354 lines
47 KiB
XML
354 lines
47 KiB
XML
<document id="10E64E804E5B712AA8BBC2124C309D17" ID-DOI="10.5281/zenodo.274183" ID-GBIF-Dataset="d1d5e1b5-3b24-47b5-8879-5ba97ea14340" ID-ISSN="1175-5326" ID-Zenodo-Dep="274183" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459826581406" checkinUser="plazi" docAuthor="Das, Indraneil, Lakim, Maklarin & Kandaung, Pius" docDate="2008" docId="0389BE6AFFF2EA3E83E9FB6B049AFE41" docLanguage="en" docName="zt01719p060.pdf" docOrigin="Zootaxa 1719" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Luperosaurus sorok Das, Lakim & Kandaung, 2008, sp. nov." docType="treatment" docVersion="6" lastPageNumber="60" masterDocId="FFB0C612FFF3EA39837EFFA60404FF86" masterDocTitle="New species of Luperosaurus (Squamata: Gekkonidae) from the Crocker Range Park, Sabah, Malaysia (Borneo)" masterLastPageNumber="60" masterPageNumber="53" pageNumber="54" updateTime="1698222331893" updateUser="plazi">
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<mods:mods id="B51D477C311435183BFFF01064092FD5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="036372A0AF86C309952C5A0A904FFAC8">
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<mods:title id="A000CC8FEB6A588F918B6EFAB8DC9375">New species of Luperosaurus (Squamata: Gekkonidae) from the Crocker Range Park, Sabah, Malaysia (Borneo)</mods:title>
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<mods:name id="34DE9321106BA93CED06E60692126232" type="personal">
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<mods:namePart id="B994AC305C26B151D85071A8567312CA">Das, Indraneil</mods:namePart>
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<mods:roleTerm id="F336D4D7BF8DE5E4BDFC7EFC96D78A90">Author</mods:roleTerm>
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<mods:namePart id="A07324A6BDF86F8AB08152BC7ABA2CA1">Lakim, Maklarin</mods:namePart>
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<mods:name id="B71C176866E3EC9C92F0EA70090218DD" type="personal">
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<mods:roleTerm id="1BE2498D592A7A0C7E8FD8514685F9B7">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="28299452DB585B29CC8E21F3C19EB847">Kandaung, Pius</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="A52F7C5164FCB7B6573A3461AD57652E">text</mods:typeOfResource>
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<mods:title id="1667E69CE1F7E9C242FD2597290FEC6E">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="A72C998364BF12298DBB86FA1D16828E">
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<mods:date id="8397628F00B5B4A92BA56D41D6C0C33C">2008</mods:date>
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<mods:detail id="05B0B410BDD91C37711B4D824A6A4F6A" type="volume">
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<mods:number id="DCA4F68986E8607007A494320DD17D31">1719</mods:number>
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<mods:classification id="26B1A24887CC8EBB89E00F3D5542E310">journal article</mods:classification>
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<mods:identifier id="AC5F4A7389449AC12CC56092349B5092" type="DOI">10.5281/zenodo.274183</mods:identifier>
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<mods:identifier id="9680BDF3C5E5B33FE94511AC2403DA10" type="GBIF-Dataset">d1d5e1b5-3b24-47b5-8879-5ba97ea14340</mods:identifier>
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<mods:identifier id="2F3603A9CC02FC0CB96D6EB16ED5F055" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="BE550FC908F7A275F7D45ED544264AF7" type="Zenodo-Dep">274183</mods:identifier>
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<treatment id="0389BE6AFFF2EA3E83E9FB6B049AFE41" ID-DOI="http://doi.org/10.5281/zenodo.6231769" ID-GBIF-Taxon="119356895" ID-Zenodo-Dep="6231769" LSID="urn:lsid:plazi:treatment:0389BE6AFFF2EA3E83E9FB6B049AFE41" httpUri="http://treatment.plazi.org/id/0389BE6AFFF2EA3E83E9FB6B049AFE41" lastPageId="7" lastPageNumber="60" pageId="1" pageNumber="54">
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<subSubSection id="C33A5CF7FFF2EA3883E9FB6B051CFA8C" pageId="1" pageNumber="54" type="nomenclature">
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<paragraph id="8B9F0F7CFFF2EA3883E9FB6B05EAFB61" blockId="1.[151,494,1229,1290]" box="[151,494,1229,1255]" pageId="1" pageNumber="54">
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<heading id="D0D7B810FFF2EA3883E9FB6B05EAFB61" bold="true" box="[151,494,1229,1255]" fontSize="11" level="1" pageId="1" pageNumber="54" reason="1">
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<emphasis id="B954D36EFFF2EA3883E9FB6B05EAFB61" bold="true" box="[151,494,1229,1255]" pageId="1" pageNumber="54">
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<taxonomicName id="4C2074FFFFF2EA3883E9FB6B058EFB61" box="[151,394,1229,1255]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="54" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
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<emphasis id="B954D36EFFF2EA3883E9FB6B058EFB61" bold="true" box="[151,394,1229,1255]" italics="true" pageId="1" pageNumber="54">Luperosaurus sorok</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A2676E15FFF2EA3882EFFB6B05EAFB61" box="[401,494,1229,1255]" pageId="1" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8B9F0F7CFFF2EA3883E9FB56051CFA8C" blockId="1.[151,494,1229,1290]" box="[151,280,1264,1290]" pageId="1" pageNumber="54">
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(
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<figureCitation id="131B13F9FFF2EA3883E1FB56050AFA8C" box="[159,270,1264,1290]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStartId-0="2.[151,255,1098,1122]" captionStartId-1="4.[151,255,1963,1987]" captionStartId-2="5.[151,255,711,735]" captionStartId-3="5.[151,255,1388,1412]" captionStartId-4="6.[151,255,893,917]" captionTargetBox-0="[269,1315,748,1097]" captionTargetBox-1="[347,1238,210,1917]" captionTargetBox-2="[524,1032,204,697]" captionTargetBox-3="[234,1344,816,1348]" captionTargetBox-4="[292,1310,188,873]" captionTargetId-0="figure@2.[264,1323,727,1099]" captionTargetId-1="figure@4.[266,1239,792,1376]" captionTargetId-2="figure@5.[524,1062,160,698]" captionTargetId-3="figure@5.[234,1368,772,1378]" captionTargetId-4="figure@6.[292,1310,170,885]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="5" captionTargetPageId-4="6" captionText-0="FIGURE 1. Holotype of Luperosaurus sorok sp. nov. (SP 06618), showing entire body in dorsal view. Scale marker = 10 mm." captionText-1="FIGURE 2. Holotype of Luperosaurus sorok sp. nov. (SP 06618), showing head in dorsal (top), ventral (middle) and lateral (bottom) views. Scale markers = 2 mm." captionText-2="FIGURE 3. Holotype of Luperosaurus sorok sp. nov. (SP 06618), showing sole and scansors of left hindlimb. Scale marker = 1 mm." captionText-3="FIGURE 4. Holotype of Luperosaurus sorok sp. nov. (SP 06618), showing ventral surface of tail. Scale marker = 4 mm." captionText-4="FIGURE 5. Holotype of Luperosaurus sorok sp. nov. (SP 06618), showing scales of dorsum at midbody. Scale marker = 2 mm." httpUri-0="https://zenodo.org/record/274184/files/figure.png" httpUri-1="https://zenodo.org/record/274185/files/figure.png" httpUri-2="https://zenodo.org/record/274186/files/figure.png" httpUri-3="https://zenodo.org/record/274187/files/figure.png" httpUri-4="https://zenodo.org/record/274188/files/figure.png" pageId="1" pageNumber="54">Figs. 1–5</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A5CF7FFF2EA3883E9FAE60513FA2C" pageId="1" pageNumber="54" type="materials_examined">
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<paragraph id="8B9F0F7CFFF2EA3883E9FAE60513FA2C" blockId="1.[151,1437,1344,2010]" pageId="1" pageNumber="54">
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<emphasis id="B954D36EFFF2EA3883E9FAE60516FADC" bold="true" box="[151,274,1344,1370]" pageId="1" pageNumber="54">
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<typeStatus id="549BB1DEFFF2EA3883E9FAE6050AFADC" box="[151,270,1344,1370]" pageId="1" pageNumber="54" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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SP 0 6618, female, from near bank of Sungai Bariawa, Crocker Range Park (05º24'177"N, 116º07'897"E;
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<figureCitation id="131B13F9FFF2EA388235FACE0592FA04" box="[331,406,1384,1410]" captionStart="FIGURE 6" captionStartId="6.[151,255,1974,1998]" captionTargetBox="[398,1192,1000,1951]" captionTargetId="figure@6.[398,1192,1000,1956]" captionTargetPageId="6" captionText="FIGURE 6. Map of Borneo, showing the type locality (indicated with a white spot) of Luperosaurus sorok sp. nov. in the Crocker Range Park, Sabah." httpUri="https://zenodo.org/record/274189/files/figure.png" pageId="1" pageNumber="54">Fig. 6</figureCitation>
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), Keningau District, Sabah,
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<collectingCountry id="F3374FECFFF2EA38818DFACE075AFA04" box="[755,862,1384,1410]" name="Malaysia" pageId="1" pageNumber="54">Malaysia</collectingCountry>
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(Borneo), collected
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<date id="FF9E29BCFFF2EA388722FACE0131FA04" box="[1116,1333,1384,1410]" pageId="1" pageNumber="54" value="2002-02-19">19 February 2002</date>
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by Pius Kandaung.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A5CF7FFF2EA3883BBFA1E05ECF96C" pageId="1" pageNumber="54" type="diagnosis">
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<paragraph id="8B9F0F7CFFF2EA3883BBFA1E05ECF96C" blockId="1.[151,1437,1344,2010]" pageId="1" pageNumber="54">
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<emphasis id="B954D36EFFF2EA3883BBFA1E0543FA54" bold="true" box="[197,327,1464,1490]" pageId="1" pageNumber="54">Diagnosis:</emphasis>
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A small species of
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<taxonomicName id="4C2074FFFFF2EA388152FA1E06CAFA57" box="[556,718,1464,1489]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
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<emphasis id="B954D36EFFF2EA388152FA1E06CAFA57" box="[556,718,1464,1489]" italics="true" pageId="1" pageNumber="54">Luperosaurus</emphasis>
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</taxonomicName>
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(SVL
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<quantity id="4CD8A299FFF2EA388060FA1E078EFA54" box="[798,906,1464,1490]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="1" pageNumber="54" unit="mm" value="34.7">34.7 mm</quantity>
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), diagnosable from congeneric species by the possession of the following combination of characters: body robust; head narrower than body; auricular opening oval-squarish; subrictal tubercles present; two intersupranasals contacting rostral; rostral contacts nares; nine supralabials to the middle of the orbital; 14/15 supralabials to posterior edge of orbit; 11 infralabials; 1–3 anteriormost chin scales subequal, larger than those to their posterior; dorsal body scales rounded, convex and granular; ca. 104 dorsal scale rows; 45 ventral scale rows; ca. 149 midbody scale rows; ventrolateral body tubercles spinose; three distal scansors deeply notched; tail fringe with distinct serrations; and pale rounded spots present on middorsum.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33A5CF7FFF2EA3A83BBF95105A6FA94" lastPageId="3" lastPageNumber="56" pageId="1" pageNumber="54" type="description">
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<paragraph id="8B9F0F7CFFF2EA3B83BBF951063FFD3C" blockId="1.[151,1437,1344,2010]" lastBlockId="2.[151,1437,152,698]" lastPageId="2" lastPageNumber="55" pageId="1" pageNumber="54">
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<emphasis id="B954D36EFFF2EA3883BBF95105F6F897" bold="true" box="[197,498,1783,1809]" pageId="1" pageNumber="54">
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Description of
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<typeStatus id="549BB1DEFFF2EA388203F95105E9F897" box="[381,493,1783,1809]" pageId="1" pageNumber="54" type="holotype">holotype</typeStatus>
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:
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</emphasis>
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A small bodied species of
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<taxonomicName id="4C2074FFFFF2EA38804EF95E07D6F897" box="[816,978,1784,1809]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="1" pageNumber="54" phylum="Chordata" rank="genus">
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<emphasis id="B954D36EFFF2EA38804EF95E07D6F897" box="[816,978,1784,1809]" italics="true" pageId="1" pageNumber="54">Luperosaurus</emphasis>
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</taxonomicName>
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(SVL
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<quantity id="4CD8A299FFF2EA38875CF95E008AF894" box="[1058,1166,1784,1810]" metricMagnitude="-2" metricUnit="m" metricValue="3.47" pageId="1" pageNumber="54" unit="mm" value="34.7">34.7 mm</quantity>
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); snout short (HL/SVL ratio 0.18), fairly wide (HW/SVL ratio 0.18), depressed (HD/HL ratio 0.75); snout tip tapered and rounded in dorsal and lateral aspects; head distinct from neck; supraorbital crest prominent; interorbital region flattened, traversed by 24 scales; snout less than twice eye diameter (ED/E-S ratio 0.64); scales on snout and forehead small, rounded, keeled, with larger, scattered scales; scales on snout larger than those on occipital region; enlarged spinose tubercles in postocular region; eyes large (ED/HL ratio 0.36); orbits of eyes lacking ‘extrabrillar fringes’ (of
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<bibRefCitation id="EFB1728DFFF1EA3B820FFF3E064DFF34" author="Underwood" box="[369,585,152,178]" pageId="2" pageNumber="60" refString="Underwood, G. (1954) On the classification and evolution of geckos. Proceedings of the Zoological Society of London, 124, 469 - 492." type="journal article" year="1954">Underwood, 1954</bibRefCitation>
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); canthus rostralis weakly concave, lores sloping; pupil vertically elliptical, with crenulated edges; no elongate supraciliaries; auricular opening deep, oval-squarish (EL/E-S ratio 0.02); auricular opening with a single, pale, enlarged lobule on top of ear opening; eye to ear distance greater than diameter of eyes (E-E/ED ratio 1.32); spinose subrictal tubercles, set obliquely along mandible, forming an irregular row; no tubercles anterior to tympanum; rostral trapezoidal, lacking a rostral groove; rostral about half as deep as wide (width =
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<quantity id="4CD8A299FFF1EA3B8288FEC60657FEFC" box="[502,595,352,378]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="55" unit="mm" value="1.5">1.5 mm</quantity>
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, depth =
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<quantity id="4CD8A299FFF1EA3B81C1FEC60719FEFC" box="[703,797,352,378]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="55" unit="mm" value="0.8">0.8 mm</quantity>
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; width/depth ratio 1.9), contacted posteriorly by narial opening, an intersupranasal, and two supranasals that are in narrow contact ventro-posteriorly with Supralabial I; individual nares oval, oriented laterally, in contact with rostral, supranasal, Supralabial I, and two postnasals; two squarish intersuprarostrals bordering rostral separate supranasals; three postnasals bound nasal; three scale rows separate orbit from supralabials; supralabial IX (L/R) in midorbital position, followed by 6/5 (L/R) smaller supralabials to point of posterior orbit of eye; infralabials 11 to edge of jaws; mental subtriangular, slightly deeper than wide; 1–3 postmentals, rounded, smaller than mental, subequal, 2–3 times larger than those to their posterior; tongue narrowly elongate, with a weak median cleft; enlarged extracranial endolymphatic sac present on nuchal region.
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</paragraph>
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<caption id="DF5F5FF4FFF1EA3B83E9FBEC04EDFB02" httpUri="https://zenodo.org/record/274184/files/figure.png" pageId="2" pageNumber="55" targetBox="[269,1315,748,1097]" targetPageId="2">
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<paragraph id="8B9F0F7CFFF1EA3B83E9FBEC04EDFB02" blockId="2.[151,1436,1098,1156]" pageId="2" pageNumber="55">
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<emphasis id="B954D36EFFF1EA3B83E9FBEC051EFBE4" bold="true" box="[151,282,1098,1122]" pageId="2" pageNumber="55">FIGURE 1.</emphasis>
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Holotype of
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<taxonomicName id="4C2074FFFFF1EA3B82D5FBEC0684FBE7" box="[427,640,1098,1121]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="2" pageNumber="55" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
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<emphasis id="B954D36EFFF1EA3B82D5FBEC0684FBE7" box="[427,640,1098,1121]" italics="true" pageId="2" pageNumber="55">Luperosaurus sorok</emphasis>
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</taxonomicName>
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<emphasis id="B954D36EFFF1EA3B81F6FBEC06DBFBE4" bold="true" box="[648,735,1098,1122]" pageId="2" pageNumber="55">
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<taxonomicNameLabel id="A2676E15FFF1EA3B81F6FBEC06DBFBE4" box="[648,735,1098,1122]" pageId="2" pageNumber="55" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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(SP 06618), showing entire body in dorsal view. Scale marker = 10 mm.
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</paragraph>
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</caption>
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<paragraph id="8B9F0F7CFFF1EA3B83BBFB1E00EAF934" blockId="2.[151,1437,1208,2034]" pageId="2" pageNumber="55">Head and body robust; body dorsoventrally flattened, relatively stout, somewhat elongate (A-G/SVL ratio 0.51); dermal spines and cutaneous expansion of axilla-groin region absent; tubercles on dorsum low, rounded, in irregular rows; body scales juxtaposed, lacking interstitial granules; scales along the vertebral region not differentiated; ventrals smooth, imbricate; chin shields smaller than ventrals; scales on gular, pectoral and abdominal regions subequal; dorsal scales at midbody much smaller than ventral scales; ca. 149 transverse body scale rows; 45 enlarged ventral scales; distinct demarcation between enlarged, flattened, ventral scales and the smaller, more rounded dorsal scales on lower flanks; ventrolateral body tubercles spinose; scales on ventral surfaces of manus and pes smooth, rounded; scales on inner surface of fore- and hindlimbs conical; those on outer surface of fore- and hindlimbs weakly spinose; on preanal region, scales on anteroventral surfaces as large as those on venter of body, while those on posteroventral surfaces, closer to cloaca, much smaller; femoral scales undifferentiated in size; preanal depression or groove absent; preanofemoral pores absent, but preanofemoral scales enlarged, differentiated; cloacal orifice bowed posteriorly, bordered anteriorly with a bowed integumentary fold; postcloacal glandular openings posterior to vent absent.</paragraph>
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<paragraph id="8B9F0F7CFFF1EA3A83BBF9660682FEFC" blockId="2.[151,1437,1208,2034]" lastBlockId="3.[151,1437,152,2018]" lastPageId="3" lastPageNumber="56" pageId="2" pageNumber="55">Forelimbs moderately short, stout; forearm short; hindlimbs short and stout; tibia short (TBL/SVL ratio 0.15); digits relatively long, weakly dilated, all clawed except for Finger I, which exhibits a small, recurved claw inside its digital tip; scansors narrow, subrectangular, unnotched, except for the 3–4 most distal scansors that are distinctly notched; basal subdigital lamellae fragmented up to mid-level of each digit; enlarged lamellae (inclusive of terminal one) numbering on manus I (5/4); II (8/8); III (11/11); IV (9/11); and V (9/9); and on pes I (8/8); II (12/11); III (10/10); IV (8/10); and V (10/19); on Finger IV and Toes III and IV, non-enlarged scales reach around midlevel of respective phalange; interdigital webbing present between all adjacent phalanges, extending beyond the mid-level of Toes III and IV; relative length of digits (finger): IV> III> V> II> I; (toe): IV> III> V> II> I; anterior edges of forearm with a weak dermal fringe; anterior edge of hindlimb at both femur and tibia lacking dermal fringe; posterior edge of forearm with a weak cutaneous fringe extending from the carpal region to axilla; posterior edge of hindlimbs with a distinct dermal fringe, extending from the base of pes below Toe V to base of thighs, and is widest at level of the knee joint, and with a distinct notch ca. 10% fringe length towards inner edge; cutaneous expansions with small, subimbricate scales; a single large pale spine on outer surface of tarsus.</paragraph>
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBFE2E06ECFC8C" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">Tail, original, subcylindrical at base, with a weak basal constriction and slightly flattened dorsoventrally (TW/TD ratio = 1.8) along rest of tail body, shorter than snout-vent length (TL/SVL ratio 0.69); tail body slen- der, gradually tapering; lateral margins of tail with regular serrations, each "tooth" composed of 7–8 scales forming 18 enlarged spinous fringed structures on each side of tail, the peripheral ones, especially those towards distal portion of tail being the least serrate; tail terminates in a rounded point; a single, small, cloacal spur on each side of vent; tail covered with small, cycloid scales and weakly segmented, with 5–10 irregular, transverse rows of enlarged, spinous scales; remaining caudals small, granular; subcaudals more flattened, cycloid; median subcaudal series not transversely enlarged, only slightly larger than adjacent scales; scales on postcloacal region and base smaller than on rest of tail; a low median tail furrow but no lateral tail furrows; a small spinous postcloacal spur on each side of tail.</paragraph>
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBFCB104C8FC04" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<emphasis id="B954D36EFFF0EA3A83BBFCB105F1FCB7" bold="true" box="[197,501,791,817]" pageId="3" pageNumber="56">Measurements (in mm):</emphasis>
|
||
SVL 34.7; TL 23.9; TW 4.1; TD 2.3; BW 8.4; FA 4.4; CL 5.3; TBL 5.1; A-G 17.8; HL 6.1; HW 6.3; HD 4.6; ED 2.2; E-S 3.4; E-N 2.8; IO 3.7; EL 0.07; IN 1.5; TIVL 3.6; and TIVW
|
||
<quantity id="4CD8A299FFF0EA3A8609FCE604CDFC04" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="3" pageNumber="56" unit="mm" value="1.1">1.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBFC290767FC54" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<emphasis id="B954D36EFFF0EA3A83BBFC290540FC2F" bold="true" box="[197,324,911,937]" pageId="3" pageNumber="56">Pholidosis</emphasis>
|
||
: Supralabials (midorbital) 9/9; infralabials 11/11; interorbitals 24; dorsal scale rows ca. 104; transverse body scale rows ca. 149; and enlarged ventrals 45.
|
||
</paragraph>
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBFC7905A6FA94" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<emphasis id="B954D36EFFF0EA3A83BBFC790552FC7F" bold="true" box="[197,342,991,1017]" pageId="3" pageNumber="56">Coloration:</emphasis>
|
||
Pale gray dorsally, with dark gray double chevron-marks that are fused middorsally, numbering one on forehead, four on torso, between nuchal and caudal constriction, and one on pelvic region; the last comprising two isolated, subtriangular patches; six dark transverse caudal bands on dorsum of tail; each of these dark areas on dorsum edged with black; dorsal surfaces of limbs pale gray, with dark bands; lateral fringes of tail pale gray, paler than the body of tail; chin pale, unpigmented; throat yellowish-cream, with darker speckles; rest of venter, including pectoral and abdominal region as well as under surfaces of limbs and tail yellowish-cream, with darker variegation; and undersurface of tail yellowish-cream, with darker smudges. Color in life unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A5CF7FFF0EA3A83BBFAB90097F984" pageId="3" pageNumber="56" type="etymology">
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBFAB904ECFAE4" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<emphasis id="B954D36EFFF0EA3A83BBFAB90552FABF" bold="true" box="[197,342,1311,1337]" pageId="3" pageNumber="56">Etymology:</emphasis>
|
||
Derived from Malay, meaning "to hide", in reference to its presumed ecologically cryptic nature.
|
||
</paragraph>
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBFAC90097F984" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<emphasis id="B954D36EFFF0EA3A83BBFAC90536FA0F" bold="true" box="[197,306,1391,1417]" pageId="3" pageNumber="56">Ecology:</emphasis>
|
||
Nothing is known of the natural history of the new
|
||
<emphasis id="B954D36EFFF0EA3A80E5FAD60089FA0F" box="[923,1165,1392,1417]" italics="true" pageId="3" pageNumber="56">
|
||
<taxonomicName id="4C2074FFFFF0EA3A80E5FAD6008DFA0F" box="[923,1161,1392,1417]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="sorok">Luperosaurus sorok</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
The
|
||
<typeStatus id="549BB1DEFFF0EA3A87B4FAD60135FA0C" box="[1226,1329,1392,1418]" pageId="3" pageNumber="56" type="holotype">holotype</typeStatus>
|
||
was collected from a secondary forest within the Park, and was discovered on a wet rock, on a cloudy and windy day. Crocker Range Park itself includes a large area of primary forest, encompassing both submontane and montane vegetation, and has a rich herpetofauna (discussed in
|
||
<bibRefCitation id="EFB1728DFFF0EA3A803EFA4E07BEF984" author="Das" box="[832,954,1512,1538]" pageId="3" pageNumber="60" refString="Das, I. (2006) Crocker Range National Park, Sabah, as a refuge for Borneo's montane herpetofauna. Amphibian & Reptile Conservation, 4 (2), 3 - 11." type="journal article" year="2006">Das, 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB1728DFFF0EA3A80B9FA4E0083F984" author="Hee" box="[967,1159,1512,1538]" pageId="3" pageNumber="60" refString="Hee, K. - B. H., Sudin, A., Matsui, M. & Mohamed, M. (2004) Notes on the anurans of Crocker Range Park. In: Crocker Range Scientific Expedition 2002. pp: 103 - 112. M. Mohamed, Z. Hamzah, T. Tachi & J. Nais (Eds). Universiti Malaysia Sabah, Kota Kinabalu, Malaysia." type="journal article" year="2004">
|
||
Hee
|
||
<emphasis id="B954D36EFFF0EA3A8083FA4E003CF987" box="[1021,1080,1512,1537]" italics="true" pageId="3" pageNumber="56">et al.</emphasis>
|
||
, 2004
|
||
</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A5CF7FFF0EA3A83BBF9A90701F9D4" pageId="3" pageNumber="56" type="description">
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBF9A90701F9D4" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<emphasis id="B954D36EFFF0EA3A83BBF9A90571F9AF" bold="true" box="[197,373,1551,1577]" pageId="3" pageNumber="56">Comparisons:</emphasis>
|
||
The new species from Sabah is compared with congeners for Borneo and other parts of south-east Asia (see also
|
||
<bibRefCitation id="EFB1728DFFF0EA3A82C1F9910653F9D4" author="Rosler" box="[447,599,1591,1618]" pageId="3" pageNumber="60" refString="Rosler, H. (2000) Kommentierte Liste der rezenten, subrezenten und fossilen Gecko-Taxa (Reptilia: Gekkomorpha). Gekkota, 2, 28 - 153." type="journal article" year="2000">Rősler, 2000</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="EFB1728DFFF0EA3A811DF99E06FCF9D4" author="Kluge" box="[611,760,1592,1618]" pageId="3" pageNumber="60" refString="Kluge, A. G. (2001) Gekkotan lizard taxonomy. Hamadryad, 26 (1), 1 - 209." type="journal article" year="2001">Kluge, 2001</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33A5CF7FFF0EA3C83BBF9C60089F85F" lastPageId="5" lastPageNumber="58" pageId="3" pageNumber="56" type="reference_group">
|
||
<paragraph id="8B9F0F7CFFF0EA3A83BBF9C60566F864" blockId="3.[151,1437,152,2018]" pageId="3" pageNumber="56">
|
||
<taxonomicName id="4C2074FFFFF0EA3A83BBF9C605ABF9FF" box="[197,431,1632,1657]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
|
||
<emphasis id="B954D36EFFF0EA3A83BBF9C605ABF9FF" box="[197,431,1632,1657]" italics="true" pageId="3" pageNumber="56">Luperosaurus sorok</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954D36EFFF0EA3A82C9F9F90611F9FF" bold="true" box="[439,533,1631,1657]" pageId="3" pageNumber="56">
|
||
<taxonomicNameLabel id="A2676E15FFF0EA3A82C9F9F90611F9FF" box="[439,533,1631,1657]" pageId="3" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from
|
||
<taxonomicName id="4C2074FFFFF0EA3A81CFF9C607D9F9FC" authority="Ota, Sengoku" authorityName="Ota, Sengoku" box="[689,989,1632,1658]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="yasumai">
|
||
<emphasis id="B954D36EFFF0EA3A81CFF9C60735F9FF" box="[689,817,1632,1657]" italics="true" pageId="3" pageNumber="56">L. yasumai</emphasis>
|
||
<bibRefCitation id="EFB1728DFFF0EA3A8044F9C607D9F9FC" author="Ota" box="[826,989,1632,1658]" pageId="3" pageNumber="60" refString="Ota, H., Sengoku, S. & Hikida, T. (1996) Two new species of Luperosaurus (Reptilia: Gekkonidae) from Borneo. Copeia, 1996 (2), 433 - 439." type="journal article" year="1996">Ota, Sengoku</bibRefCitation>
|
||
</taxonomicName>
|
||
, & Hikida, 1996 (distribution: eastern Kalimantan,
|
||
<collectingCountry id="F3374FECFFF0EA3A8250F92E05A6F924" box="[302,418,1672,1698]" name="Indonesia" pageId="3" pageNumber="56">Indonesia</collectingCountry>
|
||
) by the presence of a tail fringe with distinct serrations (vs. tail fringe with an irregular outline); head narrower than body (vs. subequal); auricular opening oval-squarish (vs. horizontally elongated); intersupranasals 2 (vs. 3); subrictal tubercles present (vs. absent); and pale rounded spots on middorsum (vs. indistinct, cloudy markings on middorsum). The new species differs from
|
||
<taxonomicName id="4C2074FFFFF0EA3A8700F8A60198F89C" authority="Russell, 1979" authorityName="Russell" authorityYear="1979" box="[1150,1436,1792,1818]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="3" pageNumber="56" phylum="Chordata" rank="species" species="browni">
|
||
<emphasis id="B954D36EFFF0EA3A8700F8A600F5F89F" box="[1150,1265,1792,1817]" italics="true" pageId="3" pageNumber="56">L. browni</emphasis>
|
||
<bibRefCitation id="EFB1728DFFF0EA3A8784F8A60198F89C" author="Russell" box="[1274,1436,1792,1818]" pageId="3" pageNumber="60" refString="Russell, A. P. (1979) A new species of Luperosaurus (Gekkonidae) with comments on the genus. Herpetologica, 35, 282 - 288." type="journal article" year="1979">Russell, 1979</bibRefCitation>
|
||
</taxonomicName>
|
||
(distribution:
|
||
<collectingCountry id="F3374FECFFF0EA3A8246F88E05A4F8C4" box="[312,416,1832,1858]" name="Malaysia" pageId="3" pageNumber="56">Sarawak</collectingCountry>
|
||
, Kalimantan, West
|
||
<collectingCountry id="F3374FECFFF0EA3A81F6F88E06F7F8C4" box="[648,755,1832,1858]" name="Malaysia" pageId="3" pageNumber="56">Malaysia</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3374FECFFF0EA3A8050F88E07ADF8C4" box="[814,937,1832,1858]" name="Singapore" pageId="3" pageNumber="56">Singapore</collectingCountry>
|
||
; see
|
||
<bibRefCitation id="EFB1728DFFF0EA3A809AF88E00A4F8C4" author="Ota" box="[996,1184,1832,1858]" pageId="3" pageNumber="60" refString="Ota, H., Sengoku, S. & Hikida, T. (1996) Two new species of Luperosaurus (Reptilia: Gekkonidae) from Borneo. Copeia, 1996 (2), 433 - 439." type="journal article" year="1996">
|
||
Ota
|
||
<emphasis id="B954D36EFFF0EA3A8768F88E0056F8C7" box="[1046,1106,1832,1857]" italics="true" pageId="3" pageNumber="56">et al.</emphasis>
|
||
, 1996
|
||
</bibRefCitation>
|
||
; K. K. P.
|
||
<bibRefCitation id="EFB1728DFFF0EA3A866DF88E055CF8EC" author="Lim" pageId="3" pageNumber="60" refString="Lim, K. K. P. & Lim, N. T. - L. (2006) Luperosaurus browni (Brown's flap-legged gecko). Herpetological Review, 37 (1), 107." type="journal article" year="2006">Lim and N. T.-L. Lim, 2006</bibRefCitation>
|
||
) by the possession of convex (vs. flattened) dorsal scales; lateral edges of tail fringed with regularly arranged scales grouped into saw-like tooth (vs. isolated spinous scales); head narrower than body (vs. subequal); intersupranasals 2 (vs. 1); genials enlarged (vs. undifferentiated); and midbody scales number ca. 149 (vs. 129).
|
||
</paragraph>
|
||
<caption id="DF5F5FF4FFF7EA3D83E9F80D0668F863" httpUri="https://zenodo.org/record/274185/files/figure.png" pageId="4" pageNumber="57" targetBox="[347,1238,210,1917]" targetPageId="4">
|
||
<paragraph id="8B9F0F7CFFF7EA3D83E9F80D0668F863" blockId="4.[151,1436,1963,2021]" pageId="4" pageNumber="57">
|
||
<emphasis id="B954D36EFFF7EA3D83E9F80D051CF845" bold="true" box="[151,280,1963,1987]" pageId="4" pageNumber="57">FIGURE 2.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C2074FFFFF7EA3D82DAF80D067CF844" box="[420,632,1963,1986]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="4" pageNumber="57" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
|
||
<emphasis id="B954D36EFFF7EA3D82DAF80D067CF844" box="[420,632,1963,1986]" italics="true" pageId="4" pageNumber="57">Luperosaurus sorok</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954D36EFFF7EA3D8100F80D06D7F845" bold="true" box="[638,723,1963,1987]" pageId="4" pageNumber="57">
|
||
<taxonomicNameLabel id="A2676E15FFF7EA3D8100F80D06D7F845" box="[638,723,1963,1987]" pageId="4" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(SP 06618), showing head in dorsal (top), ventral (middle) and lateral (bottom) views. Scale markers = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F5FF4FFF6EA3C83E9FD610540FC84" httpUri="https://zenodo.org/record/274186/files/figure.png" pageId="5" pageNumber="58" targetBox="[524,1032,204,697]" targetPageId="5">
|
||
<paragraph id="8B9F0F7CFFF6EA3C83E9FD610540FC84" blockId="5.[151,1436,711,770]" pageId="5" pageNumber="58">
|
||
<emphasis id="B954D36EFFF6EA3C83E9FD61051FFD59" bold="true" box="[151,283,711,735]" pageId="5" pageNumber="58">FIGURE 3.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C2074FFFFF6EA3C82CEFD6E0682FD59" box="[432,646,712,735]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
|
||
<emphasis id="B954D36EFFF6EA3C82CEFD6E0682FD59" box="[432,646,712,735]" italics="true" pageId="5" pageNumber="58">Luperosaurus sorok</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954D36EFFF6EA3C81EEFD6106ECFD59" bold="true" box="[656,744,711,735]" pageId="5" pageNumber="58">
|
||
<taxonomicNameLabel id="A2676E15FFF6EA3C81EEFD6106ECFD59" box="[656,744,711,735]" pageId="5" pageNumber="58" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(SP 06618), showing sole and scansors of left hindlimb. Scale marker = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F5FF4FFF6EA3C83E9FACA0199FA05" box="[151,1437,1388,1412]" httpUri="https://zenodo.org/record/274187/files/figure.png" pageId="5" pageNumber="58" targetBox="[234,1344,816,1348]" targetPageId="5">
|
||
<paragraph id="8B9F0F7CFFF6EA3C83E9FACA0199FA05" blockId="5.[151,1437,1388,1412]" box="[151,1437,1388,1412]" pageId="5" pageNumber="58">
|
||
<emphasis id="B954D36EFFF6EA3C83E9FACA051DFA02" bold="true" box="[151,281,1388,1412]" pageId="5" pageNumber="58">FIGURE 4.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C2074FFFFF6EA3C82DBFACB067DFA02" box="[421,633,1389,1412]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
|
||
<emphasis id="B954D36EFFF6EA3C82DBFACB067DFA02" box="[421,633,1389,1412]" italics="true" pageId="5" pageNumber="58">Luperosaurus sorok</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2676E15FFF6EA3C8101FACA06D0FA02" box="[639,724,1388,1412]" pageId="5" pageNumber="58" rank="species">
|
||
<emphasis id="B954D36EFFF6EA3C8101FACA06D0FA02" bold="true" box="[639,724,1388,1412]" pageId="5" pageNumber="58">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
(SP 06618), showing ventral surface of tail. Scale marker = 4 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B9F0F7CFFF6EA3C83BBFA110089F85F" blockId="5.[151,1437,1463,2009]" pageId="5" pageNumber="58">
|
||
In this section, we compare the new species from Sabah with populations from Sumatra, Sulawesi, and the
|
||
<collectingCountry id="F3374FECFFF6EA3C83E9FA79051BFA7F" box="[151,287,1503,1529]" name="Philippines" pageId="5" pageNumber="58">Philippines</collectingCountry>
|
||
. Only opposing suite of characters are listed:
|
||
<taxonomicName id="4C2074FFFFF6EA3C8027FA460087FA7F" authority="Boulenger, 1920" authorityName="Boulenger" authorityYear="1920" box="[857,1155,1503,1529]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="brooksi">
|
||
<emphasis id="B954D36EFFF6EA3C8027FA4607ABFA7F" box="[857,943,1504,1529]" italics="true" pageId="5" pageNumber="58">brooksi</emphasis>
|
||
Boulenger, 1920
|
||
</taxonomicName>
|
||
(distribution: Sumatra,
|
||
<collectingCountry id="F3374FECFFF6EA3C83E9F9A1050AF9A7" box="[151,270,1543,1569]" name="Indonesia" pageId="5" pageNumber="58">Indonesia</collectingCountry>
|
||
), subrictal tubercles absent; genials small; and a single intersupranasal contacting rostral;
|
||
<taxonomicName id="4C2074FFFFF6EA3C8647F9AE055CF9CF" authority="Gaulke, Roesler" authorityName="Gaulke, Roesler" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="corfieldi">
|
||
<emphasis id="B954D36EFFF6EA3C8647F9AE0199F9A7" box="[1337,1437,1544,1569]" italics="true" pageId="5" pageNumber="58">corfieldi</emphasis>
|
||
<bibRefCitation id="EFB1728DFFF6EA3C83E9F989055CF9CF" author="Gaulke" box="[151,344,1583,1609]" pageId="5" pageNumber="60" refString="Gaulke, M., Roesler, H. & Brown, R. M. (2007) A new species of Luperosaurus (Squamata: Gekkonidae) from Panay Island, Philippines, with comments on the taxonomic status of Luperosaurus cumingii (Gray, 1845). Copeia, 2007 (2), 413 - 423." type="journal article" year="2007">Gaulke, Roesler</bibRefCitation>
|
||
</taxonomicName>
|
||
, & Brown, 2007 (distribution: Panay,
|
||
<collectingCountry id="F3374FECFFF6EA3C8057F98907B6F9CF" box="[809,946,1583,1609]" name="Philippines" pageId="5" pageNumber="58">Philippines</collectingCountry>
|
||
), wide cutanaeous expansion on anterior forelimbs present; enlarged ventrolateral tail tubercles absent; spinous scales on edges of anterior cutaneous expansions of limbs present; females with 11 preanal pores; and dorsum with three dark chevrons between fore- and hindlimbs;
|
||
<taxonomicName id="4C2074FFFFF6EA3C82F2F90E0677F947" authority="Gray, 1845" authorityName="Gray" authorityYear="1845" box="[396,627,1703,1729]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="cumingi">
|
||
<emphasis id="B954D36EFFF6EA3C82F2F90E05EEF947" box="[396,490,1704,1729]" italics="true" pageId="5" pageNumber="58">cumingi</emphasis>
|
||
<bibRefCitation id="EFB1728DFFF6EA3C828FF9010677F947" author="Gray" box="[497,627,1703,1729]" pageId="5" pageNumber="60" refString="Gray, J. E. (1845) Catalogue of the specimens of lizards in the collection of the British Museum. British Museum (Natural History), London, United Kingdom. xxviii + 289 pp." type="book" year="1845">Gray, 1845</bibRefCitation>
|
||
</taxonomicName>
|
||
(distribution: Luzon,
|
||
<collectingCountry id="F3374FECFFF6EA3C800CF90107FFF947" box="[882,1019,1703,1729]" name="Philippines" pageId="5" pageNumber="58">Philippines</collectingCountry>
|
||
), infralabials 13–15; preanofemoral pores 15–20; scansors on Toe I, 11–14; on Toe III, 13–16; ventrals small, granular; wide flaps on anterior edge of forelimbs; dermal fringe present on both anterior and posterior edges of hindlimbs; and subrictal tubercles absent;
|
||
<taxonomicName id="4C2074FFFFF6EA3C838CF8860638F8BF" authority="Brown, Supriatna" authorityName="Brown, Supriatna" box="[242,572,1823,1849]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="iskandari">
|
||
<emphasis id="B954D36EFFF6EA3C838CF8860565F8BF" box="[242,353,1824,1849]" italics="true" pageId="5" pageNumber="58">iskandari</emphasis>
|
||
<bibRefCitation id="EFB1728DFFF6EA3C8217F8B90638F8BF" author="Brown" box="[361,572,1823,1849]" pageId="5" pageNumber="60" refString="Brown, R. M., Supriatna, J. & Ota, H. (2000) Discovery of a new species of Luperosaurus (Squamata; Gekkonidae) from Sulawesi, with a phylogenetic analysis of the genus, and comments on the status of Luperosaurus serraticaudus. Copeia, 2000 (1), 191 - 209." type="journal article" year="2000">Brown, Supriatna</bibRefCitation>
|
||
</taxonomicName>
|
||
, & Ota, 2000 (distribution: Sulawesi,
|
||
<collectingCountry id="F3374FECFFF6EA3C877FF8B9007CF8BF" box="[1025,1144,1823,1849]" name="Indonesia" pageId="5" pageNumber="58">Indonesia</collectingCountry>
|
||
), dorsal body scales flat, hexagonal; anterior forelimbs with wide flaps; a single intersupranasal contacts rostral; penultimate scansors bowed; and tail lacks lateral tubercles;
|
||
<taxonomicName id="4C2074FFFFF6EA3C8122F8D60776F80F" authority="Taylor, 1918" authorityName="Taylor" authorityYear="1918" box="[604,882,1903,1929]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="joloensis">
|
||
<emphasis id="B954D36EFFF6EA3C8122F8D606C0F80F" box="[604,708,1904,1929]" italics="true" pageId="5" pageNumber="58">joloensis</emphasis>
|
||
(Taylor, 1918)
|
||
</taxonomicName>
|
||
(distribution: Jolo and Mindanao,
|
||
<collectingCountry id="F3374FECFFF6EA3C8678F8C90194F80F" box="[1286,1424,1903,1929]" name="Philippines" pageId="5" pageNumber="58">Philippines</collectingCountry>
|
||
); auricular opening narrow, oblique; enlarged genials; a single intersupranasal contacts rostral; spine-like laterals; and subrictal tubercles absent;
|
||
<taxonomicName id="4C2074FFFFF6EA3C814EF866073EF85F" authority="Brown, Diesmos" authorityName="Brown, Diesmos" box="[560,826,1983,2009]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="5" pageNumber="58" phylum="Chordata" rank="species" species="kubli">
|
||
<emphasis id="B954D36EFFF6EA3C814EF866066FF85F" box="[560,619,1984,2009]" italics="true" pageId="5" pageNumber="58">kubli</emphasis>
|
||
<bibRefCitation id="EFB1728DFFF6EA3C810CF819073EF85F" author="Brown" box="[626,826,1983,2009]" pageId="5" pageNumber="60" refString="Brown, R. M., Diesmos, A. C. & Duya, M. V. (2007) A new Luperosaurus (Squamata: Gekkonidae) from the Sierra Madre of Luzon Island, Philippines. Raffles Bulletin of Zoology, 55 (1), 167 - 174." type="journal article" year="2007">Brown, Diesmos</bibRefCitation>
|
||
</taxonomicName>
|
||
, & Duya, 2007 (distribution:
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5F5FF4FFF5EA3F83E9FCDB04D8FC32" httpUri="https://zenodo.org/record/274188/files/figure.png" pageId="6" pageNumber="59" targetBox="[292,1310,188,873]" targetPageId="6">
|
||
<paragraph id="8B9F0F7CFFF5EA3F83E9FCDB04D8FC32" blockId="6.[151,1436,893,948]" pageId="6" pageNumber="59">
|
||
<emphasis id="B954D36EFFF5EA3F83E9FCDB051CFC13" bold="true" box="[151,280,893,917]" pageId="6" pageNumber="59">FIGURE 5.</emphasis>
|
||
Holotype of
|
||
<taxonomicName id="4C2074FFFFF5EA3F82DBFCDB067CFC12" box="[421,632,893,916]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="59" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
|
||
<emphasis id="B954D36EFFF5EA3F82DBFCDB067CFC12" box="[421,632,893,916]" italics="true" pageId="6" pageNumber="59">Luperosaurus sorok</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954D36EFFF5EA3F8100FCDB06D7FC13" bold="true" box="[638,723,893,917]" pageId="6" pageNumber="59">
|
||
<taxonomicNameLabel id="A2676E15FFF5EA3F8100FCDB06D7FC13" box="[638,723,893,917]" pageId="6" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(SP 06618), showing scales of dorsum at midbody. Scale marker = 2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF5F5FF4FFF5EA3F83E9F81005E8F86B" httpUri="https://zenodo.org/record/274189/files/figure.png" pageId="6" pageNumber="59" targetBox="[398,1192,1000,1951]" targetPageId="6">
|
||
<paragraph id="8B9F0F7CFFF5EA3F83E9F81005E8F86B" blockId="6.[151,1436,1974,2029]" pageId="6" pageNumber="59">
|
||
<emphasis id="B954D36EFFF5EA3F83E9F810051FF848" bold="true" box="[151,283,1974,1998]" pageId="6" pageNumber="59">FIGURE 6.</emphasis>
|
||
Map of Borneo, showing the type locality (indicated with a white spot) of
|
||
<taxonomicName id="4C2074FFFFF5EA3F8732F8110124F848" box="[1100,1312,1975,1998]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="6" pageNumber="59" phylum="Chordata" rank="species" species="sorok" status="sp. nov.">
|
||
<emphasis id="B954D36EFFF5EA3F8732F8110124F848" box="[1100,1312,1975,1998]" italics="true" pageId="6" pageNumber="59">Luperosaurus sorok</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B954D36EFFF5EA3F8657F8100184F848" bold="true" box="[1321,1408,1974,1998]" pageId="6" pageNumber="59">
|
||
<taxonomicNameLabel id="A2676E15FFF5EA3F8657F8100184F848" box="[1321,1408,1974,1998]" pageId="6" pageNumber="59" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the Crocker Range Park, Sabah.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33A5CF7FFF4EA3E83E9FF3E049AFE41" pageId="7" pageNumber="60" type="description">
|
||
<paragraph id="8B9F0F7CFFF4EA3E83E9FF3E052BFE24" blockId="7.[151,1437,152,455]" pageId="7" pageNumber="60">
|
||
Luzon,
|
||
<collectingCountry id="F3374FECFFF4EA3E838CFF3E057FFF34" box="[242,379,152,178]" name="Philippines" pageId="7" pageNumber="60">Philippines</collectingCountry>
|
||
), a gigantic species (at
|
||
<quantity id="4CD8A299FFF4EA3E81EFFF3E070DFF34" box="[657,777,152,178]" metricMagnitude="-1" metricUnit="m" metricValue="1.054" pageId="7" pageNumber="60" unit="mm" value="105.4">105.4 mm</quantity>
|
||
SVL, is over thrice as large as the
|
||
<typeStatus id="549BB1DEFFF4EA3E87D1FF3E0111FF34" box="[1199,1301,152,178]" pageId="7" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
of the new species); a single intersupranasal contacts rostral; preanofemoral pores 16; subrictal tubercles absent; scansors on Toe I, 12; on Toe III, 16; ventrals granular to subimbricate; and tail lacks lateral tubercles;
|
||
<taxonomicName id="4C2074FFFFF4EA3E87ABFF4E0537FEAC" authority="Stejneger, 1907" authorityName="Stejneger" authorityYear="1907" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="macgregori">
|
||
<emphasis id="B954D36EFFF4EA3E87ABFF4E0159FE87" box="[1237,1373,232,257]" italics="true" pageId="7" pageNumber="60">macgregori</emphasis>
|
||
(Stejneger, 1907)
|
||
</taxonomicName>
|
||
(distribution: Babuyan and Calayan,
|
||
<collectingCountry id="F3374FECFFF4EA3E818CFEB6077FFEAC" box="[754,891,272,298]" name="Philippines" pageId="7" pageNumber="60">Philippines</collectingCountry>
|
||
), digital dilation extensive; genials small; and subrictal tubercles absent; and
|
||
<taxonomicName id="4C2074FFFFF4EA3E8170FE9E07E6FED4" authority="Brown & Alcala, 1978" authorityName="Brown & Alcala" authorityYear="1978" box="[526,994,312,338]" class="Reptilia" family="Gekkonidae" genus="Luperosaurus" kingdom="Animalia" order="Squamata" pageId="7" pageNumber="60" phylum="Chordata" rank="species" species="palawanensis">
|
||
<emphasis id="B954D36EFFF4EA3E8170FE9E06A9FED7" box="[526,685,312,337]" italics="true" pageId="7" pageNumber="60">palawanensis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFB1728DFFF4EA3E81BFFE9E07DEFED4" author="Brown" box="[705,986,312,338]" pageId="7" pageNumber="60" refString="Brown, W. C. & Alcala, A. C. (1978) Philippine lizards of the family Gekkonidae. Silliman University Natural Science Monograph Series No. 1, Dumaguete City, Philippines. v + 146 pp + (1) errata sheet." type="book" year="1978">Brown & Alcala, 1978</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
(distribution: Palawan,
|
||
<collectingCountry id="F3374FECFFF4EA3E8679FE9E0194FED4" box="[1287,1424,312,338]" name="Philippines" pageId="7" pageNumber="60">Philippines</collectingCountry>
|
||
), dorsum with spear-head-like scales; penultimate scansors bowed; supralabials 11–13; and preanofemoral pores 28–32.
|
||
</paragraph>
|
||
<paragraph id="8B9F0F7CFFF4EA3E83E9FE16049AFE41" blockId="7.[151,1437,152,455]" box="[151,158,432,455]" pageId="7" pageNumber="60">.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |