228 lines
24 KiB
XML
228 lines
24 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1111.77586" ID-GBIF-Dataset="4f820466-2eff-42bb-9c93-16143363528f" ID-PMC="PMC9848911" ID-Pensoft-Pub="1313-2970-1111-43" ID-Pensoft-UUID="6E23DAFA45395554A61E23AD4B06AD68" ID-PubMed="36760855" ID-ZooBank="3FAAEA839E8141A99B868576F8A1F33A" ModsDocID="1313-2970-1111-43" checkinTime="1657577097621" checkinUser="pensoft" docAuthor="Blahnik, Roger & Andersen, Trond" docDate="2022" docId="9E0D8C53A6545889B555D789B4633D05" docLanguage="en" docName="ZooKeys 1111: 43-198" docOrigin="ZooKeys 1111" docPubDate="2022-07-11" docSource="http://dx.doi.org/10.3897/zookeys.1111.77586" docTitle="Chimarra togoana" docType="treatment" docVersion="4" id="6E23DAFA45395554A61E23AD4B06AD68" lastPageNumber="43" masterDocId="6E23DAFA45395554A61E23AD4B06AD68" masterDocTitle="New species of the genus Chimarra Stephens from Africa (Trichoptera, Philopotamidae) and characterization of the African groups and subgroups of the genus" masterLastPageNumber="198" masterPageNumber="43" pageNumber="43" updateTime="1732488290164" updateUser="ExternalLinkService">
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<mods:title id="B2C0ED152108BB488C1DDCBE68C1FA58">New species of the genus Chimarra Stephens from Africa (Trichoptera, Philopotamidae) and characterization of the African groups and subgroups of the genus</mods:title>
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<mods:name id="B442AA0477D8E921C6AEA70B1C0EE56D" type="personal">
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<mods:namePart id="F7AE88057E32E9C99FA40721FC356146">Blahnik, Roger</mods:namePart>
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<mods:affiliation id="0139BDCC9B064440773A388CA342CCD7">Department of Entomology, University of Minnesota, 1980 Folwell Ave., 219 Hodson Hall, St. Paul, Minnesota, 55108, USA</mods:affiliation>
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<mods:namePart id="FADA5EE70FB3BC603838397B36B8E770">Andersen, Trond</mods:namePart>
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<mods:affiliation id="0F0DC3FC91525794B9E0B96AC1227ACD">Department of Natural History, University Museum of Bergen, University of Bergen, PO Box 7800, NO- 5020 Bergen, Norway</mods:affiliation>
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<mods:nameIdentifier id="CD7D5D8CDC83BD5578ADC16B68CD77FE" type="email">trond.andersen@uib.no</mods:nameIdentifier>
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<mods:title id="419C727DD16409C9C360E6438DB732CC">ZooKeys</mods:title>
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<mods:date id="A45663C15E6EF295BE4C8E010A2E38D3">2022</mods:date>
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<mods:number id="8F41FCBA1EEF2B12661A63E856C6FD86">2022-07-11</mods:number>
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<mods:number id="9A1B4E8E449CEA753FB7C194FBDB2CF4">1111</mods:number>
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<mods:url id="5EFAD1BC4D69248D9C1C1F2BA8C99A9B">http://dx.doi.org/10.3897/zookeys.1111.77586</mods:url>
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<mods:identifier id="09C5B1A6FDF85315205738FF5045AE4C" type="DOI">http://dx.doi.org/10.3897/zookeys.1111.77586</mods:identifier>
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</mods:mods>
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<treatment id="9E0D8C53A6545889B555D789B4633D05" ID-GBIF-Taxon="197225422" LSID="urn:lsid:plazi:treatment:9E0D8C53A6545889B555D789B4633D05" httpUri="http://treatment.plazi.org/id/9E0D8C53A6545889B555D789B4633D05" lastPageNumber="43" pageId="0" pageNumber="43" scope_family="Philopotamidae" scope_order="Trichoptera">
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<subSubSection id="1F6146FE05D1FE3033BD947A584A2CE3" pageId="0" pageNumber="43" type="nomenclature">
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<paragraph id="AC02A036FC869FCF4C982BF2B81CC70F" pageId="0" pageNumber="43">
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<taxonomicName id="AD5DEB090CF8CC531D8E65499DACBBFF" ID-CoL="7S83N" LSID="9E0D8C53-A654-5889-B555-D789B4633D05" authority="(Ulmer, 1907)" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra togoana" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="togoana">Chimarra togoana (Ulmer, 1907)</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection id="38CC7B08DBF730EBE2BB0CBE549F0138" pageId="0" pageNumber="43" type="description">
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<paragraph id="F125304526AA02A113D84149F711C4B3" pageId="0" pageNumber="43">
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<figureCitation id="B16511D460DD9BB92AD2716F2501C069" captionStart="Figure 14" captionStartId="F13" captionText="Figure 14. Chimarra togoana (Ulmer), ♂ genitalia A lateral B dorsal, segments IX and X C inferior appendage, ventral D inferior appendage, dorsal E phallus, lateral." figureDoi="10.3897/zookeys.1111.77586.figure14" httpUri="https://binary.pensoft.net/fig/711480" pageId="0" pageNumber="43">Fig. 14A-E</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="EA8E4D215330AACC71E24BA9ACC9BE9A" pageId="0" pageNumber="43" type="reference_group">
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||
<paragraph id="C37B848B2625725498F9AC180F34450D" pageId="0" pageNumber="43">
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<taxonomicName id="AD0696A3A288F492605EA3FD70F5E8C5" class="Insecta" family="Philopotamidae" genus="Wormaldia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Wormaldia togoana" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="togoana">Wormaldia togoana</taxonomicName>
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Ulmer, 1907: 42-43, figs 61-63.
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</paragraph>
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<paragraph id="EDC77207FFF6CDF6FDC1B0392CA0650A" pageId="0" pageNumber="43">
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<taxonomicName id="ABF84EBB283AE515A4E10165DC4A53E2" class="Insecta" family="Philopotamidae" genus="Chimarrha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarrha togoana" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="togoana">Chimarrha togoana</taxonomicName>
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(Ulmer):
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<bibRefCitation id="0756613E87394CA55D6F7D661D78AB60" DOI="https://doi.org/10.1002/mmnd.48019310102" author="Ulmer, G" journalOrPublisher="Deutsche Entomologische Zeitschrift" pageId="0" pageNumber="43" pagination="1 - 29" refId="B73" refString="Ulmer, G, 1931. Trichopteran von Afrika (hauptsachlich aus dem Britischen Museum). Deutsche Entomologische Zeitschrift 1931: 1 - 29, DOI: https://doi.org/10.1002/mmnd.48019310102" title="Trichopteran von Afrika (hauptsachlich aus dem Britischen Museum)." url="https://doi.org/10.1002/mmnd.48019310102" volume="1931" year="1931">Ulmer 1931</bibRefCitation>
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: 3.
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</paragraph>
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<paragraph id="4921A45E8FBD6B6ED6E308A26FB6DF86" pageId="0" pageNumber="43">
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<taxonomicName id="716F073D956E9C96384DB1E452680DA3" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra togoana" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="togoana">Chimarra togoana</taxonomicName>
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(Ulmer):
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<bibRefCitation id="8FC9E91ADAB993D5D4687EA0BE887934" author="Fischer, FCJ" journalOrPublisher="Memoirs of the Museum of Victoria" pageId="0" pageNumber="43" refId="B18" refString="Fischer, FCJ, 1961. Trichopterorum Catalogus, Vol. II Philopotamidae, Hydroptilidae, Stenopsychidae. Nederlandische Entomologische Vereeniging, Amsterdam." title="Trichopterorum Catalogus, Vol. II Philopotamidae, Hydroptilidae, Stenopsychidae. Nederlandische Entomologische Vereeniging, Amsterdam." year="1961">Fischer 1961</bibRefCitation>
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: 71;
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<bibRefCitation id="5DA9000A8F939F279581E5E7482C3891" DOI="https://doi.org/10.1002/mmnd.4810200410" author="Gibbs, DG" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="43" refId="B21" refString="Gibbs, DG, 1973. The Trichoptera of Ghana. Deutsche Entomologische Zeitschrift 20(4/5): 363-425. https://doi.org/10.1002/mmnd.4810200410" title="The Trichoptera of Ghana. Deutsche Entomologische Zeitschrift 20 (4 / 5): 363 - 425." url="https://doi.org/10.1002/mmnd.4810200410" year="1973">Gibbs 1973</bibRefCitation>
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: 67 (distribution: Ghana).
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="005E7530B8DD5E7EDA8E283C156E971D" pageId="0" pageNumber="43" type="materials_examined">
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<paragraph id="9ED7BCDEB393F2D513F678D8324A3ACC" pageId="0" pageNumber="43">Material examined.</paragraph>
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<paragraph id="D76F1F8C00950A591C70A8B597234B33" pageId="0" pageNumber="43">
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<materialsCitation id="938F1E46910123A7BC4706BA490EF4CD" collectingDate="1993-11-17" collectorName="J Kjaerandsen" country="Ghana" county="Reg." latitude="7.1247225" location="Agumatsa" longLatPrecision="21" longitude="0.5919444" municipality="Wli" specimenCount="3" specimenCount-female="2" specimenCount-male="1" stateProvince="Volta">
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<collectingCountry id="84957CA42F985AEAEA6C138FEFA866CF" name="Ghana">Ghana</collectingCountry>
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-
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<emphasis id="F1600726E0B89C414EF3EBB412F6B1C2" bold="true" pageId="0" pageNumber="43">
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<collectingRegion id="DE96672B7E1EABAB97D2C8FC0136748C" country="Ghana" name="Volta">Volta</collectingRegion>
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<collectingCounty id="58632B4CA4DAC38FFC90E1B87B161118">Reg.</collectingCounty>
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</emphasis>
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●
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<specimenCount id="0A3D11D08653B0AA1C252F29F4949F4A" type="male">1♂</specimenCount>
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<specimenCount id="05D6A12385B90C4BC82298A8EAFF30C4" type="female">2♀♀</specimenCount>
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;
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<collectingMunicipality id="4076A8EF7E37903C43A9CBE12EDF8C72">Wli</collectingMunicipality>
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,
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<location id="CE19EF32CDBF7F114220B48B1B5D10B0" LSID="urn:lsid:plazi:treatment:9E0D8C53A6545889B555D789B4633D05:CE19EF32CDBF7F114220B48B1B5D10B0" country="Ghana" county="Reg." latitude="7.1247225" longLatPrecision="21" longitude="0.5919444" municipality="Wli" name="Agumatsa" stateProvince="Volta">Agumatsa</location>
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waterfall, station # 3;
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<geoCoordinate id="DB6F3322F5817B745383A0DBDB8A3D51" degrees="7" direction="north" minutes="07" orientation="latitude" precision="15" seconds="29" value="7.1247225">7°07'29"N</geoCoordinate>
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,
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<geoCoordinate id="4782F89743B89B9B4A2E1585E1BEB41A" degrees="0" direction="east" minutes="35" orientation="longitude" precision="15" seconds="31" value="0.5919444">0°35'31"E</geoCoordinate>
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;
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<collectingDate id="8FDD54CA88C366F3F319E114E2DD8EEC" value="1993-11-17">17 Nov. 1993</collectingDate>
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;
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<collectorName id="84DF9A67C99F5B3E207565633983E447">
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J
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<normalizedToken id="1FB65F1E517AC27D7EF8C13C76760530" originalValue="Kjærandsen">Kjaerandsen</normalizedToken>
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</collectorName>
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leg.
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</materialsCitation>
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;
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<materialsCitation id="B94EE562DE76BEBC11FAF6CA25AC72C4" collectingDate="1993-11-11" collectingMethod="light trap" collectorName="J Kjaerandsen" country="Ghana" county="Reg." latitude="7.1247225" location="Agumatsa" longLatPrecision="21" longitude="0.5919444" municipality="Wli" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Volta">
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<collectingMethod id="DDA68ED23D8CEAF4E8E9417043D2EAB9">light trap</collectingMethod>
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; ZMBN ●
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<specimenCount id="6B3E7DEB3B17878531DBE35E570D42F3" type="male">1♂</specimenCount>
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; same collection data as for preceding; UMSP ●
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<specimenCount id="1B3E042B6AF8A898265E0BC1CCAC1780" type="female">3♀♀</specimenCount>
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; same collection data as for preceding except station # 10;
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<collectingDate id="E9394EF860B7404570C908EEC135FEB9" value="1993-11-11">11 Nov. 1993</collectingDate>
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</materialsCitation>
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;
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<materialsCitation id="19BCD08440770096F59C729838DB3C4A" collectingDate="1993-11-20" collectorName="J Kjaerandsen" country="Ghana" county="Reg." latitude="7.1247225" location="Agumatsa" longLatPrecision="21" longitude="0.5919444" municipality="Wli" specimenCount="1" specimenCount-female="1" stateProvince="Volta">
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ZMBN ●
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<specimenCount id="9984377E40F33C4596DD3E0FBFA12FEE" type="female">1♀</specimenCount>
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; same collection data as for preceding except
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<collectingDate id="323C2AF0C54E8EE769023EED146D6F32" value="1993-11-20">20 Nov. 1993</collectingDate>
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;
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<collectorName id="D4D5C2D44BBC804E704A1DCA78D18E87">
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J
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<normalizedToken id="29041C154E6A6270E9260743F3032C75" originalValue="Kjærandsen">Kjaerandsen</normalizedToken>
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</collectorName>
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leg.
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</materialsCitation>
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; UMSP.
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</paragraph>
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<caption id="393D68E78E336C7CE983C8C69CE44DCC" doi="10.3897/zookeys.1111.77586.figure14" httpUri="https://binary.pensoft.net/fig/711480" pageId="0" pageNumber="43" start="Figure 14" startId="F13">
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||
<paragraph id="EBD015517D6CFED3F1D6373D502383D1" pageId="0" pageNumber="43">
|
||
<emphasis id="8A0FC3FFB3545BE766A36E9130895576" bold="true" pageId="0" pageNumber="43">Figure 14.</emphasis>
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<taxonomicName id="C97E91238A3EEC8E2E4DADC3BD681004" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra togoana" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="togoana">
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<emphasis id="CDAD679DDA5AE978DB6AC0AF1ACE9142" italics="true" pageId="0" pageNumber="43">Chimarra togoana</emphasis>
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</taxonomicName>
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(Ulmer),
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<specimenCount id="35E643D939D6EE02D60F40A52F0CB994">♂</specimenCount>
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genitalia
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<emphasis id="114455233E2CC750E074FE7CBE33E4D3" bold="true" pageId="0" pageNumber="43">A</emphasis>
|
||
lateral
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||
<emphasis id="CC819AB04935F8854F174DC09AD3C158" bold="true" pageId="0" pageNumber="43">B</emphasis>
|
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dorsal, segments IX and X
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||
<emphasis id="1FF53E557D425D7B9D0BD550306DB641" bold="true" pageId="0" pageNumber="43">C</emphasis>
|
||
inferior appendage, ventral
|
||
<emphasis id="695C875DBB064DABFCDDF5D102A24AA5" bold="true" pageId="0" pageNumber="43">D</emphasis>
|
||
inferior appendage, dorsal
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||
<emphasis id="2136EC0E321E70DE9F61D48773D22C59" bold="true" pageId="0" pageNumber="43">E</emphasis>
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phallus, lateral.
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||
</paragraph>
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||
</caption>
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||
</subSubSection>
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||
<subSubSection id="1545E42BF5B9E05670334F18D328F2E9" pageId="0" pageNumber="43" type="diagnosis">
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||
<paragraph id="8E88359BFFC162FE3F69A7EBF4B833B2" pageId="0" pageNumber="43">Diagnosis.</paragraph>
|
||
<paragraph id="301DA0DB5A4C4E1E4AFC5F35859B3054" pageId="0" pageNumber="43">
|
||
<taxonomicName id="9534C9ECECEF171D6A39AD2506DF58F0" baseAuthorityName="Ulmer" baseAuthorityYear="1907" class="Insecta" family="Philopotamidae" genus="Chimarra" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarra togoana" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="togoana">
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<emphasis id="D8D139F19D536D801ACB4831E83195F0" italics="true" pageId="0" pageNumber="43">Chimarra togoana</emphasis>
|
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</taxonomicName>
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||
is a very distinctive species, readily identified by the elongate, apically flared shape of its inferior appendages, with a distinctive mesal cusp at approximately midlength, and the elongate ventral process of segment IX, which is somewhat inflated apically, but apparently lacks the cluster of apicoventral spines characteristic of species in the
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<taxonomicName id="D4E6A1BAEC3A2ECB89A7E0A4F0BD65C6" class="Insecta" family="Philopotamidae" genus="Chimarrha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarrha ruficeps" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="ruficeps">
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<emphasis id="05F160C2D2B5091CCECAC10BEEA4ED1B" italics="true" pageId="0" pageNumber="43">Chimarrha ruficeps</emphasis>
|
||
</taxonomicName>
|
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subgroup. It is only provisionally placed in the
|
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<taxonomicName id="BB577714B8C2F67F5736D69DC37EB3D5" class="Insecta" family="Philopotamidae" genus="Chimarrha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarrha fallax" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="fallax">
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<emphasis id="83EE7BE51238A9364902F47E94B6E77B" italics="true" pageId="0" pageNumber="43">Chimarrha fallax</emphasis>
|
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</taxonomicName>
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subgroup since some of its characters could equally well be used to place it in the
|
||
<taxonomicName id="D615B47DAFEECDE3F129A44E0072F497" class="Insecta" family="Philopotamidae" genus="Chimarrha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarrha ruficeps" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="ruficeps">
|
||
<emphasis id="B78DD236D319B71131B6EE778E825688" italics="true" pageId="0" pageNumber="43">Chimarrha ruficeps</emphasis>
|
||
</taxonomicName>
|
||
subgroup. Characters supporting the latter interpretation include the overall shape of segment IX, which is strongly produced anteroventrally and has its ventromesal margin concave, and the distinctly formed and enlarged dorsolateral apodemes of the same segment. Characters supporting its placement in the
|
||
<taxonomicName id="A27439C2C0EEAD8575A378144721E48F" class="Insecta" family="Philopotamidae" genus="Chimarrha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarrha fallax" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="fallax">
|
||
<emphasis id="EBF8D51357249EC145DBB70F7C1FBE4E" italics="true" pageId="0" pageNumber="43">Chimarrha fallax</emphasis>
|
||
</taxonomicName>
|
||
subgroup include the posteroventral projection of segment VIII. It is also possible that it belongs to a lineage basal to both of those subgroups. The rather simple tergum X, with an apicolateral cleft on each of its lateral lobes, is probably a primitive character; it may be ancestral to both subgroups, if the periphallic processes of the
|
||
<taxonomicName id="57C15F484F37642FFDF747599DECC443" class="Insecta" family="Philopotamidae" genus="Chimarrha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Chimarrha fallax" order="Trichoptera" pageId="0" pageNumber="43" phylum="Arthropoda" rank="species" species="fallax">
|
||
<emphasis id="10AA6C6706F2634CC22B153887FAD124" italics="true" pageId="0" pageNumber="43">Chimarrha fallax</emphasis>
|
||
</taxonomicName>
|
||
subgroup had their origin as a cleft in each of the lateral lobes of tergum X.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="58650FB2172E47C0BF33EE64DE3D5068" pageId="0" pageNumber="43" type="redescription">
|
||
<paragraph id="B27266557E088945AAF8B975B3B2D635" pageId="0" pageNumber="43">Redescription.</paragraph>
|
||
<paragraph id="D2951868CAEB8D53619CAAABEC3126E4" pageId="0" pageNumber="43">
|
||
<emphasis id="936A4F09066EFB2916B860EC460D430B" bold="true" italics="true" pageId="0" pageNumber="43">Adult.</emphasis>
|
||
Overall color (in alcohol) nearly uniformly yellowish brown. Head relatively short (postocular parietal sclerite ~ 1/2 diameter of eye). Palps relatively elongate; maxillary palp with 1st segment very short (approximately as long as wide), 2nd segment short (~ 3
|
||
<normalizedToken id="44D3C4696CC6D77ADA30D6705576BD2A" originalValue="×">x</normalizedToken>
|
||
1st), apex with small cluster of stiff setae, 3rd segment relatively elongate (nearly 2
|
||
<normalizedToken id="C6FA9FBE6964717481DC839E5A4E4DE6" originalValue="×">x</normalizedToken>
|
||
2nd), 4th segment short (slightly shorter than 2nd), 5th segment elongate (nearly as long as 3rd and 4th combined). Forewing length: male, 6.2-7.0 mm; female, 6.5-7.5 mm. Fore- and hind wings with forks I, II, III, and V present. Forewing with R1 somewhat sinuous, stem of Rs weakly inflected at past midlength, without node at inflection, basal fork of discoidal cell not enlarged, fork nearly symmetric, length of cell ~ 2
|
||
<normalizedToken id="09B995F538308425740A48B77BD16270" originalValue="×">x</normalizedToken>
|
||
width, fork I slightly subsessile, fork II sessile,
|
||
<emphasis id="4059D65764EDA4194251CFA39A6D61DF" italics="true" pageId="0" pageNumber="43">r</emphasis>
|
||
crossvein diagonal, intersecting discoidal cell at past midlength, just before fork I,
|
||
<emphasis id="82C9BD3CE39F925332007499E53EC88E" italics="true" pageId="0" pageNumber="43">s</emphasis>
|
||
and
|
||
<emphasis id="86FB61721A1A55C52971CF7A71A91ED6" italics="true" pageId="0" pageNumber="43">r-m</emphasis>
|
||
crossveins linear,
|
||
<emphasis id="4CFB5756067D4667A83FC90EDE93EC2F" italics="true" pageId="0" pageNumber="43">m</emphasis>
|
||
crossvein very distinctly more proximal,
|
||
<emphasis id="ADD54732AD3D084CDCCEC9E27AD8BBAA" italics="true" pageId="0" pageNumber="43">s</emphasis>
|
||
pigmented (like wing),
|
||
<emphasis id="701166972FEDB5BF76A32C96B1B7496F" italics="true" pageId="0" pageNumber="43">r-m</emphasis>
|
||
and
|
||
<emphasis id="87AC455853FAC807EB84275B5193DF2F" italics="true" pageId="0" pageNumber="43">m</emphasis>
|
||
crossveins hyaline. 2A with crossvein (apparently forked apically to 1A and 3A). Hind wing with R1 narrowly parallel to subcosta, forks I and II strongly subsessile, fork III distal and relatively wide, anal loop small. Foreleg with apical tibial spur distinct; male with foretarsi unmodified, claws small and symmetrical.
|
||
</paragraph>
|
||
<paragraph id="9435CFF7188B3ECC7B2C20F2FA134316" pageId="0" pageNumber="43">
|
||
<emphasis id="9405188A18A0E72425B7C8E5C10FD1ED" bold="true" pageId="0" pageNumber="43">
|
||
<emphasis id="EFC0CE641D12ECA378FDD5E4E7E6FA64" italics="true" pageId="0" pageNumber="43">Male genitalia</emphasis>
|
||
.
|
||
</emphasis>
|
||
Segment VIII relatively short, sternum with short, posteriorly projecting, ventromesal projection, tergum slightly longer than sternum. Segment IX, in lateral view, with anterior margin distinctly produced and rounded in ventral
|
||
<normalizedToken id="F129E829621B177B13BD709D30AB1ABC" originalValue="¼">1/4</normalizedToken>
|
||
, dorsolaterally with prominent rounded apodemes, margin strongly convex between apodemes; tergum continuous dorsally, forming deep, narrow emargination mesally between apodemes; posterior margin broadly convex; posteroventral margin with elongate, narrow, posteriorly-projecting, ventral process, apex of process slightly expanded. Segment IX, in dorsal or ventral views, with anteroventral margin strongly concave. Lateral lobes of tergum X short, each partially divided from posterior margin into dorsal and ventral lobes, dorsal lobe with two sensilla in basal half; mesal lobe of tergum X very short, membranous. Preanal appendages short and rounded, somewhat flattened, constricted basally. Inferior appendage, in lateral view, elongate, projecting, widened and flared apically, distal margin subtruncate; appendage with prominent, sclerotized mesal cusp at approximately midlength, visible in lateral view as notch on ventral margin. Phallic apparatus with phallobase tubular, with usual basodorsal expansion, apicoventral margin only weakly projecting, endotheca with three spines, one relatively elongate, curved, and strongly sclerotized, other two relatively short, asymmetrically positioned; phallotremal sclerite complex composed of short rod and ring structure with small apical sclerite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="759F4C07701C1017918200644D2454CD" pageId="0" pageNumber="43" type="distribution">
|
||
<paragraph id="9818CA323A83F8B6EBCAAAF52096FFAB" pageId="0" pageNumber="43">Distribution.</paragraph>
|
||
<paragraph id="47FA9609708F15E4AB1CACFA4231BEE0" pageId="0" pageNumber="43">Ghana, Togo.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |