337 lines
53 KiB
XML
337 lines
53 KiB
XML
<document id="D49586F31C2D0885BCACC606D75667D5" ID-CLB-Dataset="22391" ID-DOI="10.5852/ejt.2015.139" ID-GBIF-Dataset="8a8a87b6-7e06-4004-b74e-4bb7b0eca921" ID-ISSN="2118-9773" ID-Zenodo-Dep="3785506" ID-ZooBank="F7065B0E-FCCC-45EC-9C4F-19F6C9B972B3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588622268591" checkinUser="carolina" docAuthor="Vandenspiegel, D., Golovatch, S. I. & Fiemapong, A. R. Nzoko" docDate="2015" docId="69391077FFA39B0AFD86F908FCDAA3A2" docLanguage="en" docName="ejt-2015-139.pdf.imf" docOrigin="European Journal of Taxonomy 139" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Campodesmoides corniger Vandenspiegel & Golovatch & Fiemapong 2015, sp. nov." docType="treatment" docUuid="E20A8CC8-F56C-4499-B1F0-F3D541275368" docUuidSource="ZooBank" docVersion="6" lastPageNumber="6" masterDocId="9500680FFFA29B0FFFCFFFFEFFF8A610" masterDocTitle="Two new species, including one representing a new genus, of the West African millipede family Campodesmidae (Diplopoda: Polydesmida)" masterLastPageNumber="11" masterPageNumber="1" pageNumber="2" updateTime="1698838444074" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-3.0" zenodo-license-figures="CC-BY-3.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="FB497A7B4DC302D03BE76BFAD70B1705">Two new species, including one representing a new genus, of the West African millipede family Campodesmidae (Diplopoda: Polydesmida)</mods:title>
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<mods:name id="9C988785A147CAEB3C47DEB891EC6AD0" type="personal">
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<mods:namePart id="0FD49EB48D5E5D907D4BAE69230F8D1B">Vandenspiegel, D.</mods:namePart>
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<mods:affiliation id="AEB5E7DE805274FEA964FB242E3FFE05">Musée Royal de l’Afrique Centrale, B- 3480 Tervuren, Belgique. Email: didier. van. den. spiegel @ africamuseum. be (corresponding author) & urn: lsid: zoobank. org: author: CE 8 C 3 D 01 - 28 AD- 43 F 7 - 9 D 4 F- 04802 E 68 CB 1 A</mods:affiliation>
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<mods:roleTerm id="833ADCCAD47C150200CAFE4BA59E7265">Author</mods:roleTerm>
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<mods:namePart id="8E64EF2211FEA2DA4E2C685CA05D7561">Golovatch, S. I.</mods:namePart>
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<mods:affiliation id="EF8BA7BCE22EDEF5152B57465400411F">Institute for Problems of Ecology and Evolution, Russian Academy of Sciences, Leninsky prospekt 33, Moscow 119071, Russia. Email: sgolovatch @ yandex. ru (corresponding author) & urn: lsid: zoobank. org: author: 71532 F 45 - BDD 5 - 415 D-BC 54 - 86256 E 5 D 5 D 4 A</mods:affiliation>
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<mods:namePart id="E389BBF9252C5A3CF40C553B39D6D733">Fiemapong, A. R. Nzoko</mods:namePart>
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<mods:affiliation id="69CF126D0BB9AB49A750241DDB0E2C0F">Laboratoire de Zoologie, Université Yaoundé 1, BP 812, Cameroon. Email: armandnzoko @ gmail. com & urn: lsid: zoobank. org: author: ECDC 7821 - D 8 DA- 49 D 4 - 9 D 6 A- 13 A 6 E 732863 E</mods:affiliation>
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<mods:title id="68DF4605D221042ADB7344166A5F2663">European Journal of Taxonomy</mods:title>
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<mods:date id="890C60D85591B9F950ED737B7CB50ED4">2015</mods:date>
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<mods:number id="822BFDF1B55AB4497E57179F4F9CF1EF">2015-09-28</mods:number>
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<mods:number id="5B5C9185DA5B319EA35A5381599A940A">139</mods:number>
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<treatment id="69391077FFA39B0AFD86F908FCDAA3A2" ID-DOI="http://doi.org/10.5281/zenodo.3794591" ID-GBIF-Taxon="163943974" ID-Zenodo-Dep="3794591" LSID="urn:lsid:zoobank.org:act:E20A8CC8-F56C-4499-B1F0-F3D541275368" httpUri="http://treatment.plazi.org/id/69391077FFA39B0AFD86F908FCDAA3A2" lastPageId="5" lastPageNumber="6" pageId="1" pageNumber="2">
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<subSubSection id="A98AF2EAFFA39B0EFD86F908FC13A101" box="[585,1003,1782,1809]" pageId="1" pageNumber="2" type="nomenclature">
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<paragraph id="E12FA161FFA39B0EFD86F908FC13A101" blockId="1.[372,1215,1782,1879]" box="[585,1003,1782,1809]" pageId="1" pageNumber="2">
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<heading id="BA67160DFFA39B0EFD86F908FC13A101" box="[585,1003,1782,1809]" centered="true" fontSize="11" level="2" pageId="1" pageNumber="2" reason="2">
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<taxonomicName id="2690DAE2FFA39B0EFD86F908FC71A100" authority="Vandenspiegel & Golovatch & Fiemapong, 2015" authorityName="Vandenspiegel & Golovatch & Fiemapong" authorityYear="2015" box="[585,905,1782,1809]" class="Diplopoda" family="Campodesmidae" genus="Campodesmoides" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="corniger" status="sp. nov.">
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<emphasis id="D3E47D73FFA39B0EFD86F908FC71A100" bold="true" box="[585,905,1782,1809]" italics="true" pageId="1" pageNumber="2">Campodesmoides corniger</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="C8D7C008FFA39B0EFC5FF909FC13A101" box="[912,1003,1783,1809]" pageId="1" pageNumber="2" rank="species">sp. nov.</taxonomicNameLabel>
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</heading>
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</paragraph>
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<subSubSection id="A98AF2EAFFA39B0EFEBBF8E4FCB5A147" pageId="1" pageNumber="2" type="description">
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<uri id="9501AD63FFA39B0EFEBBF8E4FB47A125" box="[372,1215,1818,1845]" pageId="1" pageNumber="2">
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urn:lsid:zoobank.org:act:
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<uuid id="95369BB4FFA39B0EFD5AF8E4FB47A125" box="[661,1215,1818,1845]" pageId="1" pageNumber="2">E20A8CC8-F56C-4499-B1F0-F3D541275368</uuid>
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</uri>
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</paragraph>
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<paragraph id="E12FA161FFA39B0EFD28F8C3FCB5A147" blockId="1.[372,1215,1782,1879]" box="[743,845,1853,1879]" pageId="1" pageNumber="2">
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<figureCitation id="79ABBDE4FFA39B0EFD28F8C3FCB5A147" box="[743,845,1853,1879]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="2.[189,232,1965,1991]" captionStartId-1="3.[189,231,1824,1850]" captionStartId-2="4.[189,232,1930,1956]" captionTargetBox-0="[189,1399,897,1925]" captionTargetBox-1="[189,1399,643,1783]" captionTargetBox-2="[189,1384,1326,1890]" captionTargetId-0="figure@2.[189,1399,897,1925]" captionTargetId-1="figure@3.[189,1399,643,1783]" captionTargetId-2="figure@4.[189,1399,1326,1890]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." captionText-1="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." captionText-2="Fig. 3. Campodesmoides corniger sp. nov., ♂, holotype. A–B. Left gonopod, mesal and lateral views, respectively. C–E. Right gonopod, mesal, lateral and ventral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi-0="http://doi.org/10.5281/zenodo.3785508" figureDoi-1="http://doi.org/10.5281/zenodo.3785510" figureDoi-2="http://doi.org/10.5281/zenodo.3785512" httpUri-0="https://zenodo.org/record/3785508/files/figure.png" httpUri-1="https://zenodo.org/record/3785510/files/figure.png" httpUri-2="https://zenodo.org/record/3785512/files/figure.png" pageId="1" pageNumber="2">Figs 1–3</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A98AF2EAFFA39B0DFF72F880FB25A7C5" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="diagnosis">
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<paragraph id="E12FA161FFA39B0EFF72F880FECEA188" blockId="1.[189,310,1918,1944]" box="[189,310,1918,1944]" pageId="1" pageNumber="2">
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<heading id="BA67160DFFA39B0EFF72F880FECEA188" bold="true" box="[189,310,1918,1944]" fontSize="11" level="3" pageId="1" pageNumber="2" reason="3">
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<emphasis id="D3E47D73FFA39B0EFF72F880FECEA188" bold="true" box="[189,310,1918,1944]" pageId="1" pageNumber="2">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E12FA161FFA39B0DFF72F850FA8FA736" blockId="1.[189,1399,1965,2027]" lastBlockId="2.[189,1399,267,294]" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2">
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A species of
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<taxonomicName id="2690DAE2FFA39B0EFE93F853FDD8A1D7" authorityName="Cook" authorityYear="1895" box="[348,544,1965,1991]" class="Diplopoda" family="Campodesmidae" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Campodesmidae</taxonomicName>
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which differs from other species of the genus
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<taxonomicName id="2690DAE2FFA39B0EFB97F853FAFBA1D7" authorityName="Cook" authorityYear="1896" box="[1112,1283,1965,1991]" class="Diplopoda" family="Campodesmidae" genus="Campodesmus" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="genus">
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<emphasis id="D3E47D73FFA39B0EFB97F853FAFBA1D7" box="[1112,1283,1965,1991]" italics="true" pageId="1" pageNumber="2">Campodesmus</emphasis>
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</taxonomicName>
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, the only hitherto recognized genus of
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<taxonomicName id="2690DAE2FFA39B0EFDD2F82FFD1CA1FB" authorityName="Cook" authorityYear="1895" box="[541,740,2001,2027]" class="Diplopoda" family="Campodesmidae" kingdom="Animalia" order="Polydesmida" pageId="1" pageNumber="2" phylum="Arthropoda" rank="family">Campodesmidae</taxonomicName>
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, by the long and only slightly clavate antennae, which reach behind segment 3 when stretched dorsally; the considerably longer legs which in the male are
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</paragraph>
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<paragraph id="E12FA161FFA09B0DFF72FED0FB25A7C5" blockId="2.[189,1399,302,470]" pageId="2" pageNumber="3">
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about 1.5 times as long as midbody height; the normal pore formula: 5, 7, 9, 10, 12, 13, 15–19, the ozopores being borne on conspicuous porosteles (versus reduced to 5, 7 without porosteles); the more dorsally situated paranota which are set at the upper ¼ of midbody height (versus about half of midbody height); the nearly flat sterna and the gonopod telopodites being suberect,
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<emphasis id="D3E47D73FFA09B0DFBE3FE67FB8DA7A2" box="[1068,1141,409,434]" italics="true" pageId="2" pageNumber="3">in situ</emphasis>
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directed forward, not crossing mesally, each with a clearly twisted solenophore devoid of a distinct solenomere.
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</paragraph>
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</subSubSection>
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<subSubSection id="A98AF2EAFFA09B0DFF72FE03FC75A478" pageId="2" pageNumber="3" type="etymology">
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<paragraph id="E12FA161FFA09B0DFF72FE03FEBCA407" blockId="2.[189,324,509,535]" box="[189,324,509,535]" pageId="2" pageNumber="3">
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<heading id="BA67160DFFA09B0DFF72FE03FEBCA407" bold="true" box="[189,324,509,535]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="D3E47D73FFA09B0DFF72FE03FEBCA407" bold="true" box="[189,324,509,535]" pageId="2" pageNumber="3">Etymology</emphasis>
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</heading>
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</paragraph>
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<paragraph id="E12FA161FFA09B0DFF72FDD2FC75A478" blockId="2.[189,1398,555,617]" pageId="2" pageNumber="3">To emphasize the high, vertical, paramedian horns present on metatergum 2, these growing increasingly small on metatergum 3 and, especially, 4; noun in apposition.</paragraph>
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</subSubSection>
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<subSubSection id="A98AF2EAFFA09B0CFF72FD6EFE6DA79E" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3" type="materials_examined">
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<paragraph id="E12FA161FFA09B0DFF72FD6EFE52A4BA" blockId="2.[189,426,656,682]" box="[189,426,656,682]" pageId="2" pageNumber="3">
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<heading id="BA67160DFFA09B0DFF72FD6EFE52A4BA" bold="true" box="[189,426,656,682]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
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<emphasis id="D3E47D73FFA09B0DFF72FD6EFE52A4BA" bold="true" box="[189,426,656,682]" pageId="2" pageNumber="3">Material examined</emphasis>
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</heading>
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</paragraph>
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<materialsCitation id="51F8AB3CFFA09B0DFF25FD41FE31A50F" ID-GBIF-Occurrence="2610058767" collectingDate="2014-04-26" collectionCode="MRAC" collectorName="A. Nzoko" country="Cameroon" elevation="830" latitude="3.8666668" location="Ongot" longLatPrecision="1307" longitude="11.35" pageId="2" pageNumber="3" specimenCode="MRAC 22679" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<paragraph id="E12FA161FFA09B0DFF25FD41FEA4A4C9" blockId="2.[189,1398,703,799]" box="[234,348,703,729]" pageId="2" pageNumber="3">
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<emphasis id="D3E47D73FFA09B0DFF25FD41FEA4A4C9" bold="true" box="[234,348,703,729]" pageId="2" pageNumber="3">
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<typeStatus id="3E2B1FC3FFA09B0DFF25FD41FEA4A4C9" box="[234,348,703,729]" pageId="2" pageNumber="3" type="holotype">Holotype</typeStatus>
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</emphasis>
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</paragraph>
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<paragraph id="E12FA161FFA09B0DFF72FD1CFE31A50F" blockId="2.[189,1398,703,799]" pageId="2" pageNumber="3">
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<collectingCountry id="9987E1F1FFA09B0DFF72FD1CFE9DA4EC" box="[189,357,738,764]" name="Cameroon" pageId="2" pageNumber="3">CAMEROON</collectingCountry>
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:
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<specimenCount id="F7966AE8FFA09B0DFEB7FD1CFE68A4EC" box="[376,400,738,764]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
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,
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<location id="E44FF7BAFFA09B0DFE6EFD1CFE15A4EC" LSID="urn:lsid:plazi:treatment:69391077FFA39B0AFD86F908FCDAA3A2:E44FF7BAFFA09B0DFE6EFD1CFE15A4EC" box="[417,493,738,764]" country="Cameroon" latitude="3.8666668" longLatPrecision="1307" longitude="11.35" name="Ongot" pageId="2" pageNumber="3">Ongot</location>
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,
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<geoCoordinate id="84A4C7A6FFA09B0DFE32FD1CFD97A4EC" box="[509,623,737,764]" degrees="03" direction="north" minutes="52" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="3.8666668">N 03°52’</geoCoordinate>
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,
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,
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<emphasis id="D3E47D73FFA09B0DFCC2FD1DFCD9A4EC" box="[781,801,739,764]" italics="true" pageId="2" pageNumber="3">c.</emphasis>
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<elevation id="6ABD4652FFA09B0DFCE2FD1CFC46A4EC" box="[813,958,738,764]" metricMagnitude="2" metricUnit="m" metricValue="8.3" pageId="2" pageNumber="3" unit="m" value="830.0">
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a.s.l.
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</elevation>
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, open terrain,
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<date id="952E87A1FFA09B0DFBBDFD1CFAEEA4EC" box="[1138,1302,738,764]" pageId="2" pageNumber="3" value="2014-04-26">
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<collectingDate id="856A7E49FFA09B0DFBBDFD1CFAEEA4EC" box="[1138,1302,738,764]" pageId="2" pageNumber="3" value="2014-04-26">26 Apr. 2014</collectingDate>
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</date>
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, leg.
|
||
<collectorName id="4C65C4B7FFA09B0DFA95FD1CFEF6A50F" pageId="2" pageNumber="3">A. Nzoko</collectorName>
|
||
,
|
||
<specimenCode id="B136091AFFA09B0DFED5FCFBFE3DA50F" box="[282,453,773,799]" collectionCode="MRAC" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="2" pageNumber="3" type="Museum">MRAC 22679</specimenCode>
|
||
.
|
||
</paragraph>
|
||
</materialsCitation>
|
||
<caption id="B5EFF1E9FFA09B0DFF72F853FEE5A1FB" ID-DOI="http://doi.org/10.5281/zenodo.3785508" ID-Zenodo-Dep="3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="2" pageNumber="3" startId="2.[189,232,1965,1991]" targetBox="[189,1399,897,1925]" targetPageId="2">
|
||
<paragraph id="E12FA161FFA09B0DFF72F853FEE5A1FB" blockId="2.[189,1398,1965,2027]" pageId="2" pageNumber="3">
|
||
<emphasis id="D3E47D73FFA09B0DFF72F853FEF2A1D7" bold="true" box="[189,266,1965,1991]" pageId="2" pageNumber="3">Fig. 1.</emphasis>
|
||
Habitus of
|
||
<taxonomicName id="2690DAE2FFA09B0DFE5EF853FD33A1D7" authority="Vandenspiegel & Golovatch & Fiemapong, 2015" authorityName="Vandenspiegel & Golovatch & Fiemapong" authorityYear="2015" box="[401,715,1965,1991]" class="Diplopoda" family="Campodesmidae" genus="Campodesmoides" kingdom="Animalia" order="Polydesmida" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="corniger" status="sp. nov.">
|
||
<emphasis id="D3E47D73FFA09B0DFE5EF853FD33A1D7" box="[401,715,1965,1991]" italics="true" pageId="2" pageNumber="3">Campodesmoides corniger</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C8D7C008FFA09B0DFD1DF850FCD6A1D8" box="[722,814,1966,1992]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♂, holotype, lateral view. Photo by J. Brecko, not to scale.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E12FA161FFA19B0CFF25FEF5FE9FA735" blockId="3.[189,1399,267,399]" box="[234,359,267,293]" pageId="3" pageNumber="4">
|
||
<emphasis id="D3E47D73FFA19B0CFF25FEF5FE9FA735" bold="true" box="[234,359,267,293]" pageId="3" pageNumber="4">
|
||
<typeStatus id="3E2B1FC3FFA19B0CFF25FEF5FE9FA735" box="[234,359,267,293]" pageId="3" pageNumber="4" type="paratype">Paratypes</typeStatus>
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="E12FA161FFA19B0CFF72FED0FE6DA79E" blockId="3.[189,1399,267,399]" pageId="3" pageNumber="4">
|
||
<collectingCountry id="9987E1F1FFA19B0CFF72FED0FE9DA758" box="[189,357,302,328]" name="Cameroon" pageId="3" pageNumber="4">CAMEROON</collectingCountry>
|
||
:
|
||
<specimenCount id="F7966AE8FFA19B0CFEBAFED0FE5AA759" box="[373,418,302,329]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
|
||
, same locality, together with
|
||
<typeStatus id="3E2B1FC3FFA19B0CFD30FED0FC91A758" box="[767,873,302,328]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
,
|
||
<collectionCode id="878139A4FFA19B0CFCBAFED0FC33A758" box="[885,971,302,328]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="3" pageNumber="4" type="Museum">MRAC</collectionCode>
|
||
22.681;
|
||
<specimenCount id="F7966AE8FFA19B0CFBFBFED0FB98A759" box="[1076,1120,302,329]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
|
||
, same locality,
|
||
<date id="952E87A1FFA19B0CFAD6FED0FF03A77B" pageId="3" pageNumber="4" value="2014-11-29">29 Nov. 2014</date>
|
||
,
|
||
<collectionCode id="878139A4FFA19B0CFECAFEAFFEA3A77B" box="[261,347,337,363]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="3" pageNumber="4" type="Museum">MRAC</collectionCode>
|
||
22.678;
|
||
<specimenCount id="F7966AE8FFA19B0CFE70FEAFFE12A77C" box="[447,490,337,364]" pageId="3" pageNumber="4" type="female">1 ♀</specimenCount>
|
||
, Egoutadjap,
|
||
<geoCoordinate id="84A4C7A6FFA19B0CFD45FEACFD0FA77B" box="[650,759,337,364]" degrees="02" direction="north" minutes="43" orientation="latitude" pageId="3" pageNumber="4" precision="925" value="2.7166667">N 02°43’</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="84A4C7A6FFA19B0CFCCDFEACFC92A77B" box="[770,874,337,364]" degrees="11" direction="east" minutes="04" orientation="longitude" pageId="3" pageNumber="4" precision="925" value="11.066667">E 11°04’</geoCoordinate>
|
||
, forest,
|
||
<date id="952E87A1FFA19B0CFC09FEAFFB9BA77B" box="[966,1123,337,364]" pageId="3" pageNumber="4" value="2014-12-06">06 Dec. 2014</date>
|
||
,
|
||
<collectionCode id="878139A4FFA19B0CFBA2FEAFFB3BA77B" box="[1133,1219,337,363]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de l’Afrique Centrale" pageId="3" pageNumber="4" type="Museum">MRAC</collectionCode>
|
||
22.680 (
|
||
<collectionCode id="878139A4FFA19B0CFAE7FEAFFA92A77B" box="[1320,1386,337,363]" pageId="3" pageNumber="4">SEM</collectionCode>
|
||
). All leg. A. Nzoko.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A98AF2EAFFA19B0AFF72FE48FB23A30E" lastPageId="5" lastPageNumber="6" pageId="3" pageNumber="4" type="description">
|
||
<paragraph id="E12FA161FFA19B0CFF72FE48FEB6A7C0" blockId="3.[189,334,438,464]" box="[189,334,438,464]" pageId="3" pageNumber="4">
|
||
<heading id="BA67160DFFA19B0CFF72FE48FEB6A7C0" bold="true" box="[189,334,438,464]" fontSize="11" level="3" pageId="3" pageNumber="4" reason="3">
|
||
<emphasis id="D3E47D73FFA19B0CFF72FE48FEB6A7C0" bold="true" box="[189,334,438,464]" pageId="3" pageNumber="4">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E12FA161FFA19B0BFF72FE1AFA8EA735" blockId="3.[189,1399,484,582]" lastBlockId="4.[189,1398,267,294]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
|
||
Length of
|
||
<typeStatus id="3E2B1FC3FFA19B0CFEF2FE1AFE5CA7EE" box="[317,420,484,510]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
|
||
<emphasis id="D3E47D73FFA19B0CFE61FE1BFE3AA7EE" box="[430,450,485,510]" italics="true" pageId="3" pageNumber="4">c.</emphasis>
|
||
<quantity id="26680C84FFA19B0CFE03FE1AFDDDA7EF" box="[460,549,484,511]" metricMagnitude="-2" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="4" unit="mm" value="26.0">26 mm</quantity>
|
||
, width on midbody pro- and metazonae 2.5 and 5.0 mm, respectively. female
|
||
<typeStatus id="3E2B1FC3FFA19B0CFEDAFDF6FE7DA432" box="[277,389,520,546]" pageId="3" pageNumber="4" type="paratype">paratypes</typeStatus>
|
||
<quantity id="26680C84FFA19B0CFE41FDF9FDF6A432" box="[398,526,519,546]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" metricValueMax="3.0" metricValueMin="2.5" pageId="3" pageNumber="4" unit="mm" value="27.5" valueMax="30.0" valueMin="25.0">25–30 mm</quantity>
|
||
long, 2.5–3.6 and 3.6–5.0 mm wide on pro- and metazonae, respectively. Coloration from light grey-brown (female) to blackish brown (male, female) (
|
||
<figureCitation id="79ABBDE4FFA19B0CFB82FDD5FB68A455" box="[1101,1168,555,581]" captionStart="Fig" captionStartId="2.[189,232,1965,1991]" captionTargetBox="[189,1399,897,1925]" captionTargetId="figure@2.[189,1399,897,1925]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
|
||
), darker specimens with brown to chocolate brown antennae and legs; tips of antennae, distal parts of paranota 5, 7, 9 and 13
|
||
</paragraph>
|
||
<caption id="B5EFF1E9FFA19B0CFF72F8DEFDD7A1FB" ID-DOI="http://doi.org/10.5281/zenodo.3785510" ID-Zenodo-Dep="3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,231,1824,1850]" targetBox="[189,1399,643,1783]" targetPageId="3">
|
||
<paragraph id="E12FA161FFA19B0CFF72F8DEFDD7A1FB" blockId="3.[189,1399,1824,2027]" pageId="3" pageNumber="4">
|
||
<emphasis id="D3E47D73FFA19B0CFF72F8DEFEF5A12A" bold="true" box="[189,269,1824,1850]" pageId="3" pageNumber="4">Fig. 2.</emphasis>
|
||
SEM micrographs of
|
||
<taxonomicName id="2690DAE2FFA19B0CFDD1F8DEFCA1A12A" authority="Vandenspiegel & Golovatch & Fiemapong, 2015" authorityName="Vandenspiegel & Golovatch & Fiemapong" authorityYear="2015" box="[542,857,1824,1850]" class="Diplopoda" family="Campodesmidae" genus="Campodesmoides" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="corniger" status="sp. nov.">
|
||
<emphasis id="D3E47D73FFA19B0CFDD1F8DEFCA1A12A" box="[542,857,1824,1850]" italics="true" pageId="3" pageNumber="4">Campodesmoides corniger</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C8D7C008FFA19B0CFCABF8DFFC3CA12B" box="[868,964,1825,1851]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♀, paratype, MRAC 22.680.
|
||
<emphasis id="D3E47D73FFA19B0CFAFCF8DEFAB0A12A" bold="true" box="[1331,1352,1824,1850]" pageId="3" pageNumber="4">A</emphasis>
|
||
,
|
||
<emphasis id="D3E47D73FFA19B0CFA95F8DEFA97A12A" bold="true" box="[1370,1391,1824,1850]" pageId="3" pageNumber="4">D</emphasis>
|
||
,
|
||
<emphasis id="D3E47D73FFA19B0CFF72F8BDFF2CA14D" bold="true" box="[189,212,1859,1885]" pageId="3" pageNumber="4">G</emphasis>
|
||
. Anterior part of body, lateral, dorsal and ventral views, respectively.
|
||
<emphasis id="D3E47D73FFA19B0CFBD2F8BDFBC8A14D" bold="true" box="[1053,1072,1859,1885]" pageId="3" pageNumber="4">B</emphasis>
|
||
,
|
||
<emphasis id="D3E47D73FFA19B0CFB8DF8BDFBADA14D" bold="true" box="[1090,1109,1859,1885]" pageId="3" pageNumber="4">E</emphasis>
|
||
,
|
||
<emphasis id="D3E47D73FFA19B0CFBA9F8BDFB85A14D" bold="true" box="[1126,1149,1859,1885]" pageId="3" pageNumber="4">H</emphasis>
|
||
. Midbody segments, lateral, dorsal and ventral views, respectively.
|
||
<emphasis id="D3E47D73FFA19B0CFD12F898FD0AA190" bold="true" box="[733,754,1894,1920]" pageId="3" pageNumber="4">C</emphasis>
|
||
,
|
||
<emphasis id="D3E47D73FFA19B0CFCCEF899FCEBA191" bold="true" box="[769,787,1895,1921]" pageId="3" pageNumber="4">F</emphasis>
|
||
,
|
||
<emphasis id="D3E47D73FFA19B0CFCECF899FCD6A191" bold="true" box="[803,814,1895,1921]" pageId="3" pageNumber="4">I</emphasis>
|
||
. Posterior part of body, lateral, dorsal and ventral views, respectively.
|
||
<emphasis id="D3E47D73FFA19B0CFE7BF874FE3BA1B4" bold="true" box="[436,451,1930,1956]" pageId="3" pageNumber="4">J</emphasis>
|
||
. Cross-section of a midbody segment, caudal view.
|
||
<emphasis id="D3E47D73FFA19B0CFB81F874FB9DA1B4" bold="true" box="[1102,1125,1930,1956]" pageId="3" pageNumber="4">K</emphasis>
|
||
. Limbus and adjacent structures.
|
||
<emphasis id="D3E47D73FFA19B0CFEF3F853FE81A1D7" bold="true" box="[316,377,1965,1991]" pageId="3" pageNumber="4">L–M</emphasis>
|
||
. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E12FA161FFA69B0BFF72FED0FDA2A77C" blockId="4.[189,1399,302,365]" pageId="4" pageNumber="5">together with their porosteles, as well as the remaining porosteles (male, female) or only all porosteles (female) yellowish to nearly pallid.</paragraph>
|
||
<paragraph id="E12FA161FFA69B0BFF72FE6AFE59A44E" blockId="4.[189,1399,403,606]" pageId="4" pageNumber="5">
|
||
HEAD. Except for a bare and rather densely setose clypeolabral region, very densely microdenticulatemicrotuberculate-microgranulate, interantennal isthmus almost as broad as diameter of antennal socket (
|
||
<figureCitation id="79ABBDE4FFA69B0BFF09FE27FEEAA7E4" box="[198,274,473,500]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="4" pageNumber="5">Fig. 2</figureCitation>
|
||
). Antennae long and only slightly clavate due to a somewhat higher antennomere 6,
|
||
<emphasis id="D3E47D73FFA69B0BFAE5FE24FA8EA7E3" box="[1322,1398,474,499]" italics="true" pageId="4" pageNumber="5">in situ</emphasis>
|
||
reaching behind segment 3 when stretched dorsally; only antennomere 6 with a dorso-apical group of tiny bacilliform sensilla; in length, antennomere 6> 2> 3 = 5> 1> 7; apical segment with the usual four sensory cones.
|
||
</paragraph>
|
||
<paragraph id="E12FA161FFA69B0AFF72FD7BFD1BA4BB" blockId="4.[189,1399,644,1270]" lastBlockId="5.[189,1399,267,683]" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
||
BODY. With 20 segments (male, female). In width, segment 3 = 4 <head = collum = 5–17 <2 (male), or collum <head = 3 = 4 <2 <5–17 (female); body rapidly tapering from segment 18 towards telson. Collum transversely ellipsoid, far from covering entire head from above, dorsal surface rather regularly and densely tuberculate, with two transverse rows of higher 2 + 2 and 4 + 4 tubercles in caudal half, paranota bearing 2–3 lateral teeth (
|
||
<figureCitation id="79ABBDE4FFA69B0BFDADFCEFFD57A53C" box="[610,687,785,812]" captionStart="Fig" captionStartId="2.[189,232,1965,1991]" captionTargetBox="[189,1399,897,1925]" captionTargetId="figure@2.[189,1399,897,1925]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="79ABBDE4FFA69B0BFD0FFCECFCF7A53C" box="[704,783,786,812]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="4" pageNumber="5">2A, D</figureCitation>
|
||
). Prozonae smooth and weakly shining; metazonae dull, densely microtuberculate or microgranulate all over, cerotegument well-developed, sometimes in places clothed with a crust of soil; dorsal surface usually with three regular, transverse rows of normally small tubercles or short spines; stricture very delicately striolate. Metatergum 2 with two transverse rows of spines, 2 + 2 per row, mid-dorsal pair in rear row being only slightly separated, unusually high, vertical and horn-shaped compared to very small remaining spines; paranota 2 strongly elevated, directed dorsolaterad at about 45°, each with 3–4 apical teeth (
|
||
<figureCitation id="79ABBDE4FFA69B0BFBD9FC1BFB9AA5EF" box="[1046,1122,997,1023]" captionStart="Fig" captionStartId="2.[189,232,1965,1991]" captionTargetBox="[189,1399,897,1925]" captionTargetId="figure@2.[189,1399,897,1925]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="79ABBDE4FFA69B0BFBBEFC1BFB46A5EF" box="[1137,1214,997,1023]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="4" pageNumber="5">2A, D</figureCitation>
|
||
). Tuberculation pattern persisting on metaterga 3 and 4 as well, but mid-dorsal pair of horns first considerably and then much shorter, respectively, each horn thereby carrying a subapical denticle on caudal face (
|
||
<figureCitation id="79ABBDE4FFA69B0BFADEFBD5FF2EA279" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="4" pageNumber="5">Fig. 2A, D</figureCitation>
|
||
); paranota 3–8(9) increasingly less strongly elevated to become subhorizontal (
|
||
<figureCitation id="79ABBDE4FFA69B0BFB41FBB0FB23A278" box="[1166,1243,1102,1128]" captionStart="Fig" captionStartId="2.[189,232,1965,1991]" captionTargetBox="[189,1399,897,1925]" captionTargetId="figure@2.[189,1399,897,1925]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="4" pageNumber="5">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="79ABBDE4FFA69B0BFB24FBB0FB01A278" box="[1259,1273,1102,1128]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="4" pageNumber="5">2</figureCitation>
|
||
B–C, E–F, J). Metaterga 5–19 with a faint, but visible, transverse, midtergal impression separating first transverse row of 1 + 1 (segment 5), 2 + 2 (segments 6–14) or 3 + 3 (segments 15–19) small tubercles from two further transverse rows, middle row usually consisting of slightly higher tubercles: 3 + 3 (segments 5–14) or 4(5) + 4(5) (segments 15–19); caudal row located near caudal margin, usually consisting of particularly small tubercles, yet clear enough against a microgranulate/microtuberculate background: 2 + 2 (segments 5–14) or 3(4) + 3(4) (segments 15–19). Poreless paranota usually with 4–5 distinct teeth at lateral margin, pore-bearing ones normally subtruncate, with 1–2 teeth at both fore and rear corners, and a prominent, midlength porostele (
|
||
<figureCitation id="79ABBDE4FFA79B0AFD3EFE8BFCC5A780" box="[753,829,373,400]" captionStart="Fig" captionStartId="2.[189,232,1965,1991]" captionTargetBox="[189,1399,897,1925]" captionTargetId="figure@2.[189,1399,897,1925]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="79ABBDE4FFA79B0AFC82FE88FCA3A780" box="[845,859,374,400]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">2</figureCitation>
|
||
B–C, E–F, J). Pore formula normal, ozopores always borne on somewhat mushroom-shaped porosteles. Paranota set high (mostly at about upper ¼ of body height, dorsum thus being only moderately convex), inclined caudad and increasingly clearly protruding behind rear tergal margin from segment 17 towards telson (
|
||
<figureCitation id="79ABBDE4FFA79B0AFBD0FE1EFB95A7EA" box="[1055,1133,480,506]" captionStart="Fig" captionStartId="2.[189,232,1965,1991]" captionTargetBox="[189,1399,897,1925]" captionTargetId="figure@2.[189,1399,897,1925]" captionTargetPageId="2" captionText="Fig. 1. Habitus of Campodesmoides corniger sp. nov., ♂, holotype, lateral view. Photo by J. Brecko, not to scale." figureDoi="http://doi.org/10.5281/zenodo.3785508" httpUri="https://zenodo.org/record/3785508/files/figure.png" pageId="5" pageNumber="6">Figs 1</figureCitation>
|
||
,
|
||
<figureCitation id="79ABBDE4FFA79B0AFBB0FE1EFB3FA7EA" box="[1151,1223,480,506]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">2C, F</figureCitation>
|
||
), considerably broader, and porosteles longer in male than in female. A thin dark axial line sometimes traceable through a transparent tegument. Pleurosternal carinae absent, a few small tuberculations in their place (
|
||
<figureCitation id="79ABBDE4FFA79B0AFA88FDD9FF1DA474" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">Fig. 2A</figureCitation>
|
||
). Limbus entire (
|
||
<figureCitation id="79ABBDE4FFA79B0AFE7AFDB4FDEEA474" box="[437,534,586,612]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">Fig. 2K</figureCitation>
|
||
). Epiproct short, small, subtruncate, coniform, with a pair of prominent subapical papillae (
|
||
<figureCitation id="79ABBDE4FFA79B0AFE50FD93FDD7A498" box="[415,559,621,648]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">Fig. 2C, F, I</figureCitation>
|
||
). Hypoproct nearly semi-circular with 1 + 1 caudal setae very distinctly separated and borne on minute knobs (
|
||
<figureCitation id="79ABBDE4FFA79B0AFD4CFD6FFD2DA4BB" box="[643,725,657,683]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">
|
||
Fig.
|
||
<date id="952E87A1FFA79B0AFD76FD6FFD2DA4BB" box="[697,725,657,683]" pageId="5" pageNumber="6">2I</date>
|
||
</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="B5EFF1E9FFA69B0BFF72F874FD74A1FB" ID-DOI="http://doi.org/10.5281/zenodo.3785512" ID-Zenodo-Dep="3785512" httpUri="https://zenodo.org/record/3785512/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1930,1956]" targetBox="[189,1384,1326,1890]" targetPageId="4">
|
||
<paragraph id="E12FA161FFA69B0BFF72F874FD74A1FB" blockId="4.[189,1399,1930,2027]" pageId="4" pageNumber="5">
|
||
<emphasis id="D3E47D73FFA69B0BFF72F874FEF4A1B4" bold="true" box="[189,268,1930,1956]" pageId="4" pageNumber="5">Fig. 3.</emphasis>
|
||
<taxonomicName id="2690DAE2FFA69B0BFEDBF874FDA9A1B4" authority="Vandenspiegel & Golovatch & Fiemapong, 2015" authorityName="Vandenspiegel & Golovatch & Fiemapong" authorityYear="2015" box="[276,593,1930,1956]" class="Diplopoda" family="Campodesmidae" genus="Campodesmoides" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="corniger" status="sp. nov.">
|
||
<emphasis id="D3E47D73FFA69B0BFEDBF874FDA9A1B4" box="[276,593,1930,1956]" italics="true" pageId="4" pageNumber="5">Campodesmoides corniger</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="C8D7C008FFA69B0BFD95F875FD40A1B5" box="[602,696,1931,1957]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
, ♂, holotype.
|
||
<emphasis id="D3E47D73FFA69B0BFCAEF874FC60A1B4" bold="true" box="[865,920,1930,1956]" pageId="4" pageNumber="5">A–B</emphasis>
|
||
. Left gonopod, mesal and lateral views, respectively.
|
||
<emphasis id="D3E47D73FFA69B0BFE97F853FE77A1D7" bold="true" box="[344,399,1965,1991]" pageId="4" pageNumber="5">C–E</emphasis>
|
||
. Right gonopod, mesal, lateral and ventral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="E12FA161FFA79B0AFF72FD2DFD3FA56A" blockId="5.[189,1399,723,891]" pageId="5" pageNumber="6">
|
||
STERNA. Broad, about twice as broad as coxa length, almost flat, axial impression absent (
|
||
<figureCitation id="79ABBDE4FFA79B0AFAC8FD2DFA91A4FD" box="[1287,1385,723,749]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">Fig. 2H</figureCitation>
|
||
). Gonapophyses on male coxae 2 vestigial (
|
||
<figureCitation id="79ABBDE4FFA79B0AFD0AFD08FCD0A500" box="[709,808,758,784]" captionStart="Fig" captionStartId="3.[189,231,1824,1850]" captionTargetBox="[189,1399,643,1783]" captionTargetId="figure@3.[189,1399,643,1783]" captionTargetPageId="3" captionText="Fig. 2. SEM micrographs of Campodesmoides corniger sp. nov., ♀, paratype, MRAC 22.680. A, D, G. Anterior part of body, lateral, dorsal and ventral views, respectively. B, E, H. Midbody segments, lateral, dorsal and ventral views, respectively. C, F, I. Posterior part of body, lateral, dorsal and ventral views, respectively. J. Cross-section of a midbody segment, caudal view. K. Limbus and adjacent structures. L–M. Poriferous paratergite with porostele, dorsal and lateral views, respectively. Scale bars: A–J = 0.5 mm; K–M = 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.3785510" httpUri="https://zenodo.org/record/3785510/files/figure.png" pageId="5" pageNumber="6">Fig. 2G</figureCitation>
|
||
). Legs long to very long, about 0.9–1.0 or 1.5– 1.6 times as long as midbody height (female and male, respectively), very slender; in length, femur> tarsus> prefemur = postfemur = tibia = coxa; claw very small, very slightly curved; ventral surface of tarsi densely setose, but forming no brushes.
|
||
</paragraph>
|
||
<paragraph id="E12FA161FFA79B0AFF72FC5CFB23A30E" blockId="5.[189,1399,930,1311]" pageId="5" pageNumber="6">
|
||
GONOPOD. Aperture transversely ovoid, large, its lateral and posterior edges slightly elevated, fully concealing gonocoxae and bases of telopodites. Gonopod coxites medium-sized, subcylindrical, fused at base to a small membranous sternal remnant, poorly setose distodorsally, including a pair of very closely placed, distalmost, particularly long setae. Cannulae slender, without peculiarities (
|
||
<figureCitation id="79ABBDE4FFA79B0AFB27FBF2FAD4A236" box="[1256,1324,1036,1062]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[189,1384,1326,1890]" captionTargetId="figure@4.[189,1399,1326,1890]" captionTargetPageId="4" captionText="Fig. 3. Campodesmoides corniger sp. nov., ♂, holotype. A–B. Left gonopod, mesal and lateral views, respectively. C–E. Right gonopod, mesal, lateral and ventral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3785512" httpUri="https://zenodo.org/record/3785512/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
|
||
A–B). Telopodites
|
||
<emphasis id="D3E47D73FFA79B0AFE80FBCFFE62A25A" box="[335,410,1073,1098]" italics="true" pageId="5" pageNumber="6">in situ</emphasis>
|
||
directed forward, held subparallel to each other, suberect, not crossing each other mesally. Prefemoral (= densely setose) portion (pf) subcylindrical, very long and slender, untwisted, almost not enlarged towards an oblique apical cingulum (c) delimiting it from femorite, with a long, slightly curved, distodorsal process (pfp). Femorite clearly twisted, lamellar, about half as long as pf, forming a large, rounded, apical lobe = solenophore (sph) to protect a smaller, parabasal lobe (lo) carrying terminal part of seminal groove on its distal rib. Seminal groove first running fully on mesal side of pf, then moving laterad above c onto distal rib of lo to end at tip of lo (
|
||
<figureCitation id="79ABBDE4FFA79B0AFB9FFAFAFB6CA30E" box="[1104,1172,1284,1310]" captionStart="Fig" captionStartId="4.[189,232,1930,1956]" captionTargetBox="[189,1384,1326,1890]" captionTargetId="figure@4.[189,1399,1326,1890]" captionTargetPageId="4" captionText="Fig. 3. Campodesmoides corniger sp. nov., ♂, holotype. A–B. Left gonopod, mesal and lateral views, respectively. C–E. Right gonopod, mesal, lateral and ventral views, respectively. Scale bars = 0.2 mm. Abbreviations are explained in the text." figureDoi="http://doi.org/10.5281/zenodo.3785512" httpUri="https://zenodo.org/record/3785512/files/figure.png" pageId="5" pageNumber="6">Fig. 3</figureCitation>
|
||
C–E).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A98AF2EAFFA79B0AFF72FAB8FCDAA3A2" pageId="5" pageNumber="6" type="discussion">
|
||
<paragraph id="E12FA161FFA79B0AFF72FAB8FEDCA370" blockId="5.[189,292,1350,1376]" box="[189,292,1350,1376]" pageId="5" pageNumber="6">
|
||
<heading id="BA67160DFFA79B0AFF72FAB8FEDCA370" bold="true" box="[189,292,1350,1376]" fontSize="11" level="3" pageId="5" pageNumber="6" reason="3">
|
||
<emphasis id="D3E47D73FFA79B0AFF72FAB8FEDCA370" bold="true" box="[189,292,1350,1376]" pageId="5" pageNumber="6">Remark</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="E12FA161FFA79B0AFF72FA8AFCDAA3A2" blockId="5.[189,1399,1396,1459]" pageId="5" pageNumber="6">
|
||
This new genus and species extends the range of
|
||
<taxonomicName id="2690DAE2FFA79B0AFCC7FA8AFC34A39E" authorityName="Cook" authorityYear="1895" box="[776,972,1396,1422]" class="Diplopoda" family="Campodesmidae" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="family">Campodesmidae</taxonomicName>
|
||
to
|
||
<collectingCountry id="9987E1F1FFA79B0AFC3CFA8BFB95A39F" box="[1011,1133,1397,1423]" name="Cameroon" pageId="5" pageNumber="6">Cameroon</collectingCountry>
|
||
in the east, the family also being recorded in that country for the first time.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |