378 lines
38 KiB
XML
378 lines
38 KiB
XML
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<mods:title id="9AB4227E097D7356C993A23DAE5E6566">New and interesting spiders (Arachnida, Araneae) from Vashlovani National Park and Chachuna Managed Reserve (Georgia)</mods:title>
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<mods:namePart id="0C806EB2DE64E91E39A831B982D5B172">Seropian, Armen</mods:namePart>
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<mods:affiliation id="0A25CB086D451ACBBFB19B0A88976E3C">Institute of Ecology, Ilia State University, Cholokashvili av. 3 / 5 Tbilisi, 0162, Georgia</mods:affiliation>
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<mods:namePart id="68D3754343F62A250B4549C610F544A9">Bulbulashvili, Natalia</mods:namePart>
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<mods:nameIdentifier id="15851FE51B94930818BC9DEFFF7D970C" type="ORCID">0000-0002-6802-1209</mods:nameIdentifier>
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<mods:affiliation id="D08C9AFFC1DBFDDF7B0DBAF7C43B0269">Rustaveli st. 9, 1400, Gori, Georgia</mods:affiliation>
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<mods:namePart id="2FE7758CD161A5D2E08841D27F559C5B">Makharadze, Giorgi</mods:namePart>
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<mods:nameIdentifier id="7EC171F164F003088FE87949DA84B3B7" type="ORCID">https://orcid.org/0009-0008-4443-5432</mods:nameIdentifier>
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<mods:affiliation id="1F74B404523BD8F6E2ECFC7140C2E9BD">Konstantine Eristavi st. 12, 0103, Tbilisi, Georgia</mods:affiliation>
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<mods:namePart id="CBD8E7D5B3F20F231D628235CEDA99E2">Baznikin, Andrei</mods:namePart>
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<mods:nameIdentifier id="4F6B7355859D3C799F7FA2A2BCA985B6" type="ORCID">https://orcid.org/0009-0001-9149-1735</mods:nameIdentifier>
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<mods:affiliation id="B28C60A70EC25E0EA2D88FD65DF4A3EC">Pozheshka st. 95 / 1 / 11, Belgrade, Serbia</mods:affiliation>
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<mods:title id="036F915688A4AF8D062F4C612BEA4396">Caucasiana</mods:title>
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<mods:part id="A69B55D565394FF14FD12D790775B6B8">
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<mods:date id="B87F59F4701313498C3A8A9EB18CCF50">2024</mods:date>
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<mods:number id="5789E9C09A17561CD818D357F09217C5">2024-10-01</mods:number>
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<mods:number id="041570F3C1E20B7D3BD94CBA7C695905">3</mods:number>
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<mods:start id="F5EB264E9AD0DCEC304A7110C5DBFE5F">183</mods:start>
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<mods:classification id="89F62B386D4A7D8BCB2B0FC1C37B61E9">journal article</mods:classification>
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<treatment id="703E32346F8055F9BDCEC9D7AE7BC4A0" ID-DOI="http://doi.org/10.5281/zenodo.13870374" ID-Zenodo-Dep="13870374" ID-arpha="703E3234-6F80-55F9-BDCE-C9D7AE7BC4A0" LSID="urn:lsid:plazi:treatment:703E32346F8055F9BDCEC9D7AE7BC4A0" httpUri="http://treatment.plazi.org/id/703E32346F8055F9BDCEC9D7AE7BC4A0">
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<subSubSection id="9346535C0B5B931148ABE3E1B725DBF3" type="nomenclature">
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<paragraph id="6B1D6DE0E8158BB55217D00E1614E7EB">
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<heading id="A0E53058B99EF86FEE49F6AEF2798F77" reason="label">**</heading>
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<taxonomicName id="AA31B4932901EFD3A9282A0CFF537E16" ID-CoL="992FL" authority="Shafaie, Nadolny & Mirshamsi, 2022" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
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<emphasis id="8E9FD731128C3F2CBD4CC3C4B405FBCF" italics="true">Lycosa soboutii</emphasis>
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<bibRefCitation id="0DE44864A3BDE5B25F633854D1A42F3A" DOI="10.11646/zootaxa.5120.4.3" author="Shafaie S & Nadolny AA & Mirshamsi O" issue="4" journalOrPublisher="Zootaxa" pagination="501-522" refId="B39" refString="Shafaie S, Nadolny AA, Mirshamsi O (2022) A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae). Zootaxa 5120 (4): 501 - 522. https: // doi. org / 10.11646 / zootaxa. 5120.4. 3" title="A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae)." volume="5120" year="2022">Shafaie, Nadolny & Mirshamsi, 2022</bibRefCitation>
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</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="7067229C7033949CEFA29553035E02D2" type="description">
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<paragraph id="4B5DF5552839CC39AB257658AEC8BCEF">
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<figureCitation id="9A0282DD4C73829A4B5688B51D5C9168" captionStart="Figures 19–20" captionStartId="F7" captionText="Figures 19–20. Lycosa soboutii, live specimens (19: female from Osiaskheoba, dorsal view; 20: male from Osiaskheoba, dorsal view)." figureDoi="10.3897/caucasiana.3.e132501.figures19-20" httpUri="https://binary.pensoft.net/fig/1146196">Figs 19</figureCitation>
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<figureCitation id="49040DE9B146B1A6D6E72A56D115F846" captionStart="Figures 21–24" captionStartId="F8" captionText="Figures 21–24. Lycosa soboutii (21: male, left palp, ventral view; 22: ditto, retrolateral view; 23: female, epigyne, ventral view; 24: endogyne, dorsal). Abbreviations: Em – embolus, Se – synembolus, Ta – tegular apophysis, Tg – tegulum, To – ventral outgrowth of tegular apophysis, Tp – distal process of tegular apophysis, Tr – tegular ridge. Scale bars: 0.5 mm." figureDoi="10.3897/caucasiana.3.e132501.figures21-24" httpUri="https://binary.pensoft.net/fig/1146197">–24</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="F900FC567940A3DF2C258473075F47F2" type="reference_group">
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<paragraph id="F491FA218EEEAD8470E80187B4242F73">
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<treatmentCitationGroup id="52C80F656182E904B2DE720FB1D54F72">
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<treatmentCitation id="D1F3462FB73101E5164443AFFEEE7994">
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<taxonomicName id="9E580AA037144209409D7FB2BDFABC7F" ID-CoL="992FL" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
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<emphasis id="F5FD9AD599A09BD690B9CAA9602DA0FF" italics="true">Lycosa soboutii</emphasis>
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</taxonomicName>
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||
<comment id="B74530F8453213F17C6583A79BBEE490">
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:
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<treatmentCitation id="0FE2CD347D01EBF1DB22CBDA9550F858" ID-DOI="http://doi.org/10.5281/zenodo.6392784" ID-Zenodo-Dep="6392784" LSID="urn:lsid:zoobank.org:act:EF28289D-EA16-4003-9056-1B52032E83BF" author="Shafaie S & Nadolny AA & Mirshamsi O" httpUri="http://treatment.plazi.org/id/CF7F87B8FF9BFFE0FF2152A1FC83A1E3" page="502" year="2022">
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<bibRefCitation id="A2F28D26F29BD37C669E2FD2CB5CC836" DOI="10.11646/zootaxa.5120.4.3" author="Shafaie S & Nadolny AA & Mirshamsi O" etAl="et al." firstAuthor="Shafaie" issue="4" journalOrPublisher="Zootaxa" page="502" pagination="501-522" refId="B39" refString="Shafaie S, Nadolny AA, Mirshamsi O (2022) A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae). Zootaxa 5120 (4): 501–522. https://doi.org/10.11646/zootaxa.5120.4.3" title="A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae)." volume="5120" year="2022">Shafaie et al. 2022: 502</bibRefCitation>
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</treatmentCitation>
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||
, figs 1–7 (
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<specimenCount id="0B1BE0A5BD11493B1A36A686E61EC84E" count="1" type="female">♀</specimenCount>
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).
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</comment>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="41301A28C653D214F2D34F2531E21274">
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<treatmentCitationGroup id="BB3CC0623C2A65D354BE2EA041462279">
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<treatmentCitation id="EE336609D06D0E4D8C90FE7723A73615">
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<taxonomicName id="AD27A578E4986EC736B9B53A1F8F6ACC" ID-CoL="992FL" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
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||
<emphasis id="806B2B3E2B9AAF56AB96B8F95AC4D35F" italics="true">Lycosa soboutii</emphasis>
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</taxonomicName>
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||
<comment id="D0A65BD7CAAF0AC078D5A8E2726EACD9">
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||
:
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||
<treatmentCitation id="7FD25D284457C4796E2BD9CF92973E08" author="Logunov DV" page="505" year="2023">
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||
<bibRefCitation id="17C859F7D122CA3299E4B5D233C81FB8" DOI="10.15298/arthsel.32.4.12" author="Logunov DV" firstAuthor="Logunov" issue="4" journalOrPublisher="Arthropoda Selecta" page="505" pagination="475-512" refId="B14" refString="Logunov DV (2023) Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475–512. https://doi.org/10.15298/arthsel.32.4.12" title="Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae)." volume="32" year="2023">Logunov 2023: 505</bibRefCitation>
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||
</treatmentCitation>
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||
, figs 145–148 (
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<specimenCount id="6633B881E16B0560F5E322D6AE928C12" count="1" type="female">♀</specimenCount>
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||
).
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</comment>
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</treatmentCitation>
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="SECID0E4FBG" type="material">
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<paragraph id="E77D048C460C30A8E971AF99270106D1">
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<heading id="730E19D8397D6E7AC8247EC0B67E97C0" reason="title">Material examined.</heading>
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</paragraph>
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<paragraph id="A259540D581DC5973CC2ADB4A78B0929">
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||
<materialsCitation id="1B10077D76643161F9740A7F8B29F4E6" collectingDate="2024-05-19" collectorName="Chachuna MR & Makharadze G & Seropian A" country="Georgia" latitude="41.2314" location="Osiaskheoba" longLatPrecision="6" longitude="46.1473" municipality="Dedoplistskaro" specimenCount="7" specimenCount-female="4" specimenCount-male="3" stateProvince="Kakheti">
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<collectingCountry id="3F3E047767AAC76E4FD29D23E184FDF2" name="Georgia">GEORGIA</collectingCountry>
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•
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<specimenCount id="40DADD1BC82389B6D82D03D7E2C981F1" count="3" type="male">3 ♂♂</specimenCount>
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,
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<specimenCount id="6A7CDBC0A23D21D760AACDF1F4F7C0A0" count="4" type="female">4 ♀♀</specimenCount>
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;
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<collectingRegion id="95D9823A83C0D11CE52CE81D99AC411F" country="Georgia" name="Kakheti">Kakheti</collectingRegion>
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,
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<collectingMunicipality id="24726B2296ED86857A98653871656FB6">Dedoplistskaro</collectingMunicipality>
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<abbrev id="ABBRID0EDGBG" xlink_title="municipality">mun</abbrev>
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.,
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<collectorName id="D3F3CEE9BEC2BAC1BB6E30E80359C702">Chachuna MR</collectorName>
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,
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<location id="07192ABF2726B1EC812C5DEA279B58A0" LSID="urn:lsid:plazi:treatment:703E32346F8055F9BDCEC9D7AE7BC4A0:07192ABF2726B1EC812C5DEA279B58A0" country="Georgia" latitude="41.2314" longLatPrecision="6" longitude="46.1473" municipality="Dedoplistskaro" name="Osiaskheoba" stateProvince="Kakheti">Osiaskheoba</location>
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;
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0EKGBG" specific-use="{"type":"Point","coordinates":[46.147300,41.231400]}">
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<geoCoordinate id="F81C9E0BD82B1478469054DFDD1071FB" degrees="41.2314" direction="north" orientation="latitude" precision="5" value="41.2314">N 41.2314 °</geoCoordinate>
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,
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<geoCoordinate id="D9153E29C709719DB85DA09AC5928582" degrees="46.1473" direction="east" orientation="longitude" precision="5" value="46.1473">E 46.1473 °</geoCoordinate>
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</named-content>
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||
; semidesert, at night with flashlight; leg.
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||
<collectorName id="1465990BF0DADEEADBBD46A239CACBFB">Makharadze G</collectorName>
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,
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<collectorName id="93B155A8153D11674300BA797BB08600">Seropian A</collectorName>
|
||
;
|
||
<date id="7BD2234A4F975DE51D1BAA349B2E02DC" value="2024-05-19">
|
||
<collectingDate id="477F107BA6961C3EFECE2CA7EBB99BBD" value="2024-05-19">19 May 2024</collectingDate>
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</date>
|
||
; CaBOL-IDs 1038296, 1038298, 1038073, 1038297, 1038299, 1038074, 1038075
|
||
</materialsCitation>
|
||
. •
|
||
<materialsCitation id="9AA29BFE95E8F5145329D2462732AE43" collectingDate="2024-05-20" collectorName="Vashlovani NP & Bulbulashvili N" country="Georgia" latitude="41.2382" location="Datviskhevi" longLatPrecision="6" longitude="46.3637" specimenCount="1" specimenCount-female="1" stateProvince="Kakheti">
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<specimenCount id="9EECD53CC275264D4325ABED79C0C043" count="1" type="female">1 ♀</specimenCount>
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;
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<collectingRegion id="FA28D03EE235F1AFE281CDBA5C755C24" country="Georgia" name="Kakheti">Kakheti</collectingRegion>
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,
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<collectorName id="6F383A675C0AF34AFF0E26C322916175">Vashlovani NP</collectorName>
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,
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<location id="7B64BE50B4388668A6761513CC6FBD26" LSID="urn:lsid:plazi:treatment:703E32346F8055F9BDCEC9D7AE7BC4A0:7B64BE50B4388668A6761513CC6FBD26" country="Georgia" latitude="41.2382" longLatPrecision="6" longitude="46.3637" name="Datviskhevi" stateProvince="Kakheti">Datviskhevi</location>
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;
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<named-content content-type="dwc:verbatimCoordinates" id="NCID0ESGBG" specific-use="{"type":"Point","coordinates":[46.363700,41.238200]}">
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<geoCoordinate id="45D99A12718B009CC49253E062DEA692" degrees="41.2382" direction="north" orientation="latitude" precision="5" value="41.2382">N 41.2382 °</geoCoordinate>
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,
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<geoCoordinate id="422F5A080B91C7A0B85ED9B06DCBCA70" degrees="46.3637" direction="east" orientation="longitude" precision="5" value="46.3637">E 46.3637 °</geoCoordinate>
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</named-content>
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; semidesert, lured from burrow; leg.
|
||
<collectorName id="5C00F09E7A8B38FF598A1E4B9C92244B">Bulbulashvili N</collectorName>
|
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;
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<date id="899A8FF13E6A09029AC30D4E2C0EA0CF" value="2024-05-20">
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<collectingDate id="FC90A1FF72596E3EFA039CDCF5A11D74" value="2024-05-20">20 May 2024</collectingDate>
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</date>
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; CaBOL-ID 1038300
|
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</materialsCitation>
|
||
. •
|
||
<materialsCitation id="D452F8A52859EBCD97465608F2AA41C4" collectingDate="2024-05-22" collectorName="Makharadze G & The & Ilia State University & Georgia" country="Georgia" latitude="41.1245" location="Mijniskure" longLatPrecision="6" longitude="46.6456" specimenCount="1" specimenCount-female="1" stateProvince="Kakheti">
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<specimenCount id="6914D3A97B5C7BCCD736A5D06C1BAAA5" count="1" type="female">1 ♀</specimenCount>
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||
;
|
||
<collectingRegion id="D357D0D8491D01C8748F6D50CD52CA43" country="Georgia" name="Kakheti">Kakheti</collectingRegion>
|
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,
|
||
<location id="E89C3B179AF1E8008B325DFC42290272" LSID="urn:lsid:plazi:treatment:703E32346F8055F9BDCEC9D7AE7BC4A0:E89C3B179AF1E8008B325DFC42290272" country="Georgia" latitude="41.1245" longLatPrecision="6" longitude="46.6456" name="Mijniskure" stateProvince="Kakheti">Mijniskure</location>
|
||
;
|
||
<named-content content-type="dwc:verbatimCoordinates" id="NCID0E1GBG" specific-use="{"type":"Point","coordinates":[46.645600,41.124500]}">
|
||
<geoCoordinate id="A2CA706AF11815C05556DCF8D5B99023" degrees="41.1245" direction="north" orientation="latitude" precision="5" value="41.1245">N 41.1245 °</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="CAC7B5B81052CEEE47271ED982002A2B" degrees="46.6456" direction="east" orientation="longitude" precision="5" value="46.6456">E 46.6456 °</geoCoordinate>
|
||
</named-content>
|
||
; semidesert, at night with flashlight; leg.
|
||
<collectorName id="465643AF0461B3DB6AA4790EE9ED9676">Makharadze G</collectorName>
|
||
;
|
||
<date id="68346F64368F5D7C0EF8B8BFDB10CDBC" value="2024-05-22">
|
||
<collectingDate id="9FDC2B359D6952938F6A64584BE1851F" value="2024-05-22">22 May 2024</collectingDate>
|
||
</date>
|
||
; CaBOL-ID 1038062.
|
||
<collectorName id="C7BE59E18ABB9FA68AC32A03A09E3ED6">The</collectorName>
|
||
material is deposited in the scientific collections of
|
||
<collectorName id="CF02B6497A2710F4D71892C8CE373BDC">Ilia State University</collectorName>
|
||
,
|
||
<collectorName id="91F06366FAC05113D0B3DB34BD91DA5D">Georgia</collectorName>
|
||
,
|
||
<collectingRegion id="557F70A5D50E8CC555BCC21F2AF31325" country="Georgia" name="Tbilisi">Tbilisi</collectingRegion>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="61351279D0E95A818DB7A8B09FD4D5DD">
|
||
<emphasis id="35176B54019E10CD5F4E99D58EA1D08A" bold="true">Diagnosis</emphasis>
|
||
. In having extra processes of the tegular apophysis, the males of
|
||
<taxonomicName id="7266D5BDE3CB977C09B318D1E21E3D62" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
|
||
<emphasis id="2F0D05729271497616F34FB2AE90B460" italics="true">L. soboutii</emphasis>
|
||
</taxonomicName>
|
||
are most similar to those of
|
||
<taxonomicName id="AE80C01F1B591921EE770FF52B901EF6" authority="Zamani et Nadolny, 2022" authorityName="Zamani et Nadolny" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="elymaisa">
|
||
<emphasis id="05CEE3E5B578593275165026B4C64217" italics="true">L. elymaisa</emphasis>
|
||
<bibRefCitation id="DF88D75B3DF3E5271758301D4BF2BA3E" DOI="10.13156/arac.2022.19.2.551" author="Zamani A & Nadolny AA & Dolejš P" issue="2" journalOrPublisher="Arachnology" pagination="551-573" refId="B47" refString="Zamani A, Nadolny AA, Dolejs P (2022) New data on the spider fauna of Iran (Arachnida: Araneae), part X. Arachnology 19 (2): 551 - 573. https: // doi. org / 10.13156 / arac. 2022.19. 2.551" title="New data on the spider fauna of Iran (Arachnida: Araneae), part X." volume="19" year="2022">Zamani et Nadolny, 2022</bibRefCitation>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="236B765FF9B7DC6309074E7D025FA80A" authority="Nadolny et Zamani, 2017" authorityName="Nadolny et Zamani" authorityYear="2017" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="aragogi">
|
||
<emphasis id="E0E7F4A59672D313267883244B687DF1" italics="true">L. aragogi</emphasis>
|
||
Nadolny et Zamani, 2017
|
||
</taxonomicName>
|
||
from
|
||
<collectingCountry id="6BAE650565182C3AC4DF2F7BF7669FC4" name="Iran">Iran</collectingCountry>
|
||
, from which it can be easily distinguished by the less curved tegulum (Tg) [vs strongly curved in
|
||
<taxonomicName id="337C72FB5E4DE328B855FEF81B0E98CD" authorityName="Zamani et Nadolny" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="elymaisa">
|
||
<emphasis id="2B4B39075DDC2A4DA16268C2FDE0CECB" italics="true">L. elymaisa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="7DAF914C7EC5D651A1A78D758D9C01E8" DOI="10.13156/arac.2022.19.2.551" author="Zamani" etAl="et al." firstAuthor="Zamani" issue="2" journalOrPublisher="Arachnology" pagination="551-573" refId="B47" refString="Zamani A, Nadolny AA, Dolejš P (2022) New data on the spider fauna of Iran (Arachnida: Araneae), part X. Arachnology 19 (2): 551–573. https://doi.org/10.13156/arac.2022.19.2.551" title="New data on the spider fauna of Iran (Arachnida: Araneae), part X." volume="19" year="2022">Zamani et al. 2022</bibRefCitation>
|
||
: figs 3 E, 4 A – B) and
|
||
<taxonomicName id="7C073B23B1553FE2F18197133F15CC62" authorityName="Nadolny et Zamani" authorityYear="2017" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="aragogi">
|
||
<emphasis id="14EEA1DBCFE8685B1DA16D809E3E6971" italics="true">L. aragogi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="F53A9FC809132228E97B945EAE2F0A6D" DOI="10.15298/arthsel.32.4.12" author="Logunov" firstAuthor="Logunov" issue="4" journalOrPublisher="Arthropoda Selecta" pagination="475-512" refId="B14" refString="Logunov DV (2023) Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475–512. https://doi.org/10.15298/arthsel.32.4.12" title="Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae)." volume="32" year="2023">Logunov 2023</bibRefCitation>
|
||
: figs 129, 131)], strongly pronounced sclerotized tegular ridge (Tr), more closely located embolus (Em) and synembolus (Se) [vs more diverged in
|
||
<taxonomicName id="8C5FC242A62C83ED8211591A176D4D61" authorityName="Zamani et Nadolny" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="elymaisa">
|
||
<emphasis id="9225AE8D80F7105946D91351D2BE16F1" italics="true">L. elymaisa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="4D196664850630688051A86FDA8DB5AD" DOI="10.13156/arac.2022.19.2.551" author="Zamani" etAl="et al." firstAuthor="Zamani" issue="2" journalOrPublisher="Arachnology" pagination="551-573" refId="B47" refString="Zamani A, Nadolny AA, Dolejš P (2022) New data on the spider fauna of Iran (Arachnida: Araneae), part X. Arachnology 19 (2): 551–573. https://doi.org/10.13156/arac.2022.19.2.551" title="New data on the spider fauna of Iran (Arachnida: Araneae), part X." volume="19" year="2022">Zamani et al. 2022</bibRefCitation>
|
||
: figs 3 H – E, 4 A – B) and
|
||
<taxonomicName id="CB338199B68EAAE6C76DC406CCEEA00C" authorityName="Nadolny et Zamani" authorityYear="2017" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="aragogi">
|
||
<emphasis id="B294FC0EF9DCAB57051FEAE1CAFDE6D5" italics="true">L. aragogi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="8336528EDBDC87FEBDBA60EAA631AE84" DOI="10.15298/arthsel.32.4.12" author="Logunov" firstAuthor="Logunov" issue="4" journalOrPublisher="Arthropoda Selecta" pagination="475-512" refId="B14" refString="Logunov DV (2023) Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475–512. https://doi.org/10.15298/arthsel.32.4.12" title="Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae)." volume="32" year="2023">Logunov 2023</bibRefCitation>
|
||
: fig. 135)], proximally located ventral outgrowth of tegular apophysis (To) [vs distally located in
|
||
<taxonomicName id="8324E1CB5FD69225EF123204C2FF4B9D" authorityName="Zamani et Nadolny" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="elymaisa">
|
||
<emphasis id="87F6193E1B415AD5544706A6D6B53030" italics="true">L. elymaisa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="B820D927F70A10EC2C39210AD88EEBB9" DOI="10.13156/arac.2022.19.2.551" author="Zamani" etAl="et al." firstAuthor="Zamani" issue="2" journalOrPublisher="Arachnology" pagination="551-573" refId="B47" refString="Zamani A, Nadolny AA, Dolejš P (2022) New data on the spider fauna of Iran (Arachnida: Araneae), part X. Arachnology 19 (2): 551–573. https://doi.org/10.13156/arac.2022.19.2.551" title="New data on the spider fauna of Iran (Arachnida: Araneae), part X." volume="19" year="2022">Zamani et al. 2022</bibRefCitation>
|
||
: figs 3 G – H, 4 A, C) and
|
||
<taxonomicName id="F01504F1FC102F6632BCEAE6325F128D" authorityName="Nadolny et Zamani" authorityYear="2017" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="aragogi">
|
||
<emphasis id="3A6B51E932F0D5291FCEE6B9AFB663FD" italics="true">L. aragogi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="E975AD388435030EE73D13061B4D7AA0" DOI="10.15298/arthsel.32.4.12" author="Logunov" firstAuthor="Logunov" issue="4" journalOrPublisher="Arthropoda Selecta" pagination="475-512" refId="B14" refString="Logunov DV (2023) Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475–512. https://doi.org/10.15298/arthsel.32.4.12" title="Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae)." volume="32" year="2023">Logunov 2023</bibRefCitation>
|
||
: figs 129–132)], and bifid distal process of tegular apophysis (Tp) directed ventro-posteriorly [vs spiral-shaped in
|
||
<taxonomicName id="38CA55A478DC33208BC18BDDC32B5736" authorityName="Zamani et Nadolny" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="elymaisa">
|
||
<emphasis id="F0365D7162CF3031BB1BDBD3C007B948" italics="true">L. elymaisa</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="DFF5C34D8B2346C6EC5F1B96F1D8CC6A" DOI="10.13156/arac.2022.19.2.551" author="Zamani" etAl="et al." firstAuthor="Zamani" issue="2" journalOrPublisher="Arachnology" pagination="551-573" refId="B47" refString="Zamani A, Nadolny AA, Dolejš P (2022) New data on the spider fauna of Iran (Arachnida: Araneae), part X. Arachnology 19 (2): 551–573. https://doi.org/10.13156/arac.2022.19.2.551" title="New data on the spider fauna of Iran (Arachnida: Araneae), part X." volume="19" year="2022">Zamani et al. 2022</bibRefCitation>
|
||
: figs 3 G – I, 4 A – D) and claw-shaped apically directed in
|
||
<taxonomicName id="D4302F1F98B47545CAF17C11BBCCE7E1" authorityName="Nadolny et Zamani" authorityYear="2017" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="aragogi">
|
||
<emphasis id="5341FAAC798569248B1B288FC5057DE3" italics="true">L. aragogi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation id="DA43035649FDF556DCEC7466A136D1E8" DOI="10.15298/arthsel.32.4.12" author="Logunov" firstAuthor="Logunov" issue="4" journalOrPublisher="Arthropoda Selecta" pagination="475-512" refId="B14" refString="Logunov DV (2023) Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae). Arthropoda Selecta 32 (4): 475–512. https://doi.org/10.15298/arthsel.32.4.12" title="Further notes on the fossorial wolf spiders of Middle Asia and the Near East (Aranei: Lycosidae)." volume="32" year="2023">Logunov 2023</bibRefCitation>
|
||
: figs 129–132)].
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0E3LBG" type="description">
|
||
<paragraph id="E6ADDBE2B7EA2643D8C2FF8D85113F99">
|
||
<heading id="D3B2D186C87E9D9CC400EB81902F9608" reason="title">Description.</heading>
|
||
</paragraph>
|
||
<paragraph id="52B029BCE8B39631A1E5699FD6112A80">
|
||
Male. Total length 13.87. Carapace 7.49 long, 5.41 wide. Carapace light reddish brown, covered with dense off-white setae and medially radiating darker bands (Fig.
|
||
<figureCitation id="31678480CFE5287132578C617B59DDEE" captionStart="Figures 19–20" captionStartId="F7" captionText="Figures 19–20. Lycosa soboutii, live specimens (19: female from Osiaskheoba, dorsal view; 20: male from Osiaskheoba, dorsal view)." figureDoi="10.3897/caucasiana.3.e132501.figures19-20" httpUri="https://binary.pensoft.net/fig/1146196">20</figureCitation>
|
||
); pars thoracica 1.6 times wider than pars cephalica; pars cephalica elevated. Chelicerae brown, with 3 pro- and 3 retromarginal teeth, with dense light-yellow setae, covering ca. ½ of cheliceral length. Labium and maxillae light-brown. Sternum black, covered with black and yellow setae. Abdomen yellow, dorsally with light trident-shaped mark margined with black setae. Spinnerets yellowish, apically darkened. All legs covered with dense off-white setae (Fig.
|
||
<figureCitation id="20BE31F3702B2A28B16D4D0EBBB1DB13" captionStart="Figures 19–20" captionStartId="F7" captionText="Figures 19–20. Lycosa soboutii, live specimens (19: female from Osiaskheoba, dorsal view; 20: male from Osiaskheoba, dorsal view)." figureDoi="10.3897/caucasiana.3.e132501.figures19-20" httpUri="https://binary.pensoft.net/fig/1146196">20</figureCitation>
|
||
): Tr,
|
||
<abbrev id="ABBRID0EKMBG" xlink_title="Tibia">Ti</abbrev>
|
||
,
|
||
<abbrev id="ABBRID0EOMBG" xlink_title="Patella">Pa</abbrev>
|
||
light-yellow; Tr ventrally with black setae;
|
||
<abbrev id="ABBRID0ESMBG" xlink_title="Tarsus">Ta</abbrev>
|
||
and
|
||
<abbrev id="ABBRID0EWMBG" xlink_title="Metatarsus">Mt</abbrev>
|
||
I – II brown,
|
||
<abbrev id="ABBRID0E1MBG" xlink_title="Tarsus">Ta</abbrev>
|
||
and
|
||
<abbrev id="ABBRID0E5MBG" xlink_title="Metatarsus">Mt</abbrev>
|
||
III – IV yellow; venter of
|
||
<abbrev id="ABBRID0ECNBG" xlink_title="Patella">Pa</abbrev>
|
||
I – IV apically, and
|
||
<abbrev id="ABBRID0EGNBG" xlink_title="Tibia">Ti</abbrev>
|
||
I – IV apically and basally darkened. Leg measurements: I 21.97 (6.05 + 2.58 + 5.47 + 5.41 + 2.46), II 20.04 (5.69 + 2.35 + 4.79 + 4.59 + 2.62), III 19.58 (5.57 + 2.14 + 3.57 + 5.52 + 2.78), IV 25.92 (7.21 + 2.34 + 5.25 + 7.87 + 3.25). Palps yellowish, with brown bulb, covered with off-white setae. Palpal structure as in Figs
|
||
<figureCitation id="D63918E2A5056738F8F9BD5AEB64A3D7" captionStart="Figures 21–24" captionStartId="F8" captionText="Figures 21–24. Lycosa soboutii (21: male, left palp, ventral view; 22: ditto, retrolateral view; 23: female, epigyne, ventral view; 24: endogyne, dorsal). Abbreviations: Em – embolus, Se – synembolus, Ta – tegular apophysis, Tg – tegulum, To – ventral outgrowth of tegular apophysis, Tp – distal process of tegular apophysis, Tr – tegular ridge. Scale bars: 0.5 mm." figureDoi="10.3897/caucasiana.3.e132501.figures21-24" httpUri="https://binary.pensoft.net/fig/1146197">21–22</figureCitation>
|
||
: cymbium 1.5 times longer than wide, apex with 8–9 strong bristles; bulb slightly (≈ 1.1 times) wider than long; embolic division bares thin, long, and curved embolus, with well-developed pars pendula and embolus-like thin synembolus (Se).
|
||
</paragraph>
|
||
<paragraph id="44CD65AA7070F401CAB72322C0E51DE2">
|
||
Female. See
|
||
<bibRefCitation id="A62A7AF4065D764EBEC12274370F3CC2" DOI="10.11646/zootaxa.5120.4.3" author="Shafaie S & Nadolny AA & Mirshamsi O" etAl="et al." firstAuthor="Shafaie" issue="4" journalOrPublisher="Zootaxa" pagination="501-522" refId="B39" refString="Shafaie S, Nadolny AA, Mirshamsi O (2022) A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae). Zootaxa 5120 (4): 501–522. https://doi.org/10.11646/zootaxa.5120.4.3" title="A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae)." volume="5120" year="2022">Shafaie et al. (2022)</bibRefCitation>
|
||
. The shape of the epigyne and the endogyne of the females from
|
||
<collectingCountry id="DF209AB96677B5C7E8AF3582AC0CDBF8" name="Georgia">Georgia</collectingCountry>
|
||
(Figs
|
||
<figureCitation id="E9659F82438C4A2EE842C891BE6DCFB1" captionStart="Figures 21–24" captionStartId="F8" captionText="Figures 21–24. Lycosa soboutii (21: male, left palp, ventral view; 22: ditto, retrolateral view; 23: female, epigyne, ventral view; 24: endogyne, dorsal). Abbreviations: Em – embolus, Se – synembolus, Ta – tegular apophysis, Tg – tegulum, To – ventral outgrowth of tegular apophysis, Tp – distal process of tegular apophysis, Tr – tegular ridge. Scale bars: 0.5 mm." figureDoi="10.3897/caucasiana.3.e132501.figures21-24" httpUri="https://binary.pensoft.net/fig/1146197">23–24</figureCitation>
|
||
) leave no doubt in its identification as
|
||
<taxonomicName id="E90CD2CBBE76DB00497A3E78F976CECB" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
|
||
<emphasis id="15AE357A370FD8D6412A8FE8D8FB9CDD" italics="true">L. soboutii</emphasis>
|
||
</taxonomicName>
|
||
, recently described from
|
||
<collectingRegion id="A0D568B57BBC262C83E2DC08CC3164E9" country="Iran" name="Zanjan">Zanjan Province</collectingRegion>
|
||
in
|
||
<collectingCountry id="93844578222F8B70E39E61D1A16E361D" name="Iran">Iran</collectingCountry>
|
||
. Interestingly, the black spot entirely covering the venter of the
|
||
<typeStatus id="218AC5DFF5140177AAFE4A741B42951E" type="holotype">holotype</typeStatus>
|
||
female’s abdomen (
|
||
<bibRefCitation id="E8A6CED27B153C2C75DEBADEE06D151C" DOI="10.11646/zootaxa.5120.4.3" author="Shafaie" etAl="et al." firstAuthor="Shafaie" issue="4" journalOrPublisher="Zootaxa" pagination="501-522" refId="B39" refString="Shafaie S, Nadolny AA, Mirshamsi O (2022) A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae). Zootaxa 5120 (4): 501–522. https://doi.org/10.11646/zootaxa.5120.4.3" title="A new species of Lycosa and three new species and a new record of Karakumosa from Iran (Araneae, Lycosidae)." volume="5120" year="2022">Shafaie et al. 2022</bibRefCitation>
|
||
, fig. 2) varied in our specimens, sometimes extending only to half of the surface.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="SECID0EHOBG" type="distribution">
|
||
<paragraph id="BFF075C78A8F73F16C6D2F5EBCF47745">
|
||
<heading id="B57F6567B4C56AFD2CBAFD66A9EEE201" reason="title">Distribution.</heading>
|
||
</paragraph>
|
||
<paragraph id="CB23A5AB22BBEB03120FCF14523227BE">
|
||
<collectingCountry id="CB12781724AFDA5F003FF6A8B53BF2F2" name="Iran">Iran</collectingCountry>
|
||
(
|
||
<bibRefCitation id="2A12D7564C589F16719770D10B653118" DOI="10.7892/BORIS.105054" refId="B41" refString="World Spider Catalog (2024) World Spider Catalog. Version 23.5. Natural History Museum Bern, online at http://wsc.nmbe.ch (Accessed: 25 / 06 // 2024) https://doi.org/10.7892/BORIS.105054" url="http://wsc.nmbe.ch" year="2024">WSC 2024</bibRefCitation>
|
||
),
|
||
<collectingCountry id="F104FC9FB5344E00E40CE9AD852A984F" name="Georgia">Georgia</collectingCountry>
|
||
(present study). This is the first record from the Caucasus region.
|
||
</paragraph>
|
||
<paragraph id="B97B255934F4BABA4C2FF1BCFEA71252">
|
||
<emphasis id="00448678935005240C5918B31FA90744" bold="true">Comments</emphasis>
|
||
: At the time of capture, all specimens were in the subadult stages and then captive-reared during May.
|
||
</paragraph>
|
||
<caption id="6687D8CA9189143CE32E7A0FBC84B18B" ID-DOI="10.3897/caucasiana.3.e132501.figures19-20" ID-arpha="6C05381A-7B04-5588-9E59-02A6E8096DE4" httpUri="https://binary.pensoft.net/fig/1146196" startId="F7">
|
||
<paragraph id="4C4348DF81B2F0726F7B2CA5F5F4459C">
|
||
<label id="31B97A50903CDF19129C7B62ADF62A2E">Figures 19–20.</label>
|
||
</paragraph>
|
||
<paragraph id="C5BAC4B1C09FFAD5F96E8114BF894527">
|
||
<taxonomicName id="3FA2D9A9617AF519BDA6D16E9F5C69F8" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
|
||
<emphasis id="FBE9D075166469C766311D6383792A6C" italics="true">Lycosa soboutii</emphasis>
|
||
</taxonomicName>
|
||
, live specimens (
|
||
<emphasis id="C5C180D91F9A52F8635C01E2978F8828" bold="true">19</emphasis>
|
||
: female from Osiaskheoba, dorsal view;
|
||
<emphasis id="84C1764E483096E14D0B99FB401D8B89" bold="true">20</emphasis>
|
||
: male from Osiaskheoba, dorsal view).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="38249BA26975B6B0861AF6A590A17E69" ID-DOI="10.3897/caucasiana.3.e132501.figures21-24" ID-arpha="0CAC575A-82EF-5687-8C8F-F3DC549872E5" httpUri="https://binary.pensoft.net/fig/1146197" startId="F8">
|
||
<paragraph id="EEAB16F9B5F64C6F0B16919663446F39">
|
||
<label id="B1B0559A131669AFBE070022943A7551">Figures 21–24.</label>
|
||
</paragraph>
|
||
<paragraph id="0289FA5F497FAEEEA80DD3E7DAB4A689">
|
||
<taxonomicName id="C106EEAE7B08F655DAE80A58103DFEB6" authorityName="Shafaie, Nadolny & Mirshamsi" authorityYear="2022" class="Arachnida" family="Lycosidae" genus="Lycosa" kingdom="Animalia" order="Araneae" phylum="Arthropoda" rank="species" species="soboutii">
|
||
<emphasis id="1C8621A19877D1B934EA2EF409578D70" italics="true">Lycosa soboutii</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<emphasis id="AC5A1A748EB4513C862D98E8BC53AC1B" bold="true">21</emphasis>
|
||
: male, left palp, ventral view;
|
||
<emphasis id="EBE45EEE434971DAD058714D544F4367" bold="true">22</emphasis>
|
||
: ditto, retrolateral view;
|
||
<emphasis id="0D3186E4E4D4C0EEE4D7ECE330BE43EA" bold="true">23</emphasis>
|
||
: female, epigyne, ventral view;
|
||
<emphasis id="ACD59BCDD63510405321FBD29BBF1689" bold="true">24</emphasis>
|
||
: endogyne, dorsal). Abbreviations: Em – embolus, Se – synembolus,
|
||
<abbrev id="ABBRID0EDUAK" xlink_title="Tarsus">Ta</abbrev>
|
||
– tegular apophysis, Tg – tegulum, To – ventral outgrowth of tegular apophysis, Tp – distal process of tegular apophysis, Tr – tegular ridge. Scale bars: 0.5 mm.
|
||
</paragraph>
|
||
</caption>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |