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<document id="7B9A8E4C8D08C75E609798B058F53970" ID-CLB-Dataset="5285" ID-DOI="10.11646/zootaxa.3835.4.3" ID-GBIF-Dataset="dbf8de20-9c7a-4b66-9130-615b6b42e84e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4921623" ID-ZooBank="AD845CF3-CB19-4924-891F-330AAE283D07" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623283883149" checkinUser="felipe" docAuthor="Yoshizawa, Kazunori, Lienhard, Charles &amp; Abd, Idris" docDate="2014" docId="039F5A5FFFFAFFFFFF20FC13FE6939C8" docLanguage="en" docName="zt03835p500.pdf" docOrigin="Zootaxa 3835 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Trichadenotecnum imrum New &amp; Thornton 1976" docType="treatment" docVersion="7" lastPageNumber="474" masterDocId="FFA62227FFFFFFFAFFB7FF9FFF893E1E" masterDocTitle="Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)" masterLastPageNumber="500" masterPageNumber="469" pageNumber="474" updateTime="1699009618324" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5B0AAEA392CE58EE7C483DD3694DA724">Trichadenotecnum species from Peninsular Malaysia and Singapore (Insecta: Psocodea: ' Psocoptera': Psocidae)</mods:title>
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<mods:namePart id="5B13460D3E26D529F1F40B7F1939B237">Yoshizawa, Kazunori</mods:namePart>
<mods:affiliation id="CABFA69B9FA848E9F4184501BF8D2D84">Systematic Entomology, School of Agriculture, Hokkaido University, Sapporo 060 - 8589, Japan.</mods:affiliation>
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<mods:namePart id="375F4CB3DC5051C957E6CE8940559CBE">Lienhard, Charles</mods:namePart>
<mods:affiliation id="6253FC22B3A40742608020FBBD17877E">Muséum d'histoire naturelle, C. P. 6434, CH- 1211 Genève 6, Switzerland</mods:affiliation>
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<mods:namePart id="DE8331B53DC39040C7AB8CCBF9030FC5">Abd, Idris</mods:namePart>
<mods:affiliation id="8E548F416DCCBC58F76F933F4DEB68A5">Center for Insect Systematics, Faculty of Sciences and Technology, Universiti Kebangsaan Malaysia, 43600 Bangi, Malaysia</mods:affiliation>
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<mods:title id="C75CE8B46C2929FD5AA4856C7B1E136B">Zootaxa</mods:title>
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<treatment id="039F5A5FFFFAFFFFFF20FC13FE6939C8" ID-DOI="http://doi.org/10.5281/zenodo.4921949" ID-GBIF-Taxon="183731214" ID-Zenodo-Dep="4921949" LSID="urn:lsid:plazi:treatment:039F5A5FFFFAFFFFFF20FC13FE6939C8" httpUri="http://treatment.plazi.org/id/039F5A5FFFFAFFFFFF20FC13FE6939C8" lastPageNumber="474" pageId="5" pageNumber="474">
<subSubSection id="C32CB8C2FFFAFFFFFF20FC13FD3A3DB8" box="[151,691,908,934]" pageId="5" pageNumber="474" type="nomenclature">
<paragraph id="8B89EB49FFFAFFFFFF20FC13FD3A3DB8" blockId="5.[151,691,908,967]" box="[151,691,908,934]" pageId="5" pageNumber="474">
<heading id="D0C15C25FFFAFFFFFF20FC13FD3A3DB8" bold="true" box="[151,691,908,934]" fontSize="11" level="1" pageId="5" pageNumber="474" reason="1">
<taxonomicName id="4C3690CAFFFAFFFFFF20FC13FD3A3DB8" ID-CoL="7CQPV" authority="New &amp; Thornton" authorityName="New &amp; Thornton" authorityYear="1976" box="[151,691,908,934]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="imrum">
<emphasis id="B942375BFFFAFFFFFF20FC13FD3A3DB8" bold="true" box="[151,691,908,934]" pageId="5" pageNumber="474">
<emphasis id="B942375BFFFAFFFFFF20FC13FE5C3DB8" bold="true" box="[151,469,908,934]" italics="true" pageId="5" pageNumber="474">Trichadenotecnum imrum</emphasis>
New &amp; Thornton
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFFAFFFFFF20FC31FEB23DD9" box="[151,315,942,967]" pageId="5" pageNumber="474" type="description">
<paragraph id="8B89EB49FFFAFFFFFF20FC31FEB23DD9" blockId="5.[151,691,908,967]" box="[151,315,942,967]" pageId="5" pageNumber="474">
(
<figureCitation id="130DF7CCFFFAFFFFFF28FC31FF723DD9" box="[159,251,942,967]" captionStart="FIGURE 1.1" captionStartId="2.[151,250,1786,1808]" captionTargetBox="[151,1435,196,1746]" captionTargetId="figure-94@2.[151,1436,193,1765]" captionTargetPageId="2" captionText="FIGURE 1.1. Forewings of Trichadenotecnum spp.: A: T. circularoides, female; B: T. cinnamonum, male; C: ditto, female; D: T. imrum, male; E: T.pardus, female; F: T. yatai, holotype male; G: T. kojimai, holotype male; H: T. adika, male; I: T. endangae, holotype male; J: T. hammani, female; K: T. sp.My19, female; L: T. gombakense, female; M: T. malayense, male." figureDoi="http://doi.org/10.5281/zenodo.4921625" httpUri="https://zenodo.org/record/4921625/files/figure.png" pageId="5" pageNumber="474">Figs 1D</figureCitation>
,
<figureCitation id="130DF7CCFFFAFFFFFEBFFC31FE913DD9" box="[264,280,942,967]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">4</figureCitation>
,
<figureCitation id="130DF7CCFFFAFFFFFE93FC31FEBB3DD9" box="[292,306,942,967]" captionStart="FIGURE 5" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[153,1426,1213,1935]" captionTargetId="figure-74@6.[151,1436,1203,1948]" captionTargetPageId="6" captionText="FIGURE 5. Female genitalia of Trichadenotecnum imrum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921636" httpUri="https://zenodo.org/record/4921636/files/figure.png" pageId="5" pageNumber="474">5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFFAFFFFFF20FC69FD163AA0" pageId="5" pageNumber="474" type="reference_group">
<paragraph id="8B89EB49FFFAFFFFFF20FC69FD433A12" blockId="5.[151,714,1014,1036]" box="[151,714,1014,1036]" pageId="5" pageNumber="474">
<treatmentCitationGroup id="AB26CC67FFFAFFFFFF20FC69FD433A12" box="[151,714,1014,1036]" pageId="5" pageNumber="474">
<treatmentCitation id="0A97CD58FFFAFFFFFF20FC69FD4F3A12" author="New, T. R. &amp; Thornton, I. W. B." box="[151,710,1014,1036]" page="414" pageId="5" pageNumber="474" year="1976">
<taxonomicName id="4C3690CAFFFAFFFFFF20FC69FD4F3A12" ID-CoL="7CQPV" authority="New &amp; Thornton, 1976: 414" authorityName="New &amp; Thornton" authorityPageNumber="414" authorityYear="1976" box="[151,710,1014,1036]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="imrum">
<emphasis id="B942375BFFFAFFFFFF20FC69FE153A12" box="[151,412,1014,1036]" italics="true" pageId="5" pageNumber="474">Trichadenotecnum imrum</emphasis>
<bibRefCitation id="EFA796B8FFFAFFFFFE15FC68FD4F3A12" author="New, T. R. &amp; Thornton, I. W. B." box="[418,710,1014,1036]" pageId="5" pageNumber="474" pagination="375 - 418" refId="ref15397" refString="New, T. R. &amp; Thornton, I. W. B. (1976) Psocomorpha (Psocoptera) from the Malayan Peninsula. Oriental Insects, 9, 375 - 418. http: // dx. doi. org / 10.1080 / 00305316.1975.10434508" type="journal article" year="1976">New &amp; Thornton, 1976: 414</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8B89EB49FFFAFFFFFF20FBA6FD163AA0" blockId="5.[151,1438,1081,2006]" pageId="5" pageNumber="474">
<emphasis id="B942375BFFFAFFFFFF20FBA6FE073A4C" bold="true" box="[151,398,1081,1106]" pageId="5" pageNumber="474">Specimens examined</emphasis>
: [
<collectingCountry id="F321ABD9FFFAFFFFFE14FBA6FD933A4C" box="[419,538,1081,1106]" name="Singapore" pageId="5" pageNumber="474">Singapore</collectingCountry>
]
<specimenCount id="9D3020C0FFFAFFFFFD90FBA6FDFD3A4C" box="[551,628,1081,1106]" pageId="5" pageNumber="474" type="male">1 male</specimenCount>
, Bukit Timah forest, 24 hr survey, section trap 79 am,
<date id="FF88CD89FFFAFFFFFB6AFBA6FAD93A4C" box="[1245,1360,1081,1106]" pageId="5" pageNumber="474" value="1969-10-26">26.x.1969</date>
, DHM (MHNG);
<materialsCitation id="3B5EE114FFFAFFFFFEB8FBC2FB2C3A68" ID-GBIF-Occurrence="3326724304" box="[271,1189,1117,1142]" collectingDate="1981-11-01" collectionCode="MHNG" location="Bukit Timah forest" pageId="5" pageNumber="474" specimenCount="2" specimenCount-female="1" specimenCount-male="1">
<specimenCount id="9D3020C0FFFAFFFFFEB8FBC2FED53A68" box="[271,348,1117,1142]" pageId="5" pageNumber="474" type="male">1 male</specimenCount>
<specimenCount id="9D3020C0FFFAFFFFFED2FBC2FE433A68" box="[357,458,1117,1142]" pageId="5" pageNumber="474" type="female">1 female</specimenCount>
,
<location id="8EE9BD92FFFAFFFFFE60FBC2FD383A68" LSID="urn:lsid:plazi:treatment:039F5A5FFFFAFFFFFF20FC13FE6939C8:8EE9BD92FFFAFFFFFE60FBC2FD383A68" box="[471,689,1117,1142]" name="Bukit Timah forest" pageId="5" pageNumber="474">Bukit Timah forest</location>
, beating foliage,
<date id="FF88CD89FFFAFFFFFCC2FBC2FC683A68" box="[885,993,1117,1142]" pageId="5" pageNumber="474" value="1981-11-01">
<collectingDate id="EFCC3461FFFAFFFFFCC2FBC2FC683A68" box="[885,993,1117,1142]" pageId="5" pageNumber="474" value="1981-11-01">1.xi.1981</collectingDate>
</date>
, DHM (
<collectionCode id="ED27738CFFFAFFFFFBF7FBC2FB153A68" box="[1088,1180,1117,1142]" country="Switzerland" httpUri="http://grbio.org/cool/2jyg-xmd2" name="Museum d'Histoire Naturelle" pageId="5" pageNumber="474" type="Museum">MHNG</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5EE114FFFAFFFFFB06FBC2FD123AA0" ID-GBIF-Occurrence="3326724322" collectingDate="1987-12-17" collectionCode="SEHU" county="Lower Path" elevation="125" location="Bukit Timah Nature Reserve" municipality="Rock Path" pageId="5" pageNumber="474" specimenCount="1" specimenCount-male="1">
<specimenCount id="9D3020C0FFFAFFFFFB06FBC2FA893A68" box="[1201,1280,1117,1142]" pageId="5" pageNumber="474" type="male">1 male</specimenCount>
,
<location id="8EE9BD92FFFAFFFFFABAFBC2FEC53A87" LSID="urn:lsid:plazi:treatment:039F5A5FFFFAFFFFFF20FC13FE6939C8:8EE9BD92FFFAFFFFFABAFBC2FEC53A87" county="Lower Path" municipality="Rock Path" name="Bukit Timah Nature Reserve" pageId="5" pageNumber="474">Bukit Timah Nature Reserve</location>
, between
<location id="8EE9BD92FFFAFFFFFE71FB1EFD973A84" LSID="urn:lsid:plazi:treatment:039F5A5FFFFAFFFFFF20FC13FE6939C8:8EE9BD92FFFAFFFFFE71FB1EFD973A84" box="[454,542,1153,1178]" county="Lower Path" municipality="Rock Path" name="Summit" pageId="5" pageNumber="474">Summit</location>
hut and
<location id="8EE9BD92FFFAFFFFFD27FB1EFD723A84" LSID="urn:lsid:plazi:treatment:039F5A5FFFFAFFFFFF20FC13FE6939C8:8EE9BD92FFFAFFFFFD27FB1EFD723A84" box="[656,763,1153,1178]" county="Lower Path" municipality="Rock Path" name="Car Park" pageId="5" pageNumber="474">Car Park</location>
, along
<location id="8EE9BD92FFFAFFFFFCE1FB1DFB8E3A84" LSID="urn:lsid:plazi:treatment:039F5A5FFFFAFFFFFF20FC13FE6939C8:8EE9BD92FFFAFFFFFCE1FB1DFB8E3A84" box="[854,1031,1154,1178]" county="Lower Path" municipality="Rock Path" name="Tangga Rengas" pageId="5" pageNumber="474">Tangga Rengas</location>
trail,
<collectingMunicipality id="6BED7133FFFAFFFFFBE3FB1EFB443A84" box="[1108,1229,1153,1178]" pageId="5" pageNumber="474">Rock Path</collectingMunicipality>
and
<collectingCounty id="62E893C5FFFAFFFFFAB9FB1DFA1E3A84" box="[1294,1431,1153,1178]" pageId="5" pageNumber="474">Lower Path</collectingCounty>
,
<quantity id="4CCE46ACFFFAFFFFFF20FB3AFE9E3AA3" box="[151,279,1189,1214]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="5" pageNumber="474" unit="m" value="125.0" valueMax="140.0" valueMin="110.0">
<elevation id="001B0C7AFFFAFFFFFF20FB3AFE9E3AA3" box="[151,279,1189,1214]" metricMagnitude="2" metricUnit="m" metricValue="1.25" metricValueMax="1.4" metricValueMin="1.1" pageId="5" pageNumber="474" unit="m" value="125.0" valueMax="140.0" valueMin="110.0">110140 m</elevation>
</quantity>
, beating,
<date id="FF88CD89FFFAFFFFFE34FB3AFD8C3AA0" box="[387,517,1189,1214]" pageId="5" pageNumber="474" value="1987-12-17">
<collectingDate id="EFCC3461FFFAFFFFFE34FB3AFD8C3AA0" box="[387,517,1189,1214]" pageId="5" pageNumber="474" value="1987-12-17">17.xii.1987</collectingDate>
</date>
, CL (
<collectionCode id="ED27738CFFFAFFFFFDF4FB3AFD1B3AA0" box="[579,658,1189,1214]" pageId="5" pageNumber="474">SEHU</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFFAFFFFFF70FB56FDB73B88" pageId="5" pageNumber="474" type="description">
<paragraph id="8B89EB49FFFAFFFFFF70FB56FDB73B88" blockId="5.[151,1438,1081,2006]" pageId="5" pageNumber="474">
<emphasis id="B942375BFFFAFFFFFF70FB56FDB53AFF" box="[199,572,1225,1250]" inLineHeading="true" pageId="5" pageNumber="474" reason="1">
<emphasis id="B942375BFFFAFFFFFF70FB56FDBC3AFC" bold="true" box="[199,565,1225,1250]" pageId="5" pageNumber="474">Description of female genitalia</emphasis>
.
</emphasis>
Egg guide long (
<figureCitation id="130DF7CCFFFAFFFFFCB6FB56FCD23AFC" box="[769,859,1225,1250]" captionStart="FIGURE 5" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[153,1426,1213,1935]" captionTargetId="figure-74@6.[151,1436,1203,1948]" captionTargetPageId="6" captionText="FIGURE 5. Female genitalia of Trichadenotecnum imrum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921636" httpUri="https://zenodo.org/record/4921636/files/figure.png" pageId="5" pageNumber="474">Fig. 5A</figureCitation>
), much longer than basal width, slightly narrowing toward apex, distal margin only slightly arched, medially and laterally with longitudinal membranous regions; body of subgenital plate rather small, posterior margin with short and broad membranous region next to egg guide, anteromedially with broad membranous region. Gonapophyses (
<figureCitation id="130DF7CCFFFAFFFFFCCCFAAAFC5E3B50" box="[891,983,1333,1358]" captionStart="FIGURE 5" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[153,1426,1213,1935]" captionTargetId="figure-74@6.[151,1436,1203,1948]" captionTargetPageId="6" captionText="FIGURE 5. Female genitalia of Trichadenotecnum imrum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921636" httpUri="https://zenodo.org/record/4921636/files/figure.png" pageId="5" pageNumber="474">Fig. 5B</figureCitation>
). Ventral valve long; dorsal valve with long distal process; posterior lobe of external valve small. Spermapore plate (
<figureCitation id="130DF7CCFFFAFFFFFC4BFAC6FBDB3B6C" box="[1020,1106,1369,1394]" captionStart="FIGURE 5" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[153,1426,1213,1935]" captionTargetId="figure-74@6.[151,1436,1203,1948]" captionTargetPageId="6" captionText="FIGURE 5. Female genitalia of Trichadenotecnum imrum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921636" httpUri="https://zenodo.org/record/4921636/files/figure.png" pageId="5" pageNumber="474">Fig. 5C</figureCitation>
) lightly pigmented anteriorly and around spermapore, symmetrical.
</paragraph>
</subSubSection>
<subSubSection id="C32CB8C2FFFAFFFFFF70FA3EFE6939C8" pageId="5" pageNumber="474" type="discussion">
<paragraph id="8B89EB49FFFAFFFFFF70FA3EFE8D3838" blockId="5.[151,1438,1081,2006]" pageId="5" pageNumber="474">
<emphasis id="B942375BFFFAFFFFFF70FA3EFEB23BA7" box="[199,315,1441,1466]" inLineHeading="true" pageId="5" pageNumber="474" reason="1">
<emphasis id="B942375BFFFAFFFFFF70FA3EFEBD3BA4" bold="true" box="[199,308,1441,1466]" pageId="5" pageNumber="474">Remarks</emphasis>
.
</emphasis>
The male specimens examined here match very well with the original description of this species, including genital structures (
<figureCitation id="130DF7CCFFFAFFFFFE53FA5AFDA73BC0" box="[484,558,1477,1502]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Fig. 4</figureCitation>
). This species was originally described from a single male from
<collectingRegion id="49F225ABFFFAFFFFFA84FA5AFA113BC0" box="[1331,1432,1477,1502]" country="Malaysia" name="Selangor" pageId="5" pageNumber="474">Selangor</collectingRegion>
,
<collectingCountry id="F321ABD9FFFAFFFFFF20FA76FE88381C" box="[151,257,1513,1538]" name="Malaysia" pageId="5" pageNumber="474">Malaysia</collectingCountry>
, and is here recorded from
<collectingCountry id="F321ABD9FFFAFFFFFD99FA76FD29381C" box="[558,672,1513,1538]" name="Singapore" pageId="5" pageNumber="474">Singapore</collectingCountry>
for the first time. The female of this species is described here for the first time.
</paragraph>
<paragraph id="8B89EB49FFFAFFFFFF70F9AEFE6939C8" blockId="5.[151,1438,1081,2006]" pageId="5" pageNumber="474">
This species is apparently closely related to
<taxonomicName id="4C3690CAFFFAFFFFFD7EF9ADFCF13857" authorityName="Endang &amp; New" authorityYear="2005" box="[713,888,1586,1609]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFD7EF9ADFCF13857" box="[713,888,1586,1609]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
but can be distinguished from the latter by the flattened epiproct (
<figureCitation id="130DF7CCFFFAFFFFFED8F9CAFE453870" box="[367,460,1621,1646]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Fig. 4A</figureCitation>
), presence of the small projection placed left to the circular-plate on the hypandrium (
<figureCitation id="130DF7CCFFFAFFFFFF28F9E6FF7E388C" box="[159,247,1657,1682]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Fig. 4C</figureCitation>
), and absence of the keel on the trichobothrial field (
<figureCitation id="130DF7CCFFFAFFFFFCF2F9E6FC14388C" box="[837,925,1657,1682]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Fig. 4A</figureCitation>
) in male and the shape of the subgenital plate (
<figureCitation id="130DF7CCFFFAFFFFFF28F902FF7138A8" box="[159,248,1693,1718]" captionStart="FIGURE 5" captionStartId="6.[151,250,1970,1992]" captionTargetBox="[153,1426,1213,1935]" captionTargetId="figure-74@6.[151,1436,1203,1948]" captionTargetPageId="6" captionText="FIGURE 5. Female genitalia of Trichadenotecnum imrum, ventral view. A: subgenital plate; B: gonapophyses; C: spermapore plate." figureDoi="http://doi.org/10.5281/zenodo.4921636" httpUri="https://zenodo.org/record/4921636/files/figure.png" pageId="5" pageNumber="474">Fig. 5A</figureCitation>
) in female. Furthermore, this species is very similar to
<taxonomicName id="4C3690CAFFFAFFFFFCC0F901FC4C38A8" authorityName="Thornton" authorityYear="1961" box="[887,965,1693,1718]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="B942375BFFFAFFFFFCC0F901FC4C38A8" box="[887,965,1693,1718]" italics="true" pageId="5" pageNumber="474">T. felix</emphasis>
</taxonomicName>
in male and female genital structures, as also mentioned for
<taxonomicName id="4C3690CAFFFAFFFFFECBF95DFDA438C7" authorityName="Endang &amp; New" authorityYear="2005" box="[380,557,1730,1753]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFECBF95DFDA438C7" box="[380,557,1730,1753]" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
. These three species apparently compose a monophyletic group within the
<emphasis id="B942375BFFFAFFFFFF20F97AFE9E38E0" box="[151,279,1765,1790]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group based on having the bifurcated posterior process on the phallosome (
<figureCitation id="130DF7CCFFFAFFFFFBC2F97AFB5C38E0" box="[1141,1237,1765,1790]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Figs 4D</figureCitation>
). They also share the following character state: the trichobothrial field is divided into two regions (
<figureCitation id="130DF7CCFFFAFFFFFB83F896FB04393C" box="[1076,1165,1801,1826]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Fig. 4A</figureCitation>
) (
<bibRefCitation id="EFA796B8FFFAFFFFFB11F896FAD5393C" author="Thornton, I. W. B." box="[1190,1372,1801,1826]" pageId="5" pageNumber="474" pagination="1 - 24" refId="ref15445" refString="Thornton, I. W. B. (1961) The Trichadenotecnum group (Psocoptera: Psocidae) in Hong Kong, with descriptions of new species. Transactions of the Royal Entomological Society, London, 113, 1 - 24. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1961. tb 00797. x" type="journal article" year="1961">Thornton, 1961</bibRefCitation>
). The empty space of the trichobothrial field, observed in
<taxonomicName id="4C3690CAFFFAFFFFFD42F8B1FCCD3958" authorityName="Thornton" authorityYear="1961" box="[757,836,1837,1862]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="felix">
<emphasis id="B942375BFFFAFFFFFD42F8B1FCCD3958" box="[757,836,1837,1862]" italics="true" pageId="5" pageNumber="474">T. felix</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3690CAFFFAFFFFFC36F8B1FC60395B" authorityName="New &amp; Thornton" authorityYear="1976" box="[897,1001,1838,1861]" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="imrum">
<emphasis id="B942375BFFFAFFFFFC36F8B1FC60395B" box="[897,1001,1838,1861]" italics="true" pageId="5" pageNumber="474">T. imrum</emphasis>
</taxonomicName>
, corresponds to the keeled area of
<taxonomicName id="4C3690CAFFFAFFFFFA30F8B1FEA13977" authorityName="Endang &amp; New" authorityYear="2005" class="Insecta" family="Psocidae" genus="Trichadenotecnum" kingdom="Animalia" order="Psocodea" pageId="5" pageNumber="474" phylum="Arthropoda" rank="species" species="cinnamonum">
<emphasis id="B942375BFFFAFFFFFA30F8B1FEA13977" italics="true" pageId="5" pageNumber="474">T. cinnamonum</emphasis>
</taxonomicName>
and other species of the
<emphasis id="B942375BFFFAFFFFFDE5F8CEFD583974" box="[594,721,1873,1898]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group (
<figureCitation id="130DF7CCFFFAFFFFFC87F8CEFC043974" box="[816,909,1873,1898]" captionStart="FIGURE 2" captionStartId="4.[151,250,1061,1083]" captionTargetBox="[155,1432,194,1030]" captionTargetId="figure-65@4.[150,1436,193,1039]" captionTargetPageId="4" captionText="FIGURE 2. Male terminalia of Trichadenotecnum cinnamonum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921630" httpUri="https://zenodo.org/record/4921630/files/figure.png" pageId="5" pageNumber="474">Fig. 2A</figureCitation>
). As mentioned above, this situation strongly suggests that the presence of the trichobothrial process/keel is autapomorphic for the
<emphasis id="B942375BFFFAFFFFFB2EF8EAFA903990" box="[1177,1305,1909,1934]" italics="true" pageId="5" pageNumber="474">corniculum</emphasis>
group and plesiomorphic within this group; its absence corresponds to an incomplete reversal leaving the empty area on the trichobothrial field (
<figureCitation id="130DF7CCFFFAFFFFFECEF822FE5A39C8" box="[377,467,1981,2006]" captionStart="FIGURE 4" captionStartId="6.[151,250,1117,1139]" captionTargetBox="[154,1430,182,1095]" captionTargetId="figure-65@6.[151,1435,177,1095]" captionTargetPageId="6" captionText="FIGURE 4. Male terminalia of Trichadenotecnum imrum. A: terminalia, lateral view; B: epiproct, dorsal view; C: hypandrium, ventral view; D: phallosome, ventral view." figureDoi="http://doi.org/10.5281/zenodo.4921634" httpUri="https://zenodo.org/record/4921634/files/figure.png" pageId="5" pageNumber="474">Fig. 4A</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>