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<document id="971339211DF58F067EF10C705DA4031B" ID="10.11646/zootaxa.4236.3.4SLASH10287" ID-CLB-Dataset="36453" ID-DOI="10.11646/zootaxa.4236.3.4" ID-GBIF-Dataset="d05219de-97e3-436c-a2c0-00c63b3ce636" ID-ISSN="1175-5326" ID-Zenodo-Dep="322266" ID-ZooBank="C034B77B-BCC0-46FA-9541-12229A053852" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1487926401001" checkinUser="plazi" docAuthor="De Souza, Wagner Rafael M. &amp; Santos, Allan Paulo Moreira" docDate="2017" docId="4B1B3448FFA4AC49E7E803EDFA374C33" docLanguage="en" docName="zootaxa.4236.3.4SLASH10287.pdf" docOrigin="Zootaxa 4236 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Oxyethira (Dactylotrichia) retrosa Souza &amp; Santos, 2017, sp. nov." docType="treatment" docVersion="17" lastPageNumber="490" masterDocId="B7224C30FFA6AC4FE77F0012FFAA4D5C" masterDocTitle="Taxonomic study of the genus Oxyethira Eaton 1873 (Trichoptera: Hydroptilidae) from Northeast Brazil: Eleven new species and distributional records" masterLastPageNumber="506" masterPageNumber="484" pageNumber="486" updateTime="1698416856431" updateUser="plazi">
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<mods:title id="7341BA5A991B0BF755E1C85E0EC2AB84">Taxonomic study of the genus Oxyethira Eaton 1873 (Trichoptera: Hydroptilidae) from Northeast Brazil: Eleven new species and distributional records</mods:title>
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<mods:namePart id="268D5929271DC88D4C8FFE6D626CE68D">De Souza, Wagner Rafael M.</mods:namePart>
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<mods:namePart id="D93DAB1473448238D89D0C9C54A75CE2">Santos, Allan Paulo Moreira</mods:namePart>
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<mods:title id="D50FE6B5095F616007DF443F047B1941">Zootaxa</mods:title>
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<mods:date id="4A7C5F937AE5C3BEE373D948EB690E65">2017</mods:date>
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<mods:number id="8A04FAA8141598CE03788683605F2AFC">4236</mods:number>
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<mods:identifier id="8DE3902A10E9BB9395DE87D4B7B9837E" type="DOI">10.11646/zootaxa.4236.3.4</mods:identifier>
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<treatment id="4B1B3448FFA4AC49E7E803EDFA374C33" ID-DOI="http://doi.org/10.5281/zenodo.4513688" ID-GBIF-Taxon="127709884" ID-Zenodo-Dep="4513688" LSID="urn:lsid:plazi:treatment:4B1B3448FFA4AC49E7E803EDFA374C33" httpUri="http://treatment.plazi.org/id/4B1B3448FFA4AC49E7E803EDFA374C33" lastPageId="6" lastPageNumber="490" pageId="2" pageNumber="486">
<subSubSection id="8BA8D6D5FFA4AC4DE7E803EDFEBC4966" pageId="2" pageNumber="486" type="nomenclature">
<paragraph id="C30D855EFFA4AC4DE7E803EDFD324945" blockId="2.[151,664,1023,1083]" box="[151,664,1023,1049]" pageId="2" pageNumber="486">
<heading id="98453232FFA4AC4DE7E803EDFD324945" bold="true" box="[151,664,1023,1049]" fontSize="11" level="1" pageId="2" pageNumber="486" reason="1">
<emphasis id="F1C6594CFFA4AC4DE7E803EDFD324945" bold="true" box="[151,664,1023,1049]" pageId="2" pageNumber="486">
<taxonomicName id="04B2FEDDFFA4AC4DE7E803EDFD994945" box="[151,563,1023,1049]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="retrosa" status="sp. nov." subGenus="Dactylotrichia">
<emphasis id="F1C6594CFFA4AC4DE7E803EDFEBA4945" bold="true" box="[151,272,1023,1049]" italics="true" pageId="2" pageNumber="486">Oxyethira</emphasis>
(
<emphasis id="F1C6594CFFA4AC4DE65F03EDFE664945" bold="true" box="[288,460,1023,1049]" italics="true" pageId="2" pageNumber="486">Dactylotrichia</emphasis>
)
<emphasis id="F1C6594CFFA4AC4DE6A103EDFD994945" bold="true" box="[478,563,1023,1049]" italics="true" pageId="2" pageNumber="486">retrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA4AC4DE54503EDFD324945" box="[570,664,1023,1049]" pageId="2" pageNumber="486" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="C30D855EFFA4AC4DE7E80430FEBC4966" blockId="2.[151,664,1023,1083]" box="[151,278,1058,1083]" pageId="2" pageNumber="486">
<figureCitation id="5B8999DBFFA4AC4DE7E80430FEAB4966" box="[151,257,1058,1083]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig 3A–3</figureCitation>
D
</paragraph>
</subSubSection>
<subSubSection id="8BA8D6D5FFA4AC4DE7E80478FC894B53" pageId="2" pageNumber="486" type="description">
<paragraph id="C30D855EFFA4AC4DE7E80478FC7549FA" blockId="2.[151,1437,1130,1983]" pageId="2" pageNumber="486">
<emphasis id="F1C6594CFFA4AC4DE7E80478FE0C49DF" bold="true" box="[151,422,1130,1155]" pageId="2" pageNumber="486">
Description.
<typeStatus id="1C093BFCFFA4AC4DE64D0478FE0B49DF" box="[306,417,1130,1155]" pageId="2" pageNumber="486" type="holotype">Holotype</typeStatus>
.
</emphasis>
Length: 2.7 mm. General color pale brown (in alcohol). General pattern of Neotropical
<taxonomicName id="04B2FEDDFFA4AC4DE7E8049FFEAD49FA" box="[151,263,1165,1190]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="genus">
<emphasis id="F1C6594CFFA4AC4DE7E8049FFEAD49FA" box="[151,263,1165,1190]" italics="true" pageId="2" pageNumber="486">Oxyethira</emphasis>
</taxonomicName>
. Sternum VII with posterior ventromesal process (
<figureCitation id="5B8999DBFFA4AC4DE441049FFC1649FA" box="[830,956,1165,1190]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Figs. 3A, 3</figureCitation>
C).
</paragraph>
<paragraph id="C30D855EFFA4AC4DE7B804A0FC894B53" blockId="2.[151,1437,1130,1983]" pageId="2" pageNumber="486">
Male genitalia: Segment VIII annular, covered with setae; sternum completely divided longitudinally (
<figureCitation id="5B8999DBFFA4AC4DE21004A0FF1649B3" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3A</figureCitation>
); tergum with posterior margin broadly concave (
<figureCitation id="5B8999DBFFA4AC4DE47E04C4FCED49B3" box="[769,839,1238,1263]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3</figureCitation>
B). Segment IX retracted in segment VIII; sternum produced anteriorly and reaching anterior area of segment VI, with pair of long lateral processes at midlength projected dorsad (
<figureCitation id="5B8999DBFFA4AC4DE61E050CFE05486B" box="[353,431,1310,1335]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Figs. 3</figureCitation>
B, 3C); tergum reduced. Inferior appendages elongate, darkened apically; in ventral view fused basally to sternum IX, digitiform and diverging apically (
<figureCitation id="5B8999DBFFA4AC4DE41F0550FC104806" box="[864,954,1346,1371]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3A</figureCitation>
); in lateral view subrectangular, with same width throughout, curved evenly posterodorsad, and slightly truncate (
<figureCitation id="5B8999DBFFA4AC4DE4C50574FC554823" box="[954,1023,1382,1407]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3</figureCitation>
C). Subgenital plate conspicuous; in ventral and dorsal views with posterior margin truncate (
<figureCitation id="5B8999DBFFA4AC4DE4650598FCDC48FE" box="[794,886,1418,1443]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3A</figureCitation>
); in lateral view tall, projecting caudally beyond segment VIII and with bilobed process about same length (
<figureCitation id="5B8999DBFFA4AC4DE45305BCFCC4489B" box="[812,878,1454,1479]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3</figureCitation>
C). Bilobed process lobes short and globose, each with small apical seta (
<figureCitation id="5B8999DBFFA4AC4DE6E405C0FE7448B6" box="[411,478,1490,1515]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3</figureCitation>
B). Tergum X membranous. Phallus with apex recurved anterad dorsally, crossing its median portion (
<figureCitation id="5B8999DBFFA4AC4DE62F05E4FE394B53" box="[336,403,1526,1551]" captionStart="FIGURE 3" captionStartId="5.[151,250,1975,1997]" captionTargetBox="[235,1352,198,1953]" captionTargetId="figure@5.[235,1352,193,1953]" captionTargetPageId="5" captionText="FIGURE 3 A – 3 D. Oxyethira retrosa sp. nov. Male genitalia: 3 A, ventral; 3 B, dorsal; 3 C, left lateral; 3 D, phallus, dorsal." httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="2" pageNumber="486">Fig. 3</figureCitation>
D); ejaculatory duct not protruding.
</paragraph>
</subSubSection>
<subSubSection id="8BA8D6D5FFA4AC4DE7B80608FEE74A2A" pageId="2" pageNumber="486" type="discussion">
<paragraph id="C30D855EFFA4AC4DE7B80608FEE74A2A" blockId="2.[151,1437,1130,1983]" pageId="2" pageNumber="486">
<emphasis id="F1C6594CFFA4AC4DE7B80608FE914B6F" bold="true" box="[199,315,1562,1587]" pageId="2" pageNumber="486">Remarks.</emphasis>
This species is quite similar to others assigned to the subgenus
<emphasis id="F1C6594CFFA4AC4DE36F0608FB1E4B6F" box="[1040,1204,1562,1587]" italics="true" pageId="2" pageNumber="486">Dactylotrichia</emphasis>
due to sternum VIII completely divided and segment IX reaching beyond segments VIII–VII internally (
<bibRefCitation id="A723F8AFFFA4AC4DE3EA062CFA864B0B" author="Kelley" box="[1173,1324,1598,1623]" pageId="2" pageNumber="486" refString="Kelley, R. W. (1984) Phylogeny, morphology and classification of the micro-caddisfly genus Oxyethira Eaton (Tricboptera: Hydroptilidae). Transactions of the American Entomological Society, 110, 435 - 463." type="journal article" year="1984">Kelley 1984</bibRefCitation>
). Among
<emphasis id="F1C6594CFFA4AC4DE7E80670FE914B27" box="[151,315,1634,1659]" italics="true" pageId="2" pageNumber="486">Dactylotrichia</emphasis>
species,
<taxonomicName id="04B2FEDDFFA4AC4DE6DB0670FDC14B26" box="[420,619,1634,1659]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="retrosa" status="sp. nov.">
<emphasis id="F1C6594CFFA4AC4DE6DB0670FDC14B26" box="[420,619,1634,1659]" italics="true" pageId="2" pageNumber="486">Oxyethira retrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA4AC4DE50B0670FD654B27" box="[628,719,1634,1659]" pageId="2" pageNumber="486" rank="species">
<emphasis id="F1C6594CFFA4AC4DE50B0670FD654B27" bold="true" box="[628,719,1634,1659]" pageId="2" pageNumber="486">sp. nov.</emphasis>
</taxonomicNameLabel>
is most similar to
<taxonomicName id="04B2FEDDFFA4AC4DE4D70671FAED4B26" authority="Harris &amp; Davenport 1999" authorityName="Harris &amp; Davenport" authorityYear="1999" box="[936,1351,1634,1659]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="hozosa">
<emphasis id="F1C6594CFFA4AC4DE4D70671FBB04B27" box="[936,1050,1634,1659]" italics="true" pageId="2" pageNumber="486">O. hozosa</emphasis>
<bibRefCitation id="A723F8AFFFA4AC4DE35D0670FAED4B26" author="Harris" box="[1058,1351,1634,1659]" pageId="2" pageNumber="486" refString="Harris, S. C. &amp; Davenport, L. J. (1999) New species of Hydroptilidae (Trichoptera) from the Amazon region of northeastern Peru. Proceedings of the Entomological Society of Washington, 101, 26 - 38." type="journal article" year="1999">Harris &amp; Davenport 1999</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="04B2FEDDFFA4AC4DE2FE0671FE114BC3" authority="Kelley 1983" authorityName="Kelley" authorityYear="1983" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="scaeodactyla">
<emphasis id="F1C6594CFFA4AC4DE2FE0671FE834BC3" italics="true" pageId="2" pageNumber="486">O. scaeodactyla</emphasis>
<bibRefCitation id="A723F8AFFFA4AC4DE64E0694FE114BC3" author="Kelley" box="[305,443,1670,1695]" pageId="2" pageNumber="486" refString="Kelley, R. W. (1983) New Neotropical species of Oxyethira (Trichoptera: Hydroptilidae). Proceedings of the Entomological Society of Washington, 85 (1), 41 - 54." type="journal article" year="1983">Kelley 1983</bibRefCitation>
</taxonomicName>
because of the elongate inferior appendages, but is the inferior appendages are wider in the new species than in those other two. Moreover, segment IX also has lateral projections directed dorsad in both
<taxonomicName id="04B2FEDDFFA4AC4DE7E806DDFEA14BBA" box="[151,267,1743,1766]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="retrosa" status="sp. nov.">
<emphasis id="F1C6594CFFA4AC4DE7E806DDFEA14BBA" box="[151,267,1743,1766]" italics="true" pageId="2" pageNumber="486">O. retrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA4AC4DE66B06DCFEDA4BBB" box="[276,368,1742,1767]" pageId="2" pageNumber="486" rank="species">
<emphasis id="F1C6594CFFA4AC4DE66B06DCFEDA4BBB" bold="true" box="[276,368,1742,1767]" pageId="2" pageNumber="486">sp. nov.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="04B2FEDDFFA4AC4DE6D306DDFDB54BBB" box="[428,543,1742,1767]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="hozosa">
<emphasis id="F1C6594CFFA4AC4DE6D306DDFDB54BBB" box="[428,543,1742,1767]" italics="true" pageId="2" pageNumber="486">O. hozosa</emphasis>
</taxonomicName>
. The new species is also similar to
<taxonomicName id="04B2FEDDFFA4AC4DE4BE06DCFAF64BBB" authority="Angrisano 1995b" authorityName="Angrisano" authorityYear="1995" box="[961,1372,1742,1767]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="acegua">
<emphasis id="F1C6594CFFA4AC4DE4BE06DCFB264BBA" box="[961,1164,1742,1767]" italics="true" pageId="2" pageNumber="486">Oxyethira acegua</emphasis>
<bibRefCitation id="A723F8AFFFA4AC4DE3EA06DCFAF64BBB" author="Angrisano" box="[1173,1372,1742,1767]" pageId="2" pageNumber="486" refString="Angrisano, E. B. (1995 b) Contribucion al conocimiento de los Trichoptera del Uruguay. II. Familia Hydroptilidae. Revista Brasileira de Entomologia, 39, 501 - 516." type="journal article" year="1995" yearSuffix="b">Angrisano 1995b</bibRefCitation>
</taxonomicName>
, both species having a recurved phallic apex. However, in the new species the recurved portion crosses the phallus dorsally, whereas in
<taxonomicName id="04B2FEDDFFA4AC4DE6F90705FE574A72" box="[390,509,1815,1838]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="acegua">
<emphasis id="F1C6594CFFA4AC4DE6F90705FE574A72" box="[390,509,1815,1838]" italics="true" pageId="2" pageNumber="486">O. acegua</emphasis>
</taxonomicName>
the phallus is recurved anterad laterally.
<taxonomicName id="04B2FEDDFFA4AC4DE4A00704FB024A72" box="[991,1192,1814,1839]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="retrosa" status="sp. nov.">
<emphasis id="F1C6594CFFA4AC4DE4A00704FB024A72" box="[991,1192,1814,1839]" italics="true" pageId="2" pageNumber="486">Oxyethira retrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA4AC4DE3CD0704FAA54A73" box="[1202,1295,1814,1839]" pageId="2" pageNumber="486" rank="species">
<emphasis id="F1C6594CFFA4AC4DE3CD0704FAA54A73" bold="true" box="[1202,1295,1814,1839]" pageId="2" pageNumber="486">sp. nov.</emphasis>
</taxonomicNameLabel>
can also be identified by the tall subgenital plate produced caudad beyond segment VIII and about as long as the lobes of the bilobed process.
</paragraph>
</subSubSection>
<subSubSection id="8BA8D6D5FFA4AC4DE7B80790FDC54AE3" pageId="2" pageNumber="486" type="etymology">
<paragraph id="C30D855EFFA4AC4DE7B80790FDC54AE3" blockId="2.[151,1437,1130,1983]" pageId="2" pageNumber="486">
<emphasis id="F1C6594CFFA4AC4DE7B80790FEE44AC7" bold="true" box="[199,334,1922,1947]" pageId="2" pageNumber="486">Etymology.</emphasis>
The specific name is an allusion to the phallus shape. The word “
<taxonomicName id="04B2FEDDFFA4AC4DE33A0790FB3C4AC6" box="[1093,1174,1922,1946]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="486" phylum="Arthropoda" rank="species" species="retrosa">retrosa</taxonomicName>
” is derived from Latin “
<emphasis id="F1C6594CFFA4AC4DE7DC07B5FEAF4AE2" box="[163,261,1959,1982]" italics="true" pageId="2" pageNumber="486">retrosum</emphasis>
” that means directed backward.
</paragraph>
</subSubSection>
<caption id="97CDD5D6FFA5AC4CE7E807D7FAFF4A86" box="[151,1365,1988,2011]" httpUri="https://zenodo.org/record/322267/files/figure.png" pageId="3" pageNumber="487" targetBox="[257,1324,196,1961]" targetPageId="3">
<paragraph id="C30D855EFFA5AC4CE7E807D7FAFF4A86" blockId="3.[151,1365,1988,2011]" box="[151,1365,1988,2011]" pageId="3" pageNumber="487">
<emphasis id="F1C6594CFFA5AC4CE7E807D7FEFA4A86" bold="true" box="[151,336,1989,2011]" pageId="3" pageNumber="487">FIGURE 1A–1D.</emphasis>
<taxonomicName id="04B2FEDDFFA5AC4CE62807D6FE544A86" box="[343,510,1988,2010]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="3" pageNumber="487" phylum="Arthropoda" rank="species" species="calori" status="sp. nov.">
<emphasis id="F1C6594CFFA5AC4CE62807D6FE544A86" box="[343,510,1988,2010]" italics="true" pageId="3" pageNumber="487">Oxyethira calori</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA5AC4CE57A07D7FDFC4A86" box="[517,598,1989,2010]" pageId="3" pageNumber="487" rank="species">
<emphasis id="F1C6594CFFA5AC4CE57A07D7FDFC4A86" bold="true" box="[517,598,1989,2010]" pageId="3" pageNumber="487">sp. nov.</emphasis>
</taxonomicNameLabel>
Male genitalia: 1A, ventral; 1B, dorsal; 1C, left lateral; 1D, phallus, dorsal.
</paragraph>
</caption>
<caption id="97CDD5D6FFA2AC4BE7E807A9FAD44A8D" box="[151,1406,1979,2001]" httpUri="https://zenodo.org/record/322268/files/figure.png" pageId="4" pageNumber="488" targetBox="[237,1350,193,1958]" targetPageId="4">
<paragraph id="C30D855EFFA2AC4BE7E807A9FAD44A8D" blockId="4.[151,1406,1979,2001]" box="[151,1406,1979,2001]" pageId="4" pageNumber="488">
<emphasis id="F1C6594CFFA2AC4BE7E807A9FEFB4A8C" bold="true" box="[151,337,1979,2001]" pageId="4" pageNumber="488">FIGURE 2A–2D.</emphasis>
<taxonomicName id="04B2FEDDFFA2AC4BE62807A9FD8D4A8D" box="[343,551,1979,2001]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="4" pageNumber="488" phylum="Arthropoda" rank="species" species="iannuzzae" status="sp. nov.">
<emphasis id="F1C6594CFFA2AC4BE62807A9FD8D4A8D" box="[343,551,1979,2001]" italics="true" pageId="4" pageNumber="488">Oxyethira iannuzzae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA2AC4BE55107A9FDD44A8C" box="[558,638,1979,2000]" pageId="4" pageNumber="488" rank="species">
<emphasis id="F1C6594CFFA2AC4BE55107A9FDD44A8C" bold="true" box="[558,638,1979,2000]" pageId="4" pageNumber="488">sp. nov.</emphasis>
</taxonomicNameLabel>
Male genitalia: 2A, ventral; 2B, dorsal; 2C, left lateral; 2D, phallus, dorsal.
</paragraph>
</caption>
<caption id="97CDD5D6FFA3AC4AE7E807A5FACA4A91" box="[151,1376,1975,1997]" httpUri="https://zenodo.org/record/322269/files/figure.png" pageId="5" pageNumber="489" targetBox="[235,1352,198,1953]" targetPageId="5">
<paragraph id="C30D855EFFA3AC4AE7E807A5FACA4A91" blockId="5.[151,1376,1975,1997]" box="[151,1376,1975,1997]" pageId="5" pageNumber="489">
<emphasis id="F1C6594CFFA3AC4AE7E807A5FEFB4A90" bold="true" box="[151,337,1975,1997]" pageId="5" pageNumber="489">FIGURE 3A–3D.</emphasis>
<taxonomicName id="04B2FEDDFFA3AC4AE62807A5FDA34A91" box="[343,521,1975,1997]" class="Insecta" family="Hydroptilidae" genus="Oxyethira" kingdom="Animalia" order="Trichoptera" pageId="5" pageNumber="489" phylum="Arthropoda" rank="species" species="retrosa" status="sp. nov.">
<emphasis id="F1C6594CFFA3AC4AE62807A5FDA34A91" box="[343,521,1975,1997]" italics="true" pageId="5" pageNumber="489">Oxyethira retrosa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="EAF5E437FFA3AC4AE56F07A5FDCA4A90" box="[528,608,1975,1996]" pageId="5" pageNumber="489" rank="species">
<emphasis id="F1C6594CFFA3AC4AE56F07A5FDCA4A90" bold="true" box="[528,608,1975,1996]" pageId="5" pageNumber="489">sp. nov.</emphasis>
</taxonomicNameLabel>
Male genitalia: 3A, ventral; 3B, dorsal; 3C, left lateral; 3D, phallus, dorsal.
</paragraph>
</caption>
<subSubSection id="8BA8D6D5FFA0AC49E7B80085FA374C33" pageId="6" pageNumber="490" type="materials_examined">
<paragraph id="C30D855EFFA0AC49E7B80085FA374C33" blockId="6.[151,1437,151,368]" pageId="6" pageNumber="490">
<materialsCitation id="73DA8F03FFA0AC49E7B80085FD914DA0" ID-GBIF-Occurrence="1437079675" collectingDate="2014-06-17" collectionCode="APM, DZRJ" collectorName="Parque Nacional da Serra de Itabaiana &amp; Riacho dos Negros &amp; Santos, DM &amp; AC Domahovski &amp; Souza" country="Brazil" elevation="202" latitude="-10.747223" location="Itabaiana" longLatPrecision="21" longitude="-37.34" municipality="Examined" pageId="6" pageNumber="490" specimenCount="1" specimenCount-male="1" stateProvince="Sergipe" typeStatus="holotype">
<emphasis id="F1C6594CFFA0AC49E7B80085FCBF4DEC" bold="true" box="[199,789,151,176]" pageId="6" pageNumber="490">
<collectingMunicipality id="23691F24FFA0AC49E7B80085FEEA4DEC" box="[199,320,151,176]" pageId="6" pageNumber="490">Examined</collectingMunicipality>
material.
<typeStatus id="1C093BFCFFA0AC49E6C40085FD804DEC" box="[443,554,151,176]" pageId="6" pageNumber="490" type="holotype">Holotype</typeStatus>
.
<collectingCountry id="BBA5C5CEFFA0AC49E5490085FD0F4DEC" box="[566,677,151,176]" name="Brazil" pageId="6" pageNumber="490">BRAZIL</collectingCountry>
:
<collectingRegion id="01764BBCFFA0AC49E5CD0085FCBA4DEC" box="[690,784,151,176]" country="Brazil" name="Sergipe" pageId="6" pageNumber="490">Sergipe</collectingRegion>
:
</emphasis>
<location id="C66DD385FFA0AC49E4630085FC2C4DEC" LSID="urn:lsid:plazi:treatment:4B1B3448FFA4AC49E7E803EDFA374C33:C66DD385FFA0AC49E4630085FC2C4DEC" box="[796,902,151,176]" country="Brazil" latitude="-10.747223" longLatPrecision="21" longitude="-37.34" municipality="Examined" name="Itabaiana" pageId="6" pageNumber="490" stateProvince="Sergipe">Itabaiana</location>
,
<collectorName id="6E47E088FFA0AC49E4EE008AFAE84DEC" box="[913,1346,151,176]" pageId="6" pageNumber="490">Parque Nacional da Serra de Itabaiana</collectorName>
,
<collectorName id="6E47E088FFA0AC49E2320085FEB64D89" pageId="6" pageNumber="490">Riacho dos Negros</collectorName>
,
<geoCoordinate id="A686E399FFA0AC49E65500AEFE004D89" box="[298,426,188,213]" direction="south" orientation="latitude" pageId="6" pageNumber="490" precision="15" value="-10.747223">10°44'50''S</geoCoordinate>
,
<geoCoordinate id="A686E399FFA0AC49E6C600AEFDE84D88" box="[441,578,188,212]" direction="west" orientation="longitude" pageId="6" pageNumber="490" precision="15" value="-37.34">37°20'24''W</geoCoordinate>
,
<elevation id="489F626DFFA0AC49E52F00AEFD374D88" box="[592,669,188,212]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="6" pageNumber="490" unit="m" value="202.0">
<quantity id="044A28BBFFA0AC49E52F00AEFD374D88" box="[592,669,188,212]" metricMagnitude="2" metricUnit="m" metricValue="2.02" pageId="6" pageNumber="490" unit="m" value="202.0">202 m</quantity>
</elevation>
,
<collectingDate id="A7485A76FFA0AC49E5D300AEFC8C4D89" box="[684,806,188,213]" pageId="6" pageNumber="490" value="2014-06-17">
<date id="B70CA39EFFA0AC49E5D300AEFC8C4D89" box="[684,806,188,213]" pageId="6" pageNumber="490" value="2014-06-17">17.vi.2014</date>
</collectingDate>
,
<collectionCode id="A5A31D9BFFA0AC49E44A00AEFCD84D88" box="[821,882,188,212]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15304" name="Algonquin Provincial Park" pageId="6" pageNumber="490">APM</collectionCode>
<collectorName id="6E47E088FFA0AC49E40300AEFBAF4D88" box="[892,1029,188,212]" pageId="6" pageNumber="490">Santos, DM</collectorName>
Takiya,
<collectorName id="6E47E088FFA0AC49E31300AEFA864D89" box="[1132,1324,188,213]" pageId="6" pageNumber="490">AC Domahovski</collectorName>
&amp; WRM
<collectorName id="6E47E088FFA0AC49E7E800F6FF714DA0" box="[151,219,228,252]" pageId="6" pageNumber="490">Souza</collectorName>
cols.,
<collectingMethod id="1AF3FD49FFA0AC49E65200F1FE334DA7" box="[301,409,227,252]" pageId="6" pageNumber="490">light trap</collectingMethod>
,
<specimenCount id="D5B44ED7FFA0AC49E6D600F6FE7D4DA7" box="[425,471,228,252]" pageId="6" pageNumber="490" type="male">1 ♂</specimenCount>
(
<collectionCode id="A5A31D9BFFA0AC49E69500F6FD994DA0" box="[490,563,228,252]" pageId="6" pageNumber="490">DZRJ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="73DA8F03FFA0AC49E53500F1FB2A4DA0" ID-GBIF-Occurrence="1437079689" box="[586,1152,227,252]" collectionCode="DZRJ" country="Brazil" location="Same" pageId="6" pageNumber="490" specimenCount="1" specimenCount-male="1" stateProvince="Sergipe" typeStatus="paratype">
<emphasis id="F1C6594CFFA0AC49E53500F1FD634DA0" bold="true" box="[586,713,227,252]" pageId="6" pageNumber="490">
<typeStatus id="1C093BFCFFA0AC49E53500F1FD6E4DA0" box="[586,708,227,252]" pageId="6" pageNumber="490" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<location id="C66DD385FFA0AC49E5AB00F6FCB84DA0" LSID="urn:lsid:plazi:treatment:4B1B3448FFA4AC49E7E803EDFA374C33:C66DD385FFA0AC49E5AB00F6FCB84DA0" box="[724,786,228,252]" country="Brazil" name="Same" pageId="6" pageNumber="490" stateProvince="Sergipe">Same</location>
data as
<typeStatus id="1C093BFCFFA0AC49E40600F1FC744DA0" box="[889,990,227,252]" pageId="6" pageNumber="490" type="holotype">holotype</typeStatus>
,
<specimenCount id="D5B44ED7FFA0AC49E49100F6FBB64DA7" box="[1006,1052,228,252]" pageId="6" pageNumber="490" type="male">1 ♂</specimenCount>
(
<collectionCode id="A5A31D9BFFA0AC49E35100F6FBDD4DA0" box="[1070,1143,228,252]" pageId="6" pageNumber="490">DZRJ</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="73DA8F03FFA0AC49E3F000F1FC1C4C14" ID-GBIF-Occurrence="1437079704" collectingDate="2014-06-20" collectionCode="APM, DZRJ" collectorName="Reserva Biologica de Pedra Talhada &amp; Bica da Juliana &amp; Santos, DM &amp; AC Domahovski &amp; Souza" country="Brazil" county="Quebrangulo" latitude="-9.239362" location="Bica da Juliana" longLatPrecision="1" longitude="-36.453167" municipality="Reserva Biologica de Pedra Talhada" pageId="6" pageNumber="490" specimenCount="2" specimenCount-male="2" stateProvince="Alagoas" typeStatus="paratype">
<emphasis id="F1C6594CFFA0AC49E3F000F1FB524DA0" bold="true" box="[1167,1272,227,252]" pageId="6" pageNumber="490">
<collectingRegion id="01764BBCFFA0AC49E3F000F1FB584DA0" box="[1167,1266,227,252]" country="Brazil" name="Alagoas" pageId="6" pageNumber="490">Alagoas</collectingRegion>
:
</emphasis>
<collectingCounty id="2A6CFDD2FFA0AC49E27C00F1FA324DA0" box="[1283,1432,227,252]" pageId="6" pageNumber="490">Quebrangulo</collectingCounty>
,
<collectingMunicipality id="23691F24FFA0AC49E7E8011AFD934C7C" box="[151,569,263,288]" pageId="6" pageNumber="490">
<collectorName id="6E47E088FFA0AC49E7E8011AFD934C7C" box="[151,569,263,288]" pageId="6" pageNumber="490">Reserva Biológica de Pedra Talhada</collectorName>
</collectingMunicipality>
,
<location id="C66DD385FFA0AC49E5390115FD534C7C" LSID="urn:lsid:plazi:treatment:4B1B3448FFA4AC49E7E803EDFA374C33:C66DD385FFA0AC49E5390115FD534C7C" box="[582,761,263,288]" country="Brazil" county="Quebrangulo" latitude="-9.239362" longLatPrecision="1" longitude="-36.453167" municipality="Reserva Biologica de Pedra Talhada" name="Bica da Juliana" pageId="6" pageNumber="490" stateProvince="Alagoas">
<collectorName id="6E47E088FFA0AC49E5390115FD534C7C" box="[582,761,263,288]" pageId="6" pageNumber="490">Bica da Juliana</collectorName>
</location>
,
<geoCoordinate id="A686E399FFA0AC49E479011AFC274C7C" box="[774,909,264,288]" direction="south" orientation="latitude" pageId="6" pageNumber="490" precision="1" value="-9.239362">9°14'21,7''S</geoCoordinate>
,
<geoCoordinate id="A686E399FFA0AC49E4E40115FB924C7C" box="[923,1080,263,288]" direction="west" orientation="longitude" pageId="6" pageNumber="490" precision="1" value="-36.453167">36°27'11,4''W</geoCoordinate>
,
<collectingDate id="A7485A76FFA0AC49E3390115FB6A4C7C" box="[1094,1216,263,288]" pageId="6" pageNumber="490" value="2014-06-20">
<date id="B70CA39EFFA0AC49E3390115FB6A4C7C" box="[1094,1216,263,288]" pageId="6" pageNumber="490" value="2014-06-20">20.vi.2014</date>
</collectingDate>
,
<collectionCode id="A5A31D9BFFA0AC49E3B1011AFAA14C7C" box="[1230,1291,264,288]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15304" name="Algonquin Provincial Park" pageId="6" pageNumber="490">APM</collectionCode>
<collectorName id="6E47E088FFA0AC49E26B011AFA364C43" box="[1300,1436,264,288]" pageId="6" pageNumber="490">Santos, DM</collectorName>
Takiya,
<collectorName id="6E47E088FFA0AC49E78D0122FE1A4C14" box="[242,432,303,328]" pageId="6" pageNumber="490">AC Domahovski</collectorName>
&amp; WRM
<collectorName id="6E47E088FFA0AC49E55B0122FDC24C14" box="[548,616,304,328]" pageId="6" pageNumber="490">Souza</collectorName>
cols.,
<collectingMethod id="1AF3FD49FFA0AC49E5CB013DFCB74C1B" box="[692,797,303,328]" pageId="6" pageNumber="490">light trap</collectingMethod>
,
<specimenCount id="D5B44ED7FFA0AC49E4560122FCFF4C1B" box="[809,853,304,328]" pageId="6" pageNumber="490" type="male">2 ♂</specimenCount>
(
<collectionCode id="A5A31D9BFFA0AC49E41A0122FC044C14" box="[869,942,304,328]" pageId="6" pageNumber="490">DZRJ</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="73DA8F03FFA0AC49E4BE013DFA374C33" ID-GBIF-Occurrence="1437079717" collectingDate="2013-08-28" collectorName="Reserva Biologica de Pedra Talhada &amp; Lima" country="Brazil" county="Quebrangulo" elevation="661" latitude="-9.257222" location="Rio Caranguejo" longLatPrecision="21" longitude="-36.431667" municipality="Reserva Biologica de Pedra Talhada" pageId="6" pageNumber="490" specimenCount="2" specimenCount-male="2" stateProvince="Alagoas" typeStatus="paratype">
<collectingCounty id="2A6CFDD2FFA0AC49E4BE013DFBFC4C14" box="[961,1110,303,328]" pageId="6" pageNumber="490">Quebrangulo</collectingCounty>
,
<collectorName id="6E47E088FFA0AC49E31C0122FF584C2C" pageId="6" pageNumber="490">
<collectingMunicipality id="23691F24FFA0AC49E31C0122FF584C2C" pageId="6" pageNumber="490">Reserva Biológica de Pedra Talhada</collectingMunicipality>
</collectorName>
,
<location id="C66DD385FFA0AC49E7820145FE1A4C2C" LSID="urn:lsid:plazi:treatment:4B1B3448FFA4AC49E7E803EDFA374C33:C66DD385FFA0AC49E7820145FE1A4C2C" box="[253,432,343,368]" country="Brazil" county="Quebrangulo" latitude="-9.257222" longLatPrecision="21" longitude="-36.431667" municipality="Reserva Biologica de Pedra Talhada" name="Rio Caranguejo" pageId="6" pageNumber="490" stateProvince="Alagoas">Rio Caranguejo</location>
,
<geoCoordinate id="A686E399FFA0AC49E6C40145FD844C2C" box="[443,558,343,368]" direction="south" orientation="latitude" pageId="6" pageNumber="490" precision="15" value="-9.257222">9°15'26''S</geoCoordinate>
,
<geoCoordinate id="A686E399FFA0AC49E5460145FD6B4C2C" box="[569,705,343,368]" direction="west" orientation="longitude" pageId="6" pageNumber="490" precision="15" value="-36.431667">36°25'54''W</geoCoordinate>
,
<elevation id="489F626DFFA0AC49E5B20145FCBC4C33" box="[717,790,343,368]" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" pageId="6" pageNumber="490" unit="m" value="661.0">
<quantity id="044A28BBFFA0AC49E5B20145FCBC4C33" box="[717,790,343,368]" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" pageId="6" pageNumber="490" unit="m" value="661.0">661 m</quantity>
</elevation>
,
<collectingDate id="A7485A76FFA0AC49E45E0145FC014C2C" box="[801,939,343,368]" pageId="6" pageNumber="490" value="2013-08-28">
<date id="B70CA39EFFA0AC49E45E0145FC014C2C" box="[801,939,343,368]" pageId="6" pageNumber="490" value="2013-08-28">28.viii.2013</date>
</collectingDate>
, LRC
<collectorName id="6E47E088FFA0AC49E48B0145FB854C2C" box="[1012,1071,343,368]" pageId="6" pageNumber="490">Lima</collectorName>
col.,
<collectingMethod id="1AF3FD49FFA0AC49E3120145FB7F4C33" box="[1133,1237,343,368]" pageId="6" pageNumber="490">light trap</collectingMethod>
,
<specimenCount id="D5B44ED7FFA0AC49E39F0145FAA04C33" box="[1248,1290,343,367]" pageId="6" pageNumber="490" type="male">2 ♂</specimenCount>
(CE-UFPE).
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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