageNumber="473" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFBFFFEFE4EF8CFFCCA3978" box="[505,835,1872,1894]" italics="true" pageId="4" pageNumber="473">Trichadenotecnum cinnamonum</emphasis>
</taxonomicName>
, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate.
</paragraph>
</caption>
<paragraph id="8B89EB49FFFAFFFFFF70FF08FD613FB5" blockId="5.[151,1438,151,824]" pageId="5" pageNumber="474">
Originally, the 
<emphasis id="B942375BFFFAFFFFFED9FF08FE643EAE" box="[366,493,151,176]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group was characterized by the presence of a process arising from the trichobothrial field of the male paraproct (Yoshizawa, 2003). Later, absence of this structure in 
<taxonomicName id="4C3690CAFFFAFFFFFBEFFF22FAEC3ECB" authority="Thornton, 1961" authorityName="Thornton" authorityYear="1961" box="[1112,1381,188,213]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="B942375BFFFAFFFFFBEFFF22FB2E3ECB" box="[1112,1191,188,213]" italics="true" pageId="5" pageNumber="474">T. felix</emphasis>
<bibRefCitation id="EFA796B8FFFAFFFFFB04FF23FAEC3ECB" author="Thornton, I. W. B." box="[1203,1381,188,213]" pageId="5" pageNumber="474" pagination="1 - 24" refId="ref15445" refString="Thornton, I. W. B. (1961) The Trichadenotecnum group (Psocoptera: Psocidae) in Hong Kong, with descriptions of new species. Transactions of the Royal Entomological Society, London, 113, 1 - 24. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1961. tb 00797. x" type="journal article" year="1961">Thornton, 1961</bibRefCitation>
</taxonomicName>
was reported, although the hypandrial structure strongly suggests its placement to the 
<emphasis id="B942375BFFFAFFFFFB9BFF40FB223EE6" box="[1068,1195,223,248]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group (
<bibRefCitation id="EFA796B8FFFAFFFFFAB4FF40FEC13F03" author="Yoshizawa, K. &amp; Lienhard, C." pageId="5" pageNumber="474" pagination="121 - 149" refId="ref15885" refString="Yoshizawa, K. &amp; Lienhard, C. (2004) Systematics of Trichadenotecnum (Psocoptera: Psocidae) in Hong Kong. Publicaciones Especiales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, 20, 121 - 149." type="journal article" year="2004">Yoshizawa &amp; Lienhard, 2004</bibRefCitation>
). The hypandrial structure of 
<taxonomicName id="4C3690CAFFFAFFFFFD22FE9AFCCB3F02" authorityName="Endang &amp; New" authorityYear="2005" box="[661,834,261,284]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFD22FE9AFCCB3F02" box="[661,834,261,284]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
is more similar to that of 
<taxonomicName id="4C3690CAFFFAFFFFFBDFFE9AFB3D3F03" authorityName="Thornton" authorityYear="1961" box="[1128,1204,260,285]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="B942375BFFFAFFFFFBDFFE9AFB3D3F03" box="[1128,1204,260,285]" italics="true" pageId="5" pageNumber="474">T. felix</emphasis>
</taxonomicName>
than to species with the trichobothrial process, but 
<taxonomicName id="4C3690CAFFFAFFFFFDBBFEB6FD493F21" authorityName="Endang &amp; New" authorityYear="2005" box="[524,704,296,320]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFDBBFEB6FD493F21" box="[524,704,296,320]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
has a keel on the trichobothrial field which is most likely homologous with the trichobothrial process. This suggests that the trichobothrial process is secondarily absent in 
<taxonomicName id="4C3690CAFFFAFFFFFA30FED2FF4E3F96" authorityName="Thornton" authorityYear="1961" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="B942375BFFFAFFFFFA30FED2FF4E3F96" italics="true" pageId="5" pageNumber="474">T. felix</emphasis>
</taxonomicName>
, and its presence is autapomorphic for the 
<emphasis id="B942375BFFFAFFFFFD64FEF0FCDB3F96" box="[723,850,367,392]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group. The assignment of 
<taxonomicName id="4C3690CAFFFAFFFFFB2DFEEEFAC53F99" authorityName="Endang &amp; New" authorityYear="2005" box="[1178,1356,368,392]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFB2DFEEEFAC53F99" box="[1178,1356,368,392]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
to the 
<emphasis id="B942375BFFFAFFFFFF20FE0BFE9E3FB3" box="[151,279,404,429]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group can be warranted by this structure.
</paragraph>
<paragraph id="8B89EB49FFFAFFFFFF70FE28FB7D3D29" blockId="5.[151,1438,151,824]" pageId="5" pageNumber="474">
The female specimen examined here differs significantly from the description of the 
<typeStatus id="548D55EBFFFAFFFFFB37FE27FB693FCE" box="[1152,1248,440,464]" pageId="5" pageNumber="474" type="paratype">paratype</typeStatus>
female given by 
<bibRefCitation id="EFA796B8FFFAFFFFFF20FE43FE253FEB" author="Endang, S. K. &amp; New, T. R." box="[151,428,476,501]" pageId="5" pageNumber="474" pagination="15 - 38" refId="ref14985" refString="Endang, S. K. &amp; New, T. R. (2005) The genus Trichadenotecnum (Insecta: Psocoptera: Psocidae) in Sumatra, Indonesia, with description of thirteen new species. Records of the Australian Museum, 57, 15 - 38. http: // dx. doi. org / 10.3853 / j. 0067 - 1975.57.2005.1437" type="journal article" year="2005">Endang &amp; New (2005)</bibRefCitation>
; this clearly shows that they belong to different species. The 
<typeStatus id="548D55EBFFFAFFFFFB2BFE43FB743FEA" box="[1180,1277,476,500]" pageId="5" pageNumber="474" type="paratype">paratype</typeStatus>
female of 
<taxonomicName id="4C3690CAFFFAFFFFFA30FE42FEA13C09" authorityName="Endang &amp; New" authorityYear="2005" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFA30FE42FEA13C09" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
also looks significantly different from the 
<typeStatus id="548D55EBFFFAFFFFFCADFE60FCF53C06" box="[794,892,511,536]" pageId="5" pageNumber="474" type="holotype">holotype</typeStatus>
male in forewing markings (heavily spotted in female 
<typeStatus id="548D55EBFFFAFFFFFF5EFDBBFEC33C22" box="[233,330,548,572]" pageId="5" pageNumber="474" type="paratype">paratype</typeStatus>
whereas much more sparsely spotted in 
<typeStatus id="548D55EBFFFAFFFFFCBAFDBBFCE63C23" box="[781,879,548,573]" pageId="5" pageNumber="474" type="holotype">holotype</typeStatus>
male). In contrast, the female identified here as 
<taxonomicName id="4C3690CAFFFAFFFFFA30FDBAFEA13C41" authorityName="Endang &amp; New" authorityYear="2005" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFA30FDBAFEA13C41" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
is more similar to the 
<typeStatus id="548D55EBFFFAFFFFFD93FDD8FD0F3C7E" box="[548,646,583,608]" pageId="5" pageNumber="474" type="holotype">holotype</typeStatus>
male of this species in the forewing markings (
<figureCitation id="130DF7CCFFFAFFFFFB2CFDD8FB533C7E" box="[1179,1242,583,608]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="5" pageNumber="474">Fig. 1</figureCitation>
BC). In addition, the female genital structures of the specimen examined here (
<figureCitation id="130DF7CCFFFAFFFFFCD7FDF3FC2C3C9B" box="[864,933,620,645]" captionStart="FIGURE 3" captionStartId="4.[151,250,1872,1894]" captionTargetBox="[155,1427,1171,1846]" captionTargetId="figure-74@4.[151,1435,1163,1850]" captionTargetPageId="4" captionText="FIGURE 3. Female genitalia of Trichadenotecnum cinnamonum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921632" httpUri="https://zenodo.org/record/4921632/files/figure.png" pageId="5" pageNumber="474">Fig. 3</figureCitation>
) are very similar to those of 
<taxonomicName id="4C3690CAFFFAFFFFFB4AFDF2FAC23C9B" authorityName="Thornton" authorityYear="1961" box="[1277,1355,620,645]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="B942375BFFFAFFFFFB4AFDF2FAC23C9B" box="[1277,1355,620,645]" italics="true" pageId="5" pageNumber="474">T. felix</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C3690CAFFFAFFFFFA30FDF2FF693CB9" authorityName="New &amp; Thornton" authorityYear="1976" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="imrum">
<emphasis id="B942375BFFFAFFFFFA30FDF2FF693CB9" italics="true" pageId="5" pageNumber="474">T. imrum</emphasis>
</taxonomicName>
, both of which are members of the 
<emphasis id="B942375BFFFAFFFFFD3DFD10FC803CB6" box="[650,777,655,680]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group. In contrast, the genital structures of the 
<typeStatus id="548D55EBFFFAFFFFFA8BFD0FFA153CB6" box="[1340,1436,656,680]" pageId="5" pageNumber="474" type="paratype">paratype</typeStatus>
female, especially the absence of the posterior lobe of the external valve of gonapophyses, are more similar to the species of the 
<emphasis id="B942375BFFFAFFFFFE8DFD48FE5F3CEE" box="[314,470,727,752]" italics="true" pageId="5" pageNumber="474">spiniserrulum</emphasis>
group (e.g., 
<taxonomicName id="4C3690CAFFFAFFFFFDDFFD48FCD63CEE" box="[616,863,727,752]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B942375BFFFAFFFFFDDFFD48FCB13CEE" box="[616,824,727,752]" italics="true" pageId="5" pageNumber="474">Trichadenotecnum</emphasis>
sp.
</taxonomicName>
My19 or 
<taxonomicName id="4C3690CAFFFAFFFFFC7BFD46FBD73CEE" authorityName="New" authorityYear="1975" box="[972,1118,727,752]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="malayense">
<emphasis id="B942375BFFFAFFFFFC7BFD46FBD73CEE" box="[972,1118,727,752]" italics="true" pageId="5" pageNumber="474">T. malayense</emphasis>
</taxonomicName>
, treated below: 
<figureCitation id="130DF7CCFFFAFFFFFAA0FD48FAE23CEE" box="[1303,1387,727,752]" captionStart="FIGURE 14" captionStartId="15.[151,250,1545,1567]" captionTargetBox="[157,1430,937,1519]" captionTargetId="figure-171@15.[151,1435,934,1523]" captionTargetPageId="15" captionText="FIGURE 14. Female genitalia of Trichadenotecnum sp.My19, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921654" httpUri="https://zenodo.org/record/4921654/files/figure.png" pageId="5" pageNumber="474">Figs 14</figureCitation>
and 
<figureCitation id="130DF7CCFFFAFFFFFF20FD63FF3C3D0B" box="[151,181,764,789]" captionStart="FIGURE 17" captionStartId="17.[151,250,1799,1821]" captionTargetBox="[155,1431,1087,1774]" captionTargetId="figure-154@17.[151,1435,1078,1777]" captionTargetPageId="17" captionText="FIGURE 17. Female genitalia of Trichadenotecnum malayense, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921660" httpUri="https://zenodo.org/record/4921660/files/figure.png" pageId="5" pageNumber="474">17</figureCitation>
, respectively). Therefore, we conclude that the female 
<typeStatus id="548D55EBFFFAFFFFFC80FD63FC1E3D0A" box="[823,919,764,788]" pageId="5" pageNumber="474" type="paratype">paratype</typeStatus>
of 
<taxonomicName id="4C3690CAFFFAFFFFFC74FD62FBFA3D0A" authorityName="Endang &amp; New" authorityYear="2005" box="[963,1139,765,788]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFC74FD62FBFA3D0A" box="[963,1139,765,788]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
was misidentified as that species, and we here provide the first description of the female genital structures of 
<taxonomicName id="4C3690CAFFFAFFFFFBF6FCBEFB643D29" authorityName="Endang &amp; New" authorityYear="2005" box="[1089,1261,800,824]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFBF6FCBEFB643D29" box="[1089,1261,800,824]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>