625 lines
78 KiB
XML
625 lines
78 KiB
XML
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<mods:title id="7C042AB583ECFD1AABF86CF8300350CC">Description of two new species forming a sympatric species pair of Crenicichla (Teleostei: Cichlidae) endemic to the Piray Guazú River in the Paraná River Basin, Misiones, Argentina and belonging to the C. mandelburgeri species complex</mods:title>
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<mods:namePart id="056C63C3742917A6B9659BFDEF8D9707">Říčan, Oldřich</mods:namePart>
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<mods:affiliation id="01CAC48E834976F110F9B761AF97000E">University of South Bohemia, Faculty of Science, Department of Zoology, Branišovská 31, 370 05, České Budějovice, Czech Republic. UNLP, Facultad de Ciencias Naturales y Museo, División Zoología Vertebrados, Paseo del Bosque, 1900 La Plata, Buenos Aires Province, Argentina. CIC, Comisión de Investigaciones Científicas de la Provincia de Buenos Aires.</mods:affiliation>
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<mods:nameIdentifier id="D02E0AF391BC6C020131C4B9A99B04E3" type="email">oldrich.rican@prf.jcu.cz</mods:nameIdentifier>
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<mods:namePart id="B15CEBF0F1A5FE6DF3C9E7E9A2FAD25C">Piálek, Lubomír</mods:namePart>
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<mods:affiliation id="B13D5E04E7E749AA3DF6582491D8BD21">University of South Bohemia, Faculty of Science, Department of Zoology, Branišovská 31, 370 05, České Budějovice, Czech Republic. UNLP, Facultad de Ciencias Naturales y Museo, División Zoología Vertebrados, Paseo del Bosque, 1900 La Plata, Buenos Aires Province, Argentina. CIC, Comisión de Investigaciones Científicas de la Provincia de Buenos Aires.</mods:affiliation>
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<mods:nameIdentifier id="182C20434F45381DFB9469E7F3756EE2" type="email">lpialek@yahoo.com</mods:nameIdentifier>
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<mods:namePart id="81E3F0874EA6028AEC7F9D35E0257A50">Almirón, Adriana</mods:namePart>
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<mods:namePart id="3992312E4D474B0722A83FC0C8DFC0F5">Casciotta, Jorge</mods:namePart>
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<mods:title id="41160C3388BDFEFEE04ACD7A650029F3">European Journal of Taxonomy</mods:title>
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<mods:date id="1163A5D3F0C889335C53B24BD1912BB8">2023</mods:date>
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<mods:number id="0EEB54BECCE34785573BA6372FEA4A2F">2023-07-07</mods:number>
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<treatment id="BF3B7739FFACFF863DB5F9ABFA81FC0B" ID-DOI="http://doi.org/10.5281/zenodo.8138241" ID-GBIF-Taxon="211053143" ID-Zenodo-Dep="8138241" LSID="urn:lsid:plazi:treatment:BF3B7739FFACFF863DB5F9ABFA81FC0B" httpUri="http://treatment.plazi.org/id/BF3B7739FFACFF863DB5F9ABFA81FC0B" lastPageId="19" lastPageNumber="57" pageId="15" pageNumber="53">
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<subSubSection id="7F8895A4FFACFF9A3DB5F9ABFCF2F9A3" pageId="15" pageNumber="53" type="nomenclature">
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<emphasis id="05E61A3DFFACFF9A3DB5F9ABFCF2F9A3" bold="true" italics="true" pageId="15" pageNumber="53">
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Keys to the species of
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<taxonomicName id="F092BDACFFACFF9A3CF3F9AAFD60F989" ID-CoL="3VK6" ID-ENA="50736" authorityName="Heckel" authorityYear="1840" box="[507,662,1590,1619]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="53" phylum="Chordata" rank="genus">Crenicichla</taxonomicName>
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from the Middle Rio
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<collectingRegion id="F55608CDFFACFF9A3ED8F9AAFBC4F989" box="[976,1074,1590,1619]" country="Brazil" name="Parana" pageId="15" pageNumber="53">Paraná</collectingRegion>
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Basin (the
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<taxonomicName id="F092BDACFFACFF9A39D4F9AAFE77F9A3" authorityName="Kullander" authorityYear="2009" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="53" phylum="Chordata" rank="species" species="mandelburgeri">Crenicichla mandelburgeri</taxonomicName>
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species group plus
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<taxonomicName id="F092BDACFFACFF9A3F8AF9C1FD0CF9A3" authorityName="Heckel" authorityYear="1840" box="[642,762,1629,1657]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="53" phylum="Chordata" rank="species" species="vittata">C. vittata</taxonomicName>
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)
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="7F8895A4FFACFF853DB5F90DFDEBFE6A" lastPageId="16" lastPageNumber="54" pageId="15" pageNumber="53" type="discussion">
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<paragraph id="372DC62FFFACFF9A3DB5F90DFCA8F92B" blockId="15.[188,1398,1681,1778]" pageId="15" pageNumber="53">
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Because of the taxonomical and morphological complexity of the
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<taxonomicName id="F092BDACFFACFF9A3EB0F90DFB71F971" authorityName="Kullander" authorityYear="2009" box="[952,1159,1681,1707]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="53" phylum="Chordata" rank="species" species="mandelburgeri">
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<emphasis id="05E61A3DFFACFF9A3EB0F90DFB71F971" box="[952,1159,1681,1707]" italics="true" pageId="15" pageNumber="53">C. mandelburgeri</emphasis>
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</taxonomicName>
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species complex we have chosen to provide two determination keys, each primarily based on a different main character. We believe this may facilitate determination of some species.
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</paragraph>
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<paragraph id="372DC62FFFACFF9A3DB5F885FD0BF88C" blockId="15.[189,1399,1817,1878]" pageId="15" pageNumber="53">The first key is more suited to the present study as it is based on the main colouration patterns, which also distinguish the two newly described species.</paragraph>
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<paragraph id="372DC62FFFACFF9A3DB5F8E1FAB6F804" blockId="15.[189,1399,1917,2014]" pageId="15" pageNumber="53">
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The second key is more generally suited to the
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<taxonomicName id="F092BDACFFACFF9A3E0FF8E2FC2CF84D" authorityName="Kullander" authorityYear="2009" box="[775,986,1917,1944]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="53" phylum="Chordata" rank="species" species="mandelburgeri">
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<emphasis id="05E61A3DFFACFF9A3E0FF8E2FC2CF84D" box="[775,986,1917,1944]" italics="true" pageId="15" pageNumber="53">C. mandelburgeri</emphasis>
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</taxonomicName>
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species complex and the existing literature on parallel evolution of ecomorphologies in
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<taxonomicName id="F092BDACFFACFF9A3E65F83CFC00F860" authorityName="Heckel" authorityYear="1840" box="[877,1014,1952,1978]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="15" pageNumber="53" phylum="Chordata" rank="genus">
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<emphasis id="05E61A3DFFACFF9A3E65F83CFC00F860" box="[877,1014,1952,1978]" italics="true" pageId="15" pageNumber="53">Crenicichla</emphasis>
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</taxonomicName>
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since it is based on oral jaw characteristics (the main charaters involved in the formation and parallel evolution of ecomorphs).
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</paragraph>
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<paragraph id="372DC62FFFB3FF853DB5FE95FDEBFE6A" blockId="16.[188,1399,265,432]" pageId="16" pageNumber="54">
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For the keys we have chosen to use only readily visible external characters. Many of the characters used in the key will work only for adult specimens (at least
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SL), especially the colour and colouration pattern characters. The main colouration pattern of the species is scored based on preserved specimens where there is less variation than in live specimens, while the colouration of the dorsal fin in females is best visible in live specimens.
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</paragraph>
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</subSubSection>
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<subSubSection id="7F8895A4FFB3FF853DB5FE4BFBF3FDCF" pageId="16" pageNumber="54" type="multiple">
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<paragraph id="372DC62FFFB3FF853DB5FE4BFBF3FDCF" blockId="16.[189,1396,470,533]" pageId="16" pageNumber="54">
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<emphasis id="05E61A3DFFB3FF853DB5FE4BFBF3FDCF" bold="true" pageId="16" pageNumber="54">
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Key to the species of
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<taxonomicName id="F092BDACFFB3FF853CB0FE4BFDB0FE28" authorityName="Heckel" authorityYear="1840" box="[440,582,471,498]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="genus">
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<emphasis id="05E61A3DFFB3FF853CB0FE4BFDB0FE28" bold="true" box="[440,582,471,498]" italics="true" pageId="16" pageNumber="54">Crenicichla</emphasis>
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</taxonomicName>
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from the Middle Rio
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<collectingRegion id="F55608CDFFB3FF853E45FE4AFC5EFE2B" box="[845,936,470,497]" country="Brazil" name="Parana" pageId="16" pageNumber="54">Paraná</collectingRegion>
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Basin (the
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<taxonomicName id="F092BDACFFB3FF853926FE4BFA82FE28" authorityName="Kullander" authorityYear="2009" box="[1070,1396,471,498]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="mandelburgeri">
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<emphasis id="05E61A3DFFB3FF853926FE4BFA82FE28" bold="true" box="[1070,1396,471,498]" italics="true" pageId="16" pageNumber="54">Crenicichla mandelburgeri</emphasis>
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</taxonomicName>
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<heading id="6C657143FFB3FF853DB5FE66FBF3FDCF" bold="true" box="[189,1029,506,533]" fontSize="11" level="3" pageId="16" pageNumber="54" reason="3">
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species group plus
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||
<taxonomicName id="F092BDACFFB3FF853CA0FE66FDE3FDCE" authorityName="Heckel" authorityYear="1840" box="[424,533,506,532]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="vittata">
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<emphasis id="05E61A3DFFB3FF853CA0FE66FDE3FDCE" bold="true" box="[424,533,506,532]" italics="true" pageId="16" pageNumber="54">C. vittata</emphasis>
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</taxonomicName>
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) based on the main colouration pattern
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</heading>
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="7F8895A4FFB3FF843DB5FDA0FA81F993" lastPageId="17" lastPageNumber="55" pageId="16" pageNumber="54" type="key">
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<key id="4303911EFFB3FF843DB5FDA0FA81F993" lastPageId="17" lastPageNumber="55" pageId="16" pageNumber="54">
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<keyStep id="8C66DE8AFFB3FF853DB5FDA0FA81FC9D" pageId="16" pageNumber="54">
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<paragraph id="372DC62FFFB3FF853DB5FDA0FA81FDA2" blockId="16.[189,1399,572,839]" pageId="16" pageNumber="54">
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<keyLead id="8C63651AFFB3FF853DB5FDA0FA81FDA2" pageId="16" pageNumber="54">1. Main colouration pattern of adults chiefly or most often composed of a dominant continuous midlateral band, in the latter case sometimes in combination with vertical bars ............................. 2</keyLead>
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</paragraph>
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<paragraph id="372DC62FFFB3FF853DB5FD1EFA81FD64" blockId="16.[189,1399,572,839]" pageId="16" pageNumber="54">
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<keyLead id="8C63651AFFB3FF853DB5FD1EFA81FD64" pageId="16" pageNumber="54">– Main colouration pattern of adults chiefly composed of midlateral blotches along the body side without any continuous midlateral band ........................................................................................... 5</keyLead>
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</paragraph>
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<paragraph id="372DC62FFFB3FF853DC9FD5BFA81FC9D" blockId="16.[189,1399,572,839]" pageId="16" pageNumber="54">
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<keyLead id="8C63651AFFB3FF853DC9FD5BFA81FC9D" pageId="16" pageNumber="54">
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– Main colouration pattern chiefly composed of paired vertical bars in young, and fragmented into dark blotches and spots all over the sides including side of belly in adults, midlateral band when present is continuous, endemic to the Acaray tributary, endemic to
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<collectingCountry id="4F8586BFFFB3FF853EF4FC97FB9FFCFF" box="[1020,1129,779,805]" name="Paraguay" pageId="16" pageNumber="54">Paraguay</collectingCountry>
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................................... ............................................................................................
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<taxonomicName id="F092BDACFFB3FF853E9FFCB1FA81FC9D" ID-CoL="6BCWQ" authority="Kullander & Lucena, 2013" authorityName="Kullander & Lucena" authorityYear="2013" box="[919,1399,812,839]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="gillmorlisi">
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<emphasis id="05E61A3DFFB3FF853E9FFCB1FBC3FC9D" bold="true" box="[919,1077,812,839]" italics="true" pageId="16" pageNumber="54">C. gillmorlisi</emphasis>
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<bibRefCitation id="5303BBDEFFB3FF853934FCB1FA81FC9D" author="Kullander S. O. & Lucena C. A. S." box="[1084,1399,813,839]" pageId="16" pageNumber="54" pagination="149 - 164" refId="ref14189" refString="Kullander S. O. & Lucena C. A. S. 2013. Crenicichla gillmforlisi, a new species of cichlid fish (Teleostei: Cichlidae) from the Parana river drainage in Paraguay. Zootaxa 3641 (2): 149 - 164. https: // doi. org / 10.11646 / zootaxa. 3641.2.3" type="journal article" year="2013">Kullander & Lucena, 2013</bibRefCitation>
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</taxonomicName>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="8C66DE8AFFB3FF853DB5FCF2FA81FBC8" pageId="16" pageNumber="54">
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<paragraph id="372DC62FFFB3FF853DB5FCF2FA81FC71" blockId="16.[189,1399,878,1043]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DB5FCF2FA81FC71" pageId="16" pageNumber="54">2. The midlateral band is always present as a dominant colouration pattern in all growth stages, moods and under other conditions, scales in E1 row 50–64 ........................................................................ 3</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB3FF853DC9FC28FA81FBC8" blockId="16.[189,1399,878,1043]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DC9FC28FA81FBC8" pageId="16" pageNumber="54">– Depending on growth stage, mood and other factors the dominant body colouration can change from a midlateral band to vertical bars, or be a combination of the two, scales in E1 row either 42–57 or 78–85 ................................................................................................................................................ 4</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB3FF853DB5FBA4FA81FB01" pageId="16" pageNumber="54">
|
||
<paragraph id="372DC62FFFB3FF853DB5FBA4FA81FBAF" blockId="16.[189,1399,1080,1245]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DB5FBA4FA81FBAF" pageId="16" pageNumber="54">
|
||
3. Breeding females with a distinct dark white-margined blotch in the dorsal fin, scales in E1 row 50–55, endemic to the Piray Guazú tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB3FF853E80FBC6FC0BFBAE" box="[904,1021,1114,1140]" name="Argentina" pageId="16" pageNumber="54">Argentina</collectingCountry>
|
||
...................
|
||
<taxonomicName id="F092BDACFFB3FF85399EFBC6FAE3FBAF" authority="Říčan & Piálek & Almirón & Casciotta, 2023" authorityName="Říčan & Piálek & Almirón & Casciotta" authorityYear="2023" box="[1174,1301,1114,1141]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="aravera" status="sp. nov.">
|
||
<emphasis id="05E61A3DFFB3FF85399EFBC6FAE3FBAF" bold="true" box="[1174,1301,1114,1141]" italics="true" pageId="16" pageNumber="54">C. aravera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED5A746FFB3FF853814FBC7FA81FBAF" box="[1308,1399,1115,1141]" pageId="16" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB3FF853DC9FBE1FA81FB01" blockId="16.[189,1399,1080,1245]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DC9FBE1FA81FB01" pageId="16" pageNumber="54">
|
||
– Breeding females with a distinct dark white-margined long band in the dorsal fin, scales in E1 row 53–64, endemic to the Urugua-í tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB3FF853E69FB03FC20FB63" box="[865,982,1183,1209]" name="Argentina" pageId="16" pageNumber="54">Argentina</collectingCountry>
|
||
...................................................... .............................................................................
|
||
<taxonomicName id="F092BDACFFB3FF853E2EFB5DFA81FB01" authority="Pialek, Casciotta, Almiron & Rican, 2019" authorityName="Pialek, Casciotta, Almiron & Rican" authorityYear="2019" box="[806,1399,1216,1245]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="yjhui">
|
||
<emphasis id="05E61A3DFFB3FF853E2EFB5DFC70FB01" bold="true" box="[806,902,1216,1243]" italics="true" pageId="16" pageNumber="54">C. yjhui</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB3FF853E86FB5DFA81FB01" author="Pialek L. & Casciotta J. & Almiron A. & Rican O." box="[910,1399,1217,1245]" pageId="16" pageNumber="54" pagination="377 - 395" refId="ref14576" refString="Pialek L., Casciotta J., Almiron A. & Rican O. 2019 b. A new pelagic predatory pike cichlid (Teleostei: Cichlidae: Crenicichla) from the C. mandelburgeri species complex with parallel and reticulate evolution. Hydrobiologia 832: 377 - 395. https: // doi. org / 10.1007 / s 10750 - 018 - 3754 - 1" type="journal article" year="2019">Piálek, Casciotta, Almirón & Říčan, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB3FF853DB5FA9DFA81F9D0" pageId="16" pageNumber="54">
|
||
<paragraph id="372DC62FFFB3FF853DB5FA9DFA81FA58" blockId="16.[189,1401,1281,1547]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DB5FA9DFA81FA58" pageId="16" pageNumber="54">
|
||
4. Midlateral band when present always fully continuous, with barely discernible fluctuation in width, scales in E1 row 78–85, widespread in the Lower Paraná and entire
|
||
<collectingCountry id="4F8586BFFFB3FF853929FAB8FB78FAE4" box="[1057,1166,1316,1342]" name="Paraguay" pageId="16" pageNumber="54">Paraguay</collectingCountry>
|
||
Basin, also present in Lower
|
||
<collectingCountry id="4F8586BFFFB3FF853C6EFADAFE3BFABA" box="[358,461,1350,1376]" name="Uruguay" pageId="16" pageNumber="54">Uruguay</collectingCountry>
|
||
and lower portions of the Middle Paraná in
|
||
<collectingCountry id="4F8586BFFFB3FF853EF0FADAFB90FABA" box="[1016,1126,1350,1376]" name="Paraguay" pageId="16" pageNumber="54">Paraguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="4F8586BFFFB3FF85397DFAD9FB36FA85" box="[1141,1216,1349,1375]" name="Brazil" pageId="16" pageNumber="54">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="4F8586BFFFB3FF8539C6FAD9FAB5FA85" box="[1230,1347,1349,1375]" name="Argentina" pageId="16" pageNumber="54">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB3FF853DE2FAF4FEA7FA58" box="[234,337,1384,1410]" name="Uruguay" pageId="16" pageNumber="54">Uruguay</collectingCountry>
|
||
.........................................................................................................
|
||
<taxonomicName id="F092BDACFFB3FF85396EFAFBFA81FA58" ID-CoL="ZCHD" authority="Heckel, 1840" authorityName="Heckel" authorityYear="1840" box="[1126,1399,1383,1410]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="05E61A3DFFB3FF85396EFAFBFB25FA5B" bold="true" box="[1126,1235,1383,1409]" italics="true" pageId="16" pageNumber="54">C. vittata</emphasis>
|
||
Heckel, 1840
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB3FF853DC9FA16FA81F9D0" blockId="16.[189,1401,1281,1547]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DC9FA16FA81F9D0" pageId="16" pageNumber="54">
|
||
– Midlateral band often semicontinuous or even distrupted (i.e., always of clearly fluctuating width), vertical bars when present always as clear double-bars, scales in E1 row 42–57, endemic to Middle Paraná main River and some non-isolated tributaries in
|
||
<collectingCountry id="4F8586BFFFB3FF853E70FA53FC13FA33" box="[888,997,1487,1513]" name="Paraguay" pageId="16" pageNumber="54">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB3FF853914FA52FB67FA32" box="[1052,1169,1486,1512]" name="Argentina" pageId="16" pageNumber="54">Argentina</collectingCountry>
|
||
............................. ......................................................................................................
|
||
<taxonomicName id="F092BDACFFB3FF853ED4FA6CFA81F9D0" ID-CoL="ZCFK" authority="Kullander, 2009" authorityName="Kullander" authorityYear="2009" box="[988,1399,1520,1547]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="05E61A3DFFB3FF853ED4FA6CFB47F9D1" bold="true" box="[988,1201,1520,1547]" italics="true" pageId="16" pageNumber="54">C. mandelburgeri</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB3FF8539B1FA6CFA81F9D0" author="Kullander S. O." box="[1209,1399,1520,1546]" pageId="16" pageNumber="54" pagination="41 - 50" refId="ref14063" refString="Kullander S. O. 2009. Crenicichla mandelburgeri, a new species of cichlid fish (Teleostei: Cichlidae) from the Parana river drainage in Paraguay. Zootaxa 2006 (1): 41 - 50. https: // doi. org / 10.11646 / zootaxa. 2006.1.3" type="journal article" year="2009">Kullander, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB3FF853DB5F9ACFA81F96B" pageId="16" pageNumber="54">
|
||
<paragraph id="372DC62FFFB3FF853DB5F9ACFA81F955" blockId="16.[189,1399,1584,1713]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DB5F9ACFA81F955" pageId="16" pageNumber="54">
|
||
5. Breeding females with a distinct dark white-margined blotch in the dorsal fin, scattered spots on body below the midlateral band in both sexes, especially in males, endemic to the Piray Guazú tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB3FF853CEBF9E9FDAEF955" box="[483,600,1653,1679]" name="Argentina" pageId="16" pageNumber="54">Argentina</collectingCountry>
|
||
.................................................................................
|
||
<taxonomicName id="F092BDACFFB3FF8539B7F9E9FAE3F955" authority="Říčan & Piálek & Almirón & Casciotta, 2023" authorityName="Říčan & Piálek & Almirón & Casciotta" authorityYear="2023" box="[1215,1301,1653,1679]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="ama" status="sp. nov.">
|
||
<emphasis id="05E61A3DFFB3FF8539B7F9E9FAE3F955" bold="true" box="[1215,1301,1653,1679]" italics="true" pageId="16" pageNumber="54">C. ama</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED5A746FFB3FF853814F9E9FA81F955" box="[1308,1399,1653,1679]" pageId="16" pageNumber="54" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB3FF853DC9F90BFA81F96B" blockId="16.[189,1399,1584,1713]" box="[193,1399,1687,1713]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DC9F90BFA81F96B" box="[193,1399,1687,1713]" pageId="16" pageNumber="54">– Breeding females with a distinct dark white-margined long band in the dorsal fin ......................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB3FF853DB5F94BFA80F882" pageId="16" pageNumber="54">
|
||
<paragraph id="372DC62FFFB3FF853DB5F94BFA81F8CE" blockId="16.[189,1400,1751,1880]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DB5F94BFA81F8CE" pageId="16" pageNumber="54">
|
||
6. Scattered spots on body below the midlateral band in both sexes, especially in males, endemic to the Iguazú,
|
||
<collectingCountry id="4F8586BFFFB3FF853C4EF965FE4DF8C9" box="[326,443,1785,1811]" name="Argentina" pageId="16" pageNumber="54">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB3FF853CFCF965FDCAF8C9" box="[500,572,1785,1811]" name="Brazil" pageId="16" pageNumber="54">Brazil</collectingCountry>
|
||
............................................................................................................ 7
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB3FF853DC9F880FA80F882" blockId="16.[189,1400,1751,1880]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DC9F880FA80F882" pageId="16" pageNumber="54">
|
||
– Without spots on body below the midlateral band in both sexes, endemic to Middle
|
||
<collectingRegion id="F55608CDFFB3FF8539D3F880FADDF8EC" box="[1243,1323,1820,1846]" country="Brazil" name="Parana" pageId="16" pageNumber="54">Paraná</collectingRegion>
|
||
Basin apart from the Iguazú ...................................................................................................................... 10
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB3FF853DB5F8E3FA81F806" pageId="16" pageNumber="54">
|
||
<paragraph id="372DC62FFFB3FF853DB5F8E3FA81F860" blockId="16.[189,1399,1918,2013]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DB5F8E3FA81F860" pageId="16" pageNumber="54">
|
||
7. Oral jaws prognathous (i.e., lower jaw projecting in front of upper jaw), mouth large, corner of mouth reaching well below eye .......................................................
|
||
<taxonomicName id="F092BDACFFB3FF853EF6F83CFA81F860" ID-CoL="6BCWX" authority="Haseman, 1911" authorityName="Haseman" authorityYear="1911" box="[1022,1399,1952,1979]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="16" pageNumber="54" phylum="Chordata" rank="species" species="iguassuensis">
|
||
<emphasis id="05E61A3DFFB3FF853EF6F83CFB4FF860" bold="true" box="[1022,1209,1952,1978]" italics="true" pageId="16" pageNumber="54">C. iguassuensis</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB3FF8539C9F83DFA81F860" author="Haseman J. D." box="[1217,1399,1952,1979]" pageId="16" pageNumber="54" pagination="329 - 373" refId="ref13834" refString="Haseman J. D. 1911. An annotated catalog of the cichlid fishes collected by the expedition of the Carnegie Museum to central South America, 1907 - 10. Annals of the Carnegie Museum 7: 329 - 373. Available from https: // www. biodiversitylibrary. org / page / 9956994 [accessed 2 Jun. 2023]." type="journal article" year="1911">Haseman, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB3FF853DC9F85FFA81F806" blockId="16.[189,1399,1918,2013]" box="[193,1399,1986,2013]" pageId="16" pageNumber="54">
|
||
<keyLead id="8C63651AFFB3FF853DC9F85FFA81F806" box="[193,1399,1986,2013]" pageId="16" pageNumber="54">– Oral jaws isognathous or hypognathous (i.e., upper jaw projecting in front of lower jaw) ............. 8</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB2FF843DB5FE95FA81FEB3" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5FE95FA81FE9D" blockId="17.[189,1399,265,362]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5FE95FA81FE9D" pageId="17" pageNumber="55">
|
||
8. Mouth large, corner of mouth reaching below vertical from anterior margin of eye, lips very thick and bulbous ..........................................
|
||
<taxonomicName id="F092BDACFFB2FF843FB4FEB0FA81FE9D" ID-CoL="ZCH9" authority="Pialek, Dragova, Casciotta, Almiron & Rican, 2015" authorityName="Pialek, Dragova, Casciotta, Almiron & Rican" authorityYear="2015" box="[700,1399,300,329]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="tuca">
|
||
<emphasis id="05E61A3DFFB2FF843FB4FEB0FCE4FE9C" bold="true" box="[700,786,300,326]" italics="true" pageId="17" pageNumber="55">C. tuca</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843E12FEB0FA81FE9D" author="Pialek L. & Dragova K. & Casciotta J. & Almiron A. & Rican O." box="[794,1399,300,329]" pageId="17" pageNumber="55" pagination="5 - 27" refId="ref14426" refString="Pialek L., Dragova K., Casciotta J., Almiron A. & Rican O. 2015. Description of two new species of Crenicichla (Teleostei: Cichlidae) from the Lower Iguazu River with a taxonomic reappraisal of C. iguassuensis, C. tesay and C. yaha. Historia Natural (Tercera Serie) 5: 5 - 27." type="journal article" year="2015">Piálek, Dragová, Casciotta, Almirón & Říčan, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9FECCFA81FEB3" blockId="17.[189,1399,265,362]" box="[193,1399,335,362]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9FECCFA81FEB3" box="[193,1399,335,362]" pageId="17" pageNumber="55">– Mouth small, not reaching eye, lips normal ..................................................................................... 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB2FF843DB5FE0DFA81FD7B" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5FE0DFA81FDE2" blockId="17.[189,1399,401,674]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5FE0DFA81FDE2" pageId="17" pageNumber="55">
|
||
9. Well-developed vertical double-bars (only species in the Iguazú with this character), anterior one or two midlateral blotches much more dominant over the remaining, which are usually only visible as connections between the double bars, a well-formed (not decomposed into spots) narrow suborbital stripe inclined significantly posteriad, near absence of dark scattered dots on flanks in males and complete absence in females ...............
|
||
<taxonomicName id="F092BDACFFB2FF843FB3FD81FA81FDE2" ID-CoL="ZCH3" authority="Pialek, Dragova, Casciotta, Almiron & Rican, 2015" authorityName="Pialek, Dragova, Casciotta, Almiron & Rican" authorityYear="2015" box="[699,1399,541,570]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="tapii">
|
||
<emphasis id="05E61A3DFFB2FF843FB3FD81FCE5FDED" bold="true" box="[699,787,541,567]" italics="true" pageId="17" pageNumber="55">C. tapii</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843E12FD82FA81FDE2" author="Pialek L. & Dragova K. & Casciotta J. & Almiron A. & Rican O." box="[794,1399,541,570]" pageId="17" pageNumber="55" pagination="5 - 27" refId="ref14426" refString="Pialek L., Dragova K., Casciotta J., Almiron A. & Rican O. 2015. Description of two new species of Crenicichla (Teleostei: Cichlidae) from the Lower Iguazu River with a taxonomic reappraisal of C. iguassuensis, C. tesay and C. yaha. Historia Natural (Tercera Serie) 5: 5 - 27." type="journal article" year="2015">Piálek, Dragová, Casciotta, Almirón & Říčan, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9FDDDFA81FD7B" blockId="17.[189,1399,401,674]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9FDDDFA81FD7B" pageId="17" pageNumber="55">
|
||
– Vertical bars and less developed double bars only visible above midlateral line, anterior four or more midlateral blotches well dominant, suborbital stripe decomposed into spots, wider, and less inclined posteriad, both sexes with dark scattered dots on flanks ...........
|
||
<taxonomicName id="F092BDACFFB2FF843EDBFD1BFA81FD7B" ID-CoL="ZCH6" authority="Casciotta & Almiron, 2008" authorityName="Casciotta & Almiron" authorityYear="2008" box="[979,1399,647,674]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="05E61A3DFFB2FF843EDBFD1BFBC7FD78" bold="true" box="[979,1073,647,674]" italics="true" pageId="17" pageNumber="55">C. tesay</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843931FD1BFA81FD7B" author="Casciotta J. & Almiron A." box="[1081,1399,647,673]" pageId="17" pageNumber="55" pagination="1 - 36" refId="ref13471" refString="Casciotta J. & Almiron A. 2008. Crenicichla tesay, a new species of cichlid (Perciformes: Labroidei) from the rio Iguazu basin in Argentina. Revue suisse de Zoologie 115: 651 - 659. https: // doi. org / 10.5962 / bhl. part. 80451 Casciotta J. R. & Arratia G. 1993. Jaws and teeth of American Cichlids (Pisces: Labroidei). Journal of Morphology 217: 1 - 36. https: // doi. org / 10.1002 / jmor. 1052170102" type="journal article" year="2008">Casciotta & Almirón, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB2FF843DB5FD55FA80FC96" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5FD55FA80FCDC" blockId="17.[189,1399,713,845]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5FD55FA80FCDC" pageId="17" pageNumber="55">10. Oral jaws prognathous (i.e., lower jaw projecting in front of upper jaw), mouth rather large (i.e., corner of mouth reaching below vertical from anterior margin of eye) ..........................................11</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9FC8CFA80FC96" blockId="17.[189,1399,713,845]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9FC8CFA80FC96" pageId="17" pageNumber="55">– Oral jaws isognathous or hypognathous (i.e., upper jaw projecting in front of lower jaw), mouth rather small (i.e., not reaching eyes) ............................................................................................... 12</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB2FF843DB5FCE8FA81FB34" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5FCE8FA81FBE4" blockId="17.[189,1401,884,1264]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5FCE8FA81FBE4" pageId="17" pageNumber="55">
|
||
11. Anterior two to three midlateral blotches dominant, background colouration light grey, midlateral blotches distinctly H-like with dorsally continuing double bars, females with a wide wide orange to red stripe along the entire length of the basal portion of the dorsal fin above which is found a shorter black band margined in white, suborbital stripe of normal length and width, composed of spots, which do not form horizontal lines, endemic to the Urugua-í tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB2FF8439A6FB9CFAD5FBC0" box="[1198,1315,1024,1050]" name="Argentina" pageId="17" pageNumber="55">Argentina</collectingCountry>
|
||
......... ..................................................................
|
||
<taxonomicName id="F092BDACFFB2FF843FDEFBB8FA81FBE4" ID-CoL="ZCHH" authority="Casciotta, Almiron, Pialek, Gomez & Rican, 2010" authorityName="Casciotta, Almiron, Pialek, Gomez & Rican" authorityYear="2010" box="[726,1399,1060,1088]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="ypo">
|
||
<emphasis id="05E61A3DFFB2FF843FDEFBB8FCD5FBE4" bold="true" box="[726,803,1060,1086]" italics="true" pageId="17" pageNumber="55">C. ypo</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843E22FBB8FA81FBE4" author="Casciotta J. & Almiron A. & Pialek L. & Gomez S. & Rican O." box="[810,1399,1060,1088]" pageId="17" pageNumber="55" pagination="643 - 648" refId="ref13623" refString="Casciotta J., Almiron A., Pialek L., Gomez S. & Rican O. 2010. Crenicichla ypo (Teleostei: Cichlidae), a new species from the middle Parana basin in Misiones, Argentina. Neotropical Ichthyology 8: 643 - 648. https: // doi. org / 10.1590 / S 1679 - 62252010000300009" type="journal article" year="2010">Casciotta, Almirón, Piálek, Gómez & Říčan, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9FBDBFA81FB34" blockId="17.[189,1401,884,1264]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9FBDBFA81FB34" pageId="17" pageNumber="55">
|
||
– All midlateral blotches well developed, background colouration very dark grey to blackish, midlateral blotches square-like with dorsally continuing single wide bars, females only with the shorter black band margined in white in the dorsal fin without the orange to red band, suborbital stripe short and wide, ventrally widest, composed of spots which do form horizontal lines, endemic to the Piray Miní tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB2FF843CEBFB48FDAEFB34" box="[483,600,1236,1262]" name="Argentina" pageId="17" pageNumber="55">Argentina</collectingCountry>
|
||
..............................
|
||
<emphasis id="05E61A3DFFB2FF843E4BFB48FC71FB34" bold="true" box="[835,903,1235,1262]" italics="true" pageId="17" pageNumber="55">C. hu</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843E86FB48FA81FB34" author="Casciotta J. & Almiron A. & Pialek L. & Gomez S. & Rican O." box="[910,1399,1236,1264]" pageId="17" pageNumber="55" pagination="643 - 648" refId="ref13623" refString="Casciotta J., Almiron A., Pialek L., Gomez S. & Rican O. 2010. Crenicichla ypo (Teleostei: Cichlidae), a new species from the middle Parana basin in Misiones, Argentina. Neotropical Ichthyology 8: 643 - 648. https: // doi. org / 10.1590 / S 1679 - 62252010000300009" type="journal article" year="2010">Piálek, Říčan, Casciotta & Almirón, 2010</bibRefCitation>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB2FF843DB5FA89FA81F993" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5FA89FA81FA43" blockId="17.[189,1399,1301,1611]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5FA89FA81FA43" pageId="17" pageNumber="55">
|
||
12. Anterior two to three midlateral blotches usually more dominant, midlateral blotches distinctly H-like with dorsally continuing double bars, suborbital stripe shorter and wider, composed of spots which tend to form horizontal lines, scales in E1 row 48–51 (mean 50), endemic to the Urugua-í tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB2FF843CEBFAE3FDAEFA43" box="[483,600,1407,1433]" name="Argentina" pageId="17" pageNumber="55">Argentina</collectingCountry>
|
||
....................................
|
||
<taxonomicName id="F092BDACFFB2FF843E79FAE3FA81FA43" ID-CoL="ZCHG" authority="Casciotta, Almiron & Gomez, 2006" authorityName="Casciotta, Almiron & Gomez" authorityYear="2006" box="[881,1399,1406,1433]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="05E61A3DFFB2FF843E79FAE3FC38FA43" bold="true" box="[881,974,1406,1433]" italics="true" pageId="17" pageNumber="55">C. yaha</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843EDDFAE3FA81FA43" author="Casciotta J. & Almiron A. E. & Gomez S. E." box="[981,1399,1407,1433]" pageId="17" pageNumber="55" pagination="107 - 112" refId="ref13569" refString="Casciotta J., Almiron A. E. & Gomez S. E. 2006. Crenicichla yaha sp. n. (Perciformes: Labroidei: Cichlidae), a new species from the rio Iguazu and arroyo Urugua-i basins, northeastern Argentina. Zoologische Abhandlungen 56: 107 - 112." type="journal article" year="2006">Casciotta, Almirón & Gómez, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9FA3EFA81F993" blockId="17.[189,1399,1301,1611]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9FA3EFA81F993" pageId="17" pageNumber="55">
|
||
– All midlateral blotches equally developed, midlateral blotches square-like with dorsally continuing single wide bars, suborbital stripe of normal length and width, composed of spots which tend to form horizontal lines, scales in E1 row 54–60 (mean 56–57), endemic to the lower portion of the Middle Paraná and some non-isolated tributaries in
|
||
<collectingCountry id="4F8586BFFFB2FF843FE5F990FCACF9FC" box="[749,858,1548,1574]" name="Paraguay" pageId="17" pageNumber="55">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB2FF843E9AF997FBF1F9FF" box="[914,1031,1547,1573]" name="Argentina" pageId="17" pageNumber="55">Argentina</collectingCountry>
|
||
................................................ .............................................
|
||
<taxonomicName id="F092BDACFFB2FF843F48F9B3FA81F993" ID-CoL="ZCH2" authority="Casciotta, Almiron, Aichino, Gomez, Pialek & Rican, 2013" authorityName="Casciotta, Almiron, Aichino, Gomez, Pialek & Rican" authorityYear="2013" box="[576,1399,1582,1611]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="taikyra">
|
||
<emphasis id="05E61A3DFFB2FF843F48F9B3FD4EF993" bold="true" box="[576,696,1582,1609]" italics="true" pageId="17" pageNumber="55">C. taikyra</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843FB7F9B3FA81F993" author="Casciotta J. & Almiron A. & Aichino D. & Gomez S. & Pialek L. & Rican O." box="[703,1399,1583,1611]" pageId="17" pageNumber="55" pagination="379 - 386" refId="ref13686" refString="Casciotta J., Almiron A., Aichino D., Gomez S., Pialek L. & Rican O. 2013. Crenicichla taikyra (Teleostei: Cichlidae), a new species of pike cichlid from the middle rio Parana, Argentina. Zootaxa 3721 (4): 379 - 386. https: // doi. org / 10.11646 / zootaxa. 3721.4.5" type="journal article" year="2013">Casciotta, Almirón, Aichino, Gómez, Piálek & Říčan, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
<subSubSection id="7F8895A4FFB2FF843DB5F9ECFC36F977" pageId="17" pageNumber="55" type="multiple">
|
||
<paragraph id="372DC62FFFB2FF843DB5F9ECFC36F977" blockId="17.[189,1396,1647,1709]" pageId="17" pageNumber="55">
|
||
<emphasis id="05E61A3DFFB2FF843DB5F9ECFC36F977" bold="true" pageId="17" pageNumber="55">
|
||
Key to the species of
|
||
<taxonomicName id="F092BDACFFB2FF843CB0F9ECFDB0F951" authorityName="Heckel" authorityYear="1840" box="[440,582,1648,1675]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="genus">
|
||
<emphasis id="05E61A3DFFB2FF843CB0F9ECFDB0F951" bold="true" box="[440,582,1648,1675]" italics="true" pageId="17" pageNumber="55">Crenicichla</emphasis>
|
||
</taxonomicName>
|
||
from the Middle Rio
|
||
<collectingRegion id="F55608CDFFB2FF843E45F9F3FC5EF950" box="[845,936,1647,1674]" country="Brazil" name="Parana" pageId="17" pageNumber="55">Paraná</collectingRegion>
|
||
Basin (the
|
||
<taxonomicName id="F092BDACFFB2FF843926F9ECFA82F951" authorityName="Kullander" authorityYear="2009" box="[1070,1396,1648,1675]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="05E61A3DFFB2FF843926F9ECFA82F951" bold="true" box="[1070,1396,1648,1675]" italics="true" pageId="17" pageNumber="55">Crenicichla mandelburgeri</emphasis>
|
||
</taxonomicName>
|
||
<heading id="6C657143FFB2FF843DB5F90FFC36F977" bold="true" box="[189,960,1683,1709]" fontSize="11" level="3" pageId="17" pageNumber="55" reason="3">
|
||
species group plus
|
||
<taxonomicName id="F092BDACFFB2FF843CA0F90FFDE3F977" authorityName="Heckel" authorityYear="1840" box="[424,533,1683,1709]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="05E61A3DFFB2FF843CA0F90FFDE3F977" bold="true" box="[424,533,1683,1709]" italics="true" pageId="17" pageNumber="55">C. vittata</emphasis>
|
||
</taxonomicName>
|
||
) based on oral jaw characteristics
|
||
</heading>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="7F8895A4FFB2FF863DB5F95EFA81FC0B" lastPageId="19" lastPageNumber="57" pageId="17" pageNumber="55" type="key">
|
||
<key id="4303911EFFB2FF863DB5F95EFA81FC0B" lastPageId="19" lastPageNumber="57" pageId="17" pageNumber="55">
|
||
<keyStep id="8C66DE8AFFB2FF843DB5F95EFA81F89F" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5F95EFA81F925" blockId="17.[189,1399,1730,1862]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5F95EFA81F925" pageId="17" pageNumber="55">1. Oral jaws isognathous or hypognathous (i.e., upper jaw projecting in front of lower jaw), mouth generally rather small to small (i.e., not reaching eyes), except in one species where quite large ... 2</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9F895FA81F89F" blockId="17.[189,1399,1730,1862]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9F895FA81F89F" pageId="17" pageNumber="55">– Oral jaws prognathous (i.e., lower jaw projecting in front of upper jaw), mouth rather large to large (i.e., corner of mouth reaching below vertical from anterior margin of eye) ................................... 6</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB2FF843DB5F8F1FA81F817" pageId="17" pageNumber="55">
|
||
<paragraph id="372DC62FFFB2FF843DB5F8F1FA81F871" blockId="17.[189,1399,1901,1998]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DB5F8F1FA81F871" pageId="17" pageNumber="55">
|
||
2. Mouth large, corner of mouth reaching below vertical from anterior margin of eye, lips very thick and bulbous ..........................................
|
||
<taxonomicName id="F092BDACFFB2FF843FB4F80CFA81F871" ID-CoL="ZCH9" authority="Pialek, Dragova, Casciotta, Almiron & Rican, 2015" authorityName="Pialek, Dragova, Casciotta, Almiron & Rican" authorityYear="2015" box="[700,1399,1936,1965]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="17" pageNumber="55" phylum="Chordata" rank="species" species="tuca">
|
||
<emphasis id="05E61A3DFFB2FF843FB4F80CFCE4F870" bold="true" box="[700,786,1936,1962]" italics="true" pageId="17" pageNumber="55">C. tuca</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB2FF843E12F80CFA81F871" author="Pialek L. & Dragova K. & Casciotta J. & Almiron A. & Rican O." box="[794,1399,1936,1965]" pageId="17" pageNumber="55" pagination="5 - 27" refId="ref14426" refString="Pialek L., Dragova K., Casciotta J., Almiron A. & Rican O. 2015. Description of two new species of Crenicichla (Teleostei: Cichlidae) from the Lower Iguazu River with a taxonomic reappraisal of C. iguassuensis, C. tesay and C. yaha. Historia Natural (Tercera Serie) 5: 5 - 27." type="journal article" year="2015">Piálek, Dragová, Casciotta, Almirón & Říčan, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB2FF843DC9F828FA81F817" blockId="17.[189,1399,1901,1998]" box="[193,1399,1971,1998]" pageId="17" pageNumber="55">
|
||
<keyLead id="8C63651AFFB2FF843DC9F828FA81F817" box="[193,1399,1971,1998]" pageId="17" pageNumber="55">– Mouth small, not reaching eye, lips normal ..................................................................................... 3</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5FE95FA81FE53" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5FE95FA81FE9F" blockId="18.[189,1399,265,393]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5FE95FA81FE9F" pageId="18" pageNumber="56">
|
||
3. Scattered spots on body below the midlateral band in both sexes, especially in males (except for one species where weakly developed), endemic to the Iguazú,
|
||
<collectingCountry id="4F8586BFFFB1FF873E9BFEB6FBFEFE9E" box="[915,1032,298,324]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB1FF873948FEB6FB7EFE9E" box="[1088,1160,298,324]" name="Brazil" pageId="18" pageNumber="56">Brazil</collectingCountry>
|
||
............................ 4
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9FED1FA81FE53" blockId="18.[189,1399,265,393]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9FED1FA81FE53" pageId="18" pageNumber="56">
|
||
– Without spots on body below the midlateral band in both sexes, endemic to Middle
|
||
<collectingRegion id="F55608CDFFB1FF8739D3FED0FADDFEBC" box="[1243,1323,332,358]" country="Brazil" name="Parana" pageId="18" pageNumber="56">Paraná</collectingRegion>
|
||
Basin apart from the Iguazú ........................................................................................................................ 5
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5FE32FA81FD69" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5FE32FA81FD95" blockId="18.[189,1399,430,691]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5FE32FA81FD95" pageId="18" pageNumber="56">
|
||
4. Well-developed vertical double-bars (only species in the Iguazú with this character), anterior one or two midlateral blotches much more dominant over the remaining, which are usually only visible as connections between the double bars, a well-formed (not decomposed into spots) narrow suborbital stripe inclined significantly posteriad, near absence of dark scattered dots on flanks in males and complete absence in females ...............
|
||
<taxonomicName id="F092BDACFFB1FF873FB3FDA8FA81FD95" ID-CoL="ZCH3" authority="Pialek, Dragova, Casciotta, Almiron & Rican, 2015" authorityName="Pialek, Dragova, Casciotta, Almiron & Rican" authorityYear="2015" box="[699,1399,564,593]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="tapii">
|
||
<emphasis id="05E61A3DFFB1FF873FB3FDA8FCE5FD94" bold="true" box="[699,787,564,590]" italics="true" pageId="18" pageNumber="56">C. tapii</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF873E12FDA8FA81FD95" author="Pialek L. & Dragova K. & Casciotta J. & Almiron A. & Rican O." box="[794,1399,564,593]" pageId="18" pageNumber="56" pagination="5 - 27" refId="ref14426" refString="Pialek L., Dragova K., Casciotta J., Almiron A. & Rican O. 2015. Description of two new species of Crenicichla (Teleostei: Cichlidae) from the Lower Iguazu River with a taxonomic reappraisal of C. iguassuensis, C. tesay and C. yaha. Historia Natural (Tercera Serie) 5: 5 - 27." type="journal article" year="2015">Piálek, Dragová, Casciotta, Almirón & Říčan, 2015</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9FDCAFA81FD69" blockId="18.[189,1399,430,691]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9FDCAFA81FD69" pageId="18" pageNumber="56">
|
||
– Vertical bars and less developed double bars only visible above midlateral line, anterior four or more midlateral blotches well dominant, suborbital stripe decomposed into spots, wider, and less inclined posteriad, both sexes with dark scattered dots on flanks ...........
|
||
<taxonomicName id="F092BDACFFB1FF873EDBFD05FA81FD69" ID-CoL="ZCH6" authority="Casciotta & Almiron, 2008" authorityName="Casciotta & Almiron" authorityYear="2008" box="[979,1399,665,691]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="tesay">
|
||
<emphasis id="05E61A3DFFB1FF873EDBFD05FBC7FD69" bold="true" box="[979,1073,665,691]" italics="true" pageId="18" pageNumber="56">C. tesay</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF873931FD05FA81FD69" author="Casciotta J. & Almiron A." box="[1081,1399,665,691]" pageId="18" pageNumber="56" pagination="1 - 36" refId="ref13471" refString="Casciotta J. & Almiron A. 2008. Crenicichla tesay, a new species of cichlid (Perciformes: Labroidei) from the rio Iguazu basin in Argentina. Revue suisse de Zoologie 115: 651 - 659. https: // doi. org / 10.5962 / bhl. part. 80451 Casciotta J. R. & Arratia G. 1993. Jaws and teeth of American Cichlids (Pisces: Labroidei). Journal of Morphology 217: 1 - 36. https: // doi. org / 10.1002 / jmor. 1052170102" type="journal article" year="2008">Casciotta & Almirón, 2008</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5FD44FA81FBDA" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5FD44FA81FC8D" blockId="18.[189,1399,728,1026]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5FD44FA81FC8D" pageId="18" pageNumber="56">
|
||
5. Anterior two to three midlateral blotches usually more dominant, midlateral blotches distinctly H-like with dorsally continuing double bars, suborbital stripe shorter and wider, composed of spots which tend to form horizontal lines, scales in E1 row 48–51 (mean 50), endemic to the Urugua-í tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB1FF873CEBFCA1FDAEFC8D" box="[483,600,829,855]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
....................................
|
||
<taxonomicName id="F092BDACFFB1FF873E79FCA1FA81FC8D" ID-CoL="ZCHG" authority="Casciotta, Almiron & Gomez, 2006" authorityName="Casciotta, Almiron & Gomez" authorityYear="2006" box="[881,1399,829,856]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="yaha">
|
||
<emphasis id="05E61A3DFFB1FF873E79FCA1FC38FC82" bold="true" box="[881,974,829,856]" italics="true" pageId="18" pageNumber="56">C. yaha</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF873EDDFCA1FA81FC8D" author="Casciotta J. & Almiron A. E. & Gomez S. E." box="[981,1399,829,856]" pageId="18" pageNumber="56" pagination="107 - 112" refId="ref13569" refString="Casciotta J., Almiron A. E. & Gomez S. E. 2006. Crenicichla yaha sp. n. (Perciformes: Labroidei: Cichlidae), a new species from the rio Iguazu and arroyo Urugua-i basins, northeastern Argentina. Zoologische Abhandlungen 56: 107 - 112." type="journal article" year="2006">Casciotta, Almirón & Gómez, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9FCFCFA81FBDA" blockId="18.[189,1399,728,1026]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9FCFCFA81FBDA" pageId="18" pageNumber="56">
|
||
– All midlateral blotches equally developed, midlateral blotches square-like with dorsally continuing single wide bars, suborbital stripe of normal length and width, composed of spots which tend to form horizontal lines, scales in E1 row 54–60 (mean 56–57), endemic to the lower portion of the Middle Paraná and some non-isolated tributaries in
|
||
<collectingCountry id="4F8586BFFFB1FF873FE5FC58FCACFC04" box="[749,858,964,990]" name="Paraguay" pageId="18" pageNumber="56">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB1FF873E9AFC58FBF1FC04" box="[914,1031,964,990]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
................................................ .............................................
|
||
<taxonomicName id="F092BDACFFB1FF873F48FC79FA81FBDA" ID-CoL="ZCH2" authority="Casciotta, Almiron, Aichino, Gomez, Pialek & Rican, 2013" authorityName="Casciotta, Almiron, Aichino, Gomez, Pialek & Rican" authorityYear="2013" box="[576,1399,997,1026]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="taikyra">
|
||
<emphasis id="05E61A3DFFB1FF873F48FC79FD4EFBDA" bold="true" box="[576,696,997,1024]" italics="true" pageId="18" pageNumber="56">C. taikyra</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF873FB7FC79FA81FBDA" author="Casciotta J. & Almiron A. & Aichino D. & Gomez S. & Pialek L. & Rican O." box="[703,1399,997,1026]" pageId="18" pageNumber="56" pagination="379 - 386" refId="ref13686" refString="Casciotta J., Almiron A., Aichino D., Gomez S., Pialek L. & Rican O. 2013. Crenicichla taikyra (Teleostei: Cichlidae), a new species of pike cichlid from the middle rio Parana, Argentina. Zootaxa 3721 (4): 379 - 386. https: // doi. org / 10.11646 / zootaxa. 3721.4.5" type="journal article" year="2013">Casciotta, Almirón, Aichino, Gómez, Piálek & Říčan, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5FBB9FA81FAF1" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5FBB9FA81FBBB" blockId="18.[189,1399,1061,1323]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5FBB9FA81FBBB" pageId="18" pageNumber="56">6. Main colouration pattern of adults chiefly or most often composed of a dominant continuous midlateral band, in the latter case sometimes in combination with vertical bars ............................. 7</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DB5FBF5FA80FB7E" blockId="18.[189,1399,1061,1323]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5FBF5FA80FB7E" pageId="18" pageNumber="56">– Main colouration pattern of adults chiefly composed of midlateral blotches along the body side without any continuous midlateral band ..........................................................................................11</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9FB30FA81FAF1" blockId="18.[189,1399,1061,1323]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9FB30FA81FAF1" pageId="18" pageNumber="56">
|
||
– Main colouration pattern chiefly composed of paired vertical bars in young, and fragmented into dark blotches and spots all over the sides including side of belly in adults, midlateral band when present is continuous, endemic to the Acaray tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB1FF873EF4FB73FB9FFAD3" box="[1020,1129,1263,1289]" name="Paraguay" pageId="18" pageNumber="56">Paraguay</collectingCountry>
|
||
................................... ............................................................................................
|
||
<taxonomicName id="F092BDACFFB1FF873E9FFA8CFA81FAF1" ID-CoL="6BCWQ" authority="Kullander & Lucena, 2013" authorityName="Kullander & Lucena" authorityYear="2013" box="[919,1399,1296,1323]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="gillmorlisi">
|
||
<emphasis id="05E61A3DFFB1FF873E9FFA8CFBC3FAF1" bold="true" box="[919,1077,1296,1323]" italics="true" pageId="18" pageNumber="56">C. gillmorlisi</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF873934FA8CFA81FAF1" author="Kullander S. O. & Lucena C. A. S." box="[1084,1399,1296,1323]" pageId="18" pageNumber="56" pagination="149 - 164" refId="ref14189" refString="Kullander S. O. & Lucena C. A. S. 2013. Crenicichla gillmforlisi, a new species of cichlid fish (Teleostei: Cichlidae) from the Parana river drainage in Paraguay. Zootaxa 3641 (2): 149 - 164. https: // doi. org / 10.11646 / zootaxa. 3641.2.3" type="journal article" year="2013">Kullander & Lucena, 2013</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5FACDFA81FA2B" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5FACDFA81FA56" blockId="18.[189,1399,1360,1521]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5FACDFA81FA56" pageId="18" pageNumber="56">7. The midlateral band is always present as a dominant colouration pattern in all growth stages, moods and under other conditions, scales in E1 row 50–64 ........................................................................ 8</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9FA08FA81FA2B" blockId="18.[189,1399,1360,1521]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9FA08FA81FA2B" pageId="18" pageNumber="56">– Depending on growth stage, mood and other factors the dominant body colouration can change from a midlateral band to vertical bars, or be a combination of the two, scales in E1 row either 42–57 or 78–85 ................................................................................................................................................ 9</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5F98BFA81F96D" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5F98BFA81F989" blockId="18.[189,1399,1558,1721]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5F98BFA81F989" pageId="18" pageNumber="56">
|
||
8. Breeding females with a distinct dark white-margined blotch in the dorsal fin, scales in E1 row 50–55, endemic to the Piray Guazú tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB1FF873E80F9A4FC0BF988" box="[904,1021,1592,1618]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
...................
|
||
<taxonomicName id="F092BDACFFB1FF87399EF9A4FAE3F988" authority="Říčan & Piálek & Almirón & Casciotta, 2023" authorityName="Říčan & Piálek & Almirón & Casciotta" authorityYear="2023" box="[1174,1301,1592,1618]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="aravera" status="sp. nov.">
|
||
<emphasis id="05E61A3DFFB1FF87399EF9A4FAE3F988" bold="true" box="[1174,1301,1592,1618]" italics="true" pageId="18" pageNumber="56">C. aravera</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED5A746FFB1FF873814F9A5FA81F989" box="[1308,1399,1593,1619]" pageId="18" pageNumber="56" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9F9C6FA81F96D" blockId="18.[189,1399,1558,1721]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9F9C6FA81F96D" pageId="18" pageNumber="56">
|
||
– Breeding females with a distinct dark white-margined long band in the dorsal fin, scales in E1 row 53–64, endemic to the Urugua-í tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB1FF873E69F9E7FC20F94F" box="[865,982,1659,1685]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
...................................................... .............................................................................
|
||
<taxonomicName id="F092BDACFFB1FF873E2EF901FA81F96D" authority="Pialek, Casciotta, Almiron & Rican, 2019" authorityName="Pialek, Casciotta, Almiron & Rican" authorityYear="2019" box="[806,1399,1692,1721]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="yjhui">
|
||
<emphasis id="05E61A3DFFB1FF873E2EF901FC70F96D" bold="true" box="[806,902,1692,1719]" italics="true" pageId="18" pageNumber="56">C. yjhui</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF873E86F901FA81F96D" author="Pialek L. & Casciotta J. & Almiron A. & Rican O." box="[910,1399,1693,1721]" pageId="18" pageNumber="56" pagination="377 - 395" refId="ref14576" refString="Pialek L., Casciotta J., Almiron A. & Rican O. 2019 b. A new pelagic predatory pike cichlid (Teleostei: Cichlidae: Crenicichla) from the C. mandelburgeri species complex with parallel and reticulate evolution. Hydrobiologia 832: 377 - 395. https: // doi. org / 10.1007 / s 10750 - 018 - 3754 - 1" type="journal article" year="2019">Piálek, Casciotta, Almirón & Říčan, 2019</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB1FF873DB5F941FA81F838" pageId="18" pageNumber="56">
|
||
<paragraph id="372DC62FFFB1FF873DB5F941FA81F886" blockId="18.[189,1401,1757,2019]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DB5F941FA81F886" pageId="18" pageNumber="56">
|
||
9. Midlateral band when present always fully continuous, with barely discernible fluctuation in width, scales in E1 row 78–85, widespread in the Lower Paraná and entire
|
||
<collectingCountry id="4F8586BFFFB1FF873929F963FB78F8C3" box="[1057,1166,1791,1817]" name="Paraguay" pageId="18" pageNumber="56">Paraguay</collectingCountry>
|
||
Basin, also present in Lower
|
||
<collectingCountry id="4F8586BFFFB1FF873C6EF8BDFE3BF8E1" box="[358,461,1825,1851]" name="Uruguay" pageId="18" pageNumber="56">Uruguay</collectingCountry>
|
||
and lower portions of the Middle Paraná in
|
||
<collectingCountry id="4F8586BFFFB1FF873EF0F8BDFB90F8E1" box="[1016,1126,1825,1851]" name="Paraguay" pageId="18" pageNumber="56">Paraguay</collectingCountry>
|
||
,
|
||
<collectingCountry id="4F8586BFFFB1FF87397DF8BCFB36F8E0" box="[1141,1216,1824,1850]" name="Brazil" pageId="18" pageNumber="56">Brazil</collectingCountry>
|
||
,
|
||
<collectingCountry id="4F8586BFFFB1FF8739C6F8BCFAB5F8E0" box="[1230,1347,1824,1850]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB1FF873DE2F8DEFEA7F886" box="[234,337,1858,1884]" name="Uruguay" pageId="18" pageNumber="56">Uruguay</collectingCountry>
|
||
.........................................................................................................
|
||
<taxonomicName id="F092BDACFFB1FF87396EF8DDFA81F886" ID-CoL="ZCHD" authority="Heckel, 1840" authorityName="Heckel" authorityYear="1840" box="[1126,1399,1857,1884]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="vittata">
|
||
<emphasis id="05E61A3DFFB1FF87396EF8DDFB25F881" bold="true" box="[1126,1235,1857,1883]" italics="true" pageId="18" pageNumber="56">C. vittata</emphasis>
|
||
Heckel, 1840
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB1FF873DC9F8F8FA81F838" blockId="18.[189,1401,1757,2019]" pageId="18" pageNumber="56">
|
||
<keyLead id="8C63651AFFB1FF873DC9F8F8FA81F838" pageId="18" pageNumber="56">
|
||
– Midlateral band most often semicontinuous or even distrupted (i.e., always of clearly fluctuating width), vertical bars when present always as clear double-bars, scales in E1 row 42–57, endemic to Middle Paraná main River and some non-isolated tributaries in
|
||
<collectingCountry id="4F8586BFFFB1FF873EDCF83BFBB7F81B" box="[980,1089,1959,1985]" name="Paraguay" pageId="18" pageNumber="56">Paraguay</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB1FF873970F83AFB1BF81A" box="[1144,1261,1958,1984]" name="Argentina" pageId="18" pageNumber="56">Argentina</collectingCountry>
|
||
................ ......................................................................................................
|
||
<taxonomicName id="F092BDACFFB1FF873ED4F854FA81F838" ID-CoL="ZCFK" authority="Kullander, 2009" authorityName="Kullander" authorityYear="2009" box="[988,1399,1991,2018]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="18" pageNumber="56" phylum="Chordata" rank="species" species="mandelburgeri">
|
||
<emphasis id="05E61A3DFFB1FF873ED4F854FB47F838" bold="true" box="[988,1201,1991,2018]" italics="true" pageId="18" pageNumber="56">C. mandelburgeri</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB1FF8739B1F854FA81F838" author="Kullander S. O." box="[1209,1399,1992,2018]" pageId="18" pageNumber="56" pagination="41 - 50" refId="ref14063" refString="Kullander S. O. 2009. Crenicichla mandelburgeri, a new species of cichlid fish (Teleostei: Cichlidae) from the Parana river drainage in Paraguay. Zootaxa 2006 (1): 41 - 50. https: // doi. org / 10.11646 / zootaxa. 2006.1.3" type="journal article" year="2009">Kullander, 2009</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB0FF863DB5FE95FA80FE51" pageId="19" pageNumber="57">
|
||
<paragraph id="372DC62FFFB0FF863DB5FE95FA81FEB3" blockId="19.[189,1399,265,395]" pageId="19" pageNumber="57">
|
||
<keyLead id="8C63651AFFB0FF863DB5FE95FA81FEB3" pageId="19" pageNumber="57">
|
||
10. Breeding females with a distinct dark white-margined blotch in the dorsal fin, a few scattered spots on body below the midlateral band in both sexes, especially in males, endemic to the Piray Guazú tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB0FF863CECFED2FDAFFEB2" box="[484,601,334,360]" name="Argentina" pageId="19" pageNumber="57">Argentina</collectingCountry>
|
||
.................................................................................
|
||
<taxonomicName id="F092BDACFFB0FF8639B7FED2FAE3FEB3" authority="Říčan & Piálek & Almirón & Casciotta, 2023" authorityName="Říčan & Piálek & Almirón & Casciotta" authorityYear="2023" box="[1215,1301,334,361]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="57" phylum="Chordata" rank="species" species="ama" status="sp. nov.">
|
||
<emphasis id="05E61A3DFFB0FF8639B7FED2FAE3FEB3" bold="true" box="[1215,1301,334,361]" italics="true" pageId="19" pageNumber="57">C. ama</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="1ED5A746FFB0FF863814FED3FA81FEB3" box="[1308,1399,335,361]" pageId="19" pageNumber="57" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB0FF863DC9FEEDFA80FE51" blockId="19.[189,1399,265,395]" box="[193,1398,369,395]" pageId="19" pageNumber="57">
|
||
<keyLead id="8C63651AFFB0FF863DC9FEEDFA80FE51" box="[193,1398,369,395]" pageId="19" pageNumber="57">– Breeding females with a distinct dark white-margined long band in the dorsal fin ........................11</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB0FF863DB5FE2EFA80FDEE" pageId="19" pageNumber="57">
|
||
<paragraph id="372DC62FFFB0FF863DB5FE2EFA81FE35" blockId="19.[189,1400,434,564]" pageId="19" pageNumber="57">
|
||
<keyLead id="8C63651AFFB0FF863DB5FE2EFA81FE35" pageId="19" pageNumber="57">
|
||
11. Scattered spots on body below the midlateral band in both sexes, especially in males, endemic to the Iguazú,
|
||
<collectingCountry id="4F8586BFFFB0FF863C4EFE48FE4DFE34" box="[326,443,468,494]" name="Argentina" pageId="19" pageNumber="57">Argentina</collectingCountry>
|
||
and
|
||
<collectingCountry id="4F8586BFFFB0FF863CFCFE48FDCAFE34" box="[500,572,468,494]" name="Brazil" pageId="19" pageNumber="57">Brazil</collectingCountry>
|
||
...........................................................
|
||
<taxonomicName id="F092BDACFFB0FF863EF6FE48FA81FE35" ID-CoL="6BCWX" authority="Haseman, 1911" authorityName="Haseman" authorityYear="1911" box="[1022,1399,468,495]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="57" phylum="Chordata" rank="species" species="iguassuensis">
|
||
<emphasis id="05E61A3DFFB0FF863EF6FE48FB4FFE34" bold="true" box="[1022,1209,468,494]" italics="true" pageId="19" pageNumber="57">C. iguassuensis</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB0FF8639C9FE49FA81FE35" author="Haseman J. D." box="[1217,1399,469,495]" pageId="19" pageNumber="57" pagination="329 - 373" refId="ref13834" refString="Haseman J. D. 1911. An annotated catalog of the cichlid fishes collected by the expedition of the Carnegie Museum to central South America, 1907 - 10. Annals of the Carnegie Museum 7: 329 - 373. Available from https: // www. biodiversitylibrary. org / page / 9956994 [accessed 2 Jun. 2023]." type="journal article" year="1911">Haseman, 1911</bibRefCitation>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="372DC62FFFB0FF863DC9FE64FA80FDEE" blockId="19.[189,1400,434,564]" pageId="19" pageNumber="57">
|
||
<keyLead id="8C63651AFFB0FF863DC9FE64FA80FDEE" pageId="19" pageNumber="57">
|
||
– Without spots on body below the midlateral band in both sexes, endemic to Middle Paraná Basin apart from the Iguazú, endemic to
|
||
<collectingCountry id="4F8586BFFFB0FF863F76FD86FD05FDEE" box="[638,755,538,564]" name="Argentina" pageId="19" pageNumber="57">Argentina</collectingCountry>
|
||
................................................................................. 12
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="8C66DE8AFFB0FF863DB5FDC7FA81FC0B" pageId="19" pageNumber="57">
|
||
<paragraph id="372DC62FFFB0FF863DB5FDC7FA81FCF9" blockId="19.[189,1401,603,979]" pageId="19" pageNumber="57">
|
||
<keyLead id="8C63651AFFB0FF863DB5FDC7FA81FCF9" pageId="19" pageNumber="57">
|
||
12. Anterior two to three midlateral blotches dominant, background colouration light grey, midlateral blotches distinctly H-like with dorsally continuing double bars, females with a wide wide orange to red stripe along the entire length of the basal portion of the dorsal fin above which is found a shorter black band margined in white, suborbital stripe of normal length and width, composed of spots, which do not form horizontal lines, endemic to the Urugua-í tributary, endemic to
|
||
<collectingCountry id="4F8586BFFFB0FF8639A6FD7AFAD5FCDA" box="[1198,1315,742,768]" name="Argentina" pageId="19" pageNumber="57">Argentina</collectingCountry>
|
||
......... ..................................................................
|
||
<taxonomicName id="F092BDACFFB0FF863FDEFC94FA81FCF9" ID-CoL="ZCHH" authority="Casciotta, Almiron, Pialek, Gomez & Rican, 2010" authorityName="Casciotta, Almiron, Pialek, Gomez & Rican" authorityYear="2010" box="[726,1399,776,805]" family="Cichlidae" genus="Crenicichla" kingdom="Animalia" order="Perciformes" pageId="19" pageNumber="57" phylum="Chordata" rank="species" species="ypo">
|
||
<emphasis id="05E61A3DFFB0FF863FDEFC94FCD5FCF9" bold="true" box="[726,803,776,803]" italics="true" pageId="19" pageNumber="57">C. ypo</emphasis>
|
||
<bibRefCitation id="5303BBDEFFB0FF863E22FC95FA81FCF9" author="Casciotta J. & Almiron A. & Pialek L. & Gomez S. & Rican O." box="[810,1399,777,805]" pageId="19" pageNumber="57" pagination="643 - 648" refId="ref13623" refString="Casciotta J., Almiron A., Pialek L., Gomez S. & Rican O. 2010. Crenicichla ypo (Teleostei: Cichlidae), a new species from the middle Parana basin in Misiones, Argentina. Neotropical Ichthyology 8: 643 - 648. https: // doi. org / 10.1590 / S 1679 - 62252010000300009" type="journal article" year="2010">Casciotta, Almirón, Piálek, Gómez & Říčan, 2010</bibRefCitation>
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</taxonomicName>
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</paragraph>
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<paragraph id="372DC62FFFB0FF863DC9FCB0FA81FC0B" blockId="19.[189,1401,603,979]" pageId="19" pageNumber="57">
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– All midlateral blotches well developed, backgroundcolouration very dark grey to blackish, midlateral blotches square-like with dorsally continuing single wide bars, females only with the shorter black band margined in white in the dorsal fin without the orange to red band, suborbital stripe short and wide, ventrally widest, composed of spots which do form horizontal lines, endemic to the Piray Miní tributary, endemic to
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<collectingCountry id="4F8586BFFFB0FF863CEBFC2AFDAEFC0A" box="[483,600,950,976]" name="Argentina" pageId="19" pageNumber="57">Argentina</collectingCountry>
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..............................
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<emphasis id="05E61A3DFFB0FF863E4BFC2AFC71FC0B" bold="true" box="[835,903,950,977]" italics="true" pageId="19" pageNumber="57">C. hu</emphasis>
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<bibRefCitation id="5303BBDEFFB0FF863E86FC2AFA81FC0B" author="Casciotta J. & Almiron A. & Pialek L. & Gomez S. & Rican O." box="[910,1399,950,979]" pageId="19" pageNumber="57" pagination="643 - 648" refId="ref13623" refString="Casciotta J., Almiron A., Pialek L., Gomez S. & Rican O. 2010. Crenicichla ypo (Teleostei: Cichlidae), a new species from the middle Parana basin in Misiones, Argentina. Neotropical Ichthyology 8: 643 - 648. https: // doi. org / 10.1590 / S 1679 - 62252010000300009" type="journal article" year="2010">Piálek, Říčan, Casciotta & Almirón, 2010</bibRefCitation>
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</keyLead>
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</paragraph>
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</key>
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</subSubSection>
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</treatment>
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