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<document id="A39F136581964C9BAEA851F032F1AD64" ID-DOI="10.1206/3770.2" ID-ISSN="0003-0082" ID-Zenodo-Dep="5361898" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630237122464" checkinUser="carolina" docAuthor="Grimaldi, David, Engel, Michael S., Nascimbene, Paul c. &amp; Singh, Hukam" docDate="2013" docId="4B0C87DE5744D30C3DE3FF58FC7ED00B" docLanguage="en" docName="N3770.pdf" docOrigin="American Museum Novitates 2013 (3770)" docSource="http://www.bioone.org/doi/abs/10.1206/3770.2" docStyle="DocumentStyle:26C3580A17D4BBE58BD8D8AC6AE09AC4.6:AmMusNovit.2010-.journal_article" docStyleId="26C3580A17D4BBE58BD8D8AC6AE09AC4" docStyleName="AmMusNovit.2010-.journal_article" docStyleVersion="6" docTitle="Spiloconis glaesaria Meinander" docType="treatment" docVersion="5" lastPageNumber="4" masterDocId="B735FFA65747D30F3C08FFB9FF97D36F" masterDocTitle="Coniopterygidae (Neuroptera: Aleuropteryginae) in amber from the Eocene of India and the Miocene of Hispaniola" masterLastPageNumber="20" masterPageNumber="1" pageNumber="4" updateTime="1699194493376" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="74B656859A49850142E9120D8E1DCF27">Coniopterygidae (Neuroptera: Aleuropteryginae) in amber from the Eocene of India and the Miocene of Hispaniola</mods:title>
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<mods:namePart id="943614069B5496F4E2E6A99437E997E9">Grimaldi, David</mods:namePart>
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<mods:namePart id="9DBFDBCF19DA6BB345C39F4D0D6717BF">Engel, Michael S.</mods:namePart>
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<mods:namePart id="B70A638DE003D80EEA1BE8298B886476">Nascimbene, Paul c.</mods:namePart>
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<mods:namePart id="60A0E726B4B99A33033070528298D8D4">Singh, Hukam</mods:namePart>
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<subSubSection id="8BBF65435744D30C3DE3FF58FCC3D394" box="[491,852,225,251]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="C31A36C85744D30C3DE3FF58FCC3D394" blockId="3.[491,852,225,251]" box="[491,852,225,251]" pageId="3" pageNumber="4">
<heading id="985281A45744D30C3DE3FF58FCC3D394" box="[491,852,225,251]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="4" reason="5">
<taxonomicName id="04A54D4B5744D30C3DE3FF58FCC3D394" ID-CoL="4Z3MK" authority="Meinander" authorityName="Meinander" box="[491,852,225,251]" class="Insecta" family="Coniopterygidae" genus="Spiloconis" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="glaesaria">
<emphasis id="F1D1EADA5744D30C3DE3FF58FD5CD394" box="[491,715,225,251]" italics="true" pageId="3" pageNumber="4">Spiloconis glaesaria</emphasis>
Meinander
</taxonomicName>
</heading>
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<subSubSection id="8BBF65435744D30C3E28FEBEFCB7D24E" box="[544,800,263,289]" pageId="3" pageNumber="4" type="description">
<paragraph id="C31A36C85744D30C3E28FEBEFCB7D24E" blockId="3.[120,885,263,326]" box="[544,800,263,289]" pageId="3" pageNumber="4">
<heading id="985281A45744D30C3E28FEBEFCB7D24E" box="[544,800,263,289]" centered="true" fontSize="36" level="3" pageId="3" pageNumber="4" reason="7">
<figureCitation id="5B9E2A4D5744D30C3E28FEBEFD54D24E" box="[544,707,263,289]" captionStart="FIGURE 1" captionStartId="4.[120,206,1546,1567]" captionTargetBox="[120,1224,224,1520]" captionTargetId="figure-11@4.[120,1224,224,1520]" captionTargetPageId="4" captionText="FIGURE 1. Photomicrographs of Aleuropteryginae in amber. AD: In Miocene amber from the Dominican Republic. A. Spiloconis glaesaria Meinander, holotype, AMNH DR10-59. B. S. glaesaria, AMNH DR14-1094. C. Spiloconis oediloma Engel and Grimaldi, AMNH DR14-1427. D. Neoconis paleocaribis Grimaldi and Engel, new species, holotype, AMNH DR15-29. EF: Spiloconis eominuta Grimaldi and Engel, new species, in Early Eocene Cambay amber from India; both are males. E. Holotype, BSIP Tad 316D. F. AMNH Tad 316B." figureDoi="http://doi.org/10.5281/zenodo.5361900" httpUri="https://zenodo.org/record/5361900/files/figure.png" pageId="3" pageNumber="4">Figures 1A, B</figureCitation>
;
<figureCitation id="5B9E2A4D5744D30C3EC7FEBEFD66D24E" box="[719,753,263,289]" captionStart="FIGURE 2" captionStartId="5.[120,206,1629,1650]" captionTargetBox="[335,1070,228,1598]" captionTargetId="figure-9@5.[149,1192,227,1602]" captionTargetPageId="5" captionText="FIGURE 2. Frontal views of head of fossil species of Spiloconis. AB: Dominican amber; C: Cambay amber. A. Spiloconis oediloma Engel and Grimaldi, AMNH DR15-421. B. S. glaesaria Meinander, holotype, AMNH DR10-59. Dashed line above head approximates dorsal margin. C. S. eominuta Grimaldi and Engel, new species, BSIP Tad 316D (reconstructed, based on right half of head). Note differences in scale between both A and B and C." figureDoi="http://doi.org/10.5281/zenodo.5361902" httpUri="https://zenodo.org/record/5361902/files/figure.png" pageId="3" pageNumber="4">2B</figureCitation>
;
<figureCitation id="5B9E2A4D5744D30C3EF5FEBEFCB7D24E" box="[765,800,263,289]" captionStart="FIGURE 4" captionStartId="7.[120,206,1615,1636]" captionTargetBox="[128,1218,227,1588]" captionTargetId="figure-9@7.[120,1224,224,1589]" captionTargetPageId="7" captionText="FIGURE 4. Forewings of fossil species of Spiloconis in Dominican amber. A. Spiloconis glaesaria Meinander, AMNH DR14-1094, also showing detail of costal setae. B. S. oediloma Engel and Grimaldi, holotype, AMNH DR14-1097. C. S. oediloma, AMNH DR15-421 (slightly reconstructed from fractured specimen). Costal setae omitted in B, C, and in figure 5." figureDoi="http://doi.org/10.5281/zenodo.5361906" httpUri="https://zenodo.org/record/5361906/files/figure.png" pageId="3" pageNumber="4">4A</figureCitation>
</heading>
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<paragraph id="C31A36C85744D30C3C70FE97FCE2D229" blockId="3.[120,885,263,326]" box="[120,885,302,326]" pageId="3" pageNumber="4">
<treatmentCitationGroup id="E3B511E65744D30C3C70FE97FCE2D229" box="[120,885,302,326]" pageId="3" pageNumber="4">
<treatmentCitation id="420410D95744D30C3C70FE97FDBFD229" author="Meinander, M." box="[120,552,302,326]" page="33" pageId="3" pageNumber="4" year="1998">
<taxonomicName id="04A54D4B5744D30C3C70FE97FDBFD229" ID-CoL="4Z3MK" authority="Meinander, 1998: 33" authorityName="Meinander" authorityPageNumber="33" authorityYear="1998" box="[120,552,302,326]" class="Insecta" family="Coniopterygidae" genus="Spiloconis" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="glaesaria">
<emphasis id="F1D1EADA5744D30C3C70FE97FED3D229" box="[120,324,302,326]" italics="true" pageId="3" pageNumber="4">Spiloconis glaesaria</emphasis>
<bibRefCitation id="A7344B395744D30C3D43FE97FDBFD229" author="Meinander, M." box="[331,552,302,326]" pageId="3" pageNumber="4" pagination="33 - 36" refId="ref8444" refString="Meinander, M. 1998. Coniopterygidae (Neuroptera) in amber from the Dominican Republic. Journal of Neuropterology 1: 33 - 36." type="journal article" year="1998">Meinander, 1998: 33</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
<treatmentCitation id="420410D95744D30C3E3BFE97FCE6D229" author="Engel, M. S. &amp; D. A. Grimaldi" box="[563,881,302,326]" page="19" pageId="3" pageNumber="4" year="2007">
<bibRefCitation id="A7344B395744D30C3E3BFE97FCE6D229" author="Engel, M. S. &amp; D. A. Grimaldi" box="[563,881,302,326]" pageId="3" pageNumber="4" pagination="1 - 58" refId="ref7804" refString="Engel, M. S., and D. A. Grimaldi. 2007. The neuropterid fauna of Dominican and Mexican amber (Neuropterida: Megaloptera, Neuroptera). American Museum Novitates 3587: 1 - 58." type="journal article" year="2007">Engel and Grimaldi, 2007: 19</bibRefCitation>
</treatmentCitation>
.
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</paragraph>
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<paragraph id="C31A36C85744D30C3CA0FEDDFC93D156" blockId="3.[120,1225,356,868]" pageId="3" pageNumber="4">
DIAGNOSIS: Oral margin in frontal view broad (width 0.7× the distance between compound eyes), much of frontal portion of head membranous/lightly sclerotized, most of it covered with fine setulae; dorsal margin of head barely protruding (cf.
<taxonomicName id="04A54D4B5744D30C3F92FE16FE8AD280" authority="Engel and Grimaldi" authorityName="Engel and Grimaldi" class="Insecta" family="Coniopterygidae" genus="Spiloconis" kingdom="Animalia" order="Neuroptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="oediloma">
<emphasis id="F1D1EADA5744D30C3F92FE16FBE9D2A6" box="[922,1150,431,457]" italics="true" pageId="3" pageNumber="4">Spiloconis oediloma</emphasis>
Engel and Grimaldi
</taxonomicName>
); wing with
<emphasis id="F1D1EADA5744D30C3DBCFE6CFE4BD281" box="[436,476,469,494]" italics="true" pageId="3" pageNumber="4">r-rs</emphasis>
proximal to fork of
<emphasis id="F1D1EADA5744D30C3EDCFE6CFCD5D29C" box="[724,834,469,499]" italics="true" pageId="3" pageNumber="4">
R
<subScript id="5F21348D5744D30C3EEEFE5BFC93D29C" attach="left" box="[742,772,482,499]" fontSize="7" pageId="3" pageNumber="4">2+3</subScript>
R
<subScript id="5F21348D5744D30C3F2CFE5BFCD5D29C" attach="left" box="[804,834,482,499]" fontSize="7" pageId="3" pageNumber="4">4+5</subScript>
</emphasis>
(i.e.,
<emphasis id="F1D1EADA5744D30C3F8FFE6CFC38D281" box="[903,943,469,494]" italics="true" pageId="3" pageNumber="4">r-rs</emphasis>
connected to
<emphasis id="F1D1EADA5744D30C3854FE6CFBE0D281" box="[1116,1143,469,494]" italics="true" pageId="3" pageNumber="4">Rs</emphasis>
); L/W basal discal cell 3.2; antenna with 2122 flagellomeres (2324 antennomeres), size of basal flagellomere nearly equal to that of other flagellomeres.
</paragraph>
</subSubSection>
<subSubSection id="8BBF65435744D30C3CA0FDFCFC7ED00B" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="C31A36C85744D30C3CA0FDFCFC7ED00B" blockId="3.[120,1225,356,868]" pageId="3" pageNumber="4">
<materialsCitation id="73CD3C955744D30C3CA0FDFCFBCAD131" box="[168,1117,580,607]" collectionCode="TYPE, AND" pageId="3" pageNumber="4" specimenCount="2" typeStatus="holotype">
<collectionCode id="A5B4AE0D5744D30C3CA0FDFCFEB6D131" box="[168,289,580,607]" pageId="3" pageNumber="4">TYPE AND</collectionCode>
OTHER MATERIAL: Known only from Dominican amber.
<typeStatus id="1C1E886A5744D30C3FE5FDFDFBCAD131" box="[1005,1117,580,606]" pageId="3" pageNumber="4" type="holotype">Holotype</typeStatus>
</materialsCitation>
,
<materialsCitation id="73CD3C955744D30C3864FDFDFC72D00B" collectionCode="AMNH" pageId="3" pageNumber="4" specimenCode="DR14-1094" specimenCount="1" typeStatus="holotype">
<collectionCode id="A5B4AE0D5744D30C3864FDFDFB50D131" box="[1132,1223,580,606]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH</collectionCode>
<specimenCode id="93039EB35744D30C3C70FDD3FE96D1EB" box="[120,257,618,644]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">DR14-1094</specimenCode>
, in clear yellow amber 9 × 4 ×
<quantity id="045D9B2D5744D30C3E67FDD0FD22D1EB" box="[623,693,617,644]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
in size, embedded in EpoTek 301 resin, 12 × 3 ×
<quantity id="045D9B2D5744D30C3CA4FD36FF60D1C6" box="[172,247,655,682]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="4" unit="mm" value="5.0">5 mm</quantity>
. The amber contains many bubbles, and the
<typeStatus id="1C1E886A5744D30C3F15FD36FC12D1C6" box="[797,901,655,681]" pageId="3" pageNumber="4" type="holotype">holotype</typeStatus>
is best observed ventrally. Meinander (1988) stated that the body of the specimen was covered with “wax” (a fine layer of which indeed coats most modern dustywings), but the fossil is actually covered with a milky froth, a preservational artifact common among inclusions in amber. Another specimen was reported (
<collectionCode id="A5B4AE0D5744D30C3CE5FC9DFEDDD051" box="[237,330,804,830]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">AMNH</collectionCode>
<specimenCode id="93039EB35744D30C3D5BFC9DFE4BD051" box="[339,476,804,830]" collectionCode="AMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="3" pageNumber="4">DR14-1094</specimenCode>
) with a photograph, in
<bibRefCitation id="A7344B395744D30C3EFEFC9DFBAFD051" author="Engel, M. S. &amp; D. A. Grimaldi" box="[758,1080,804,830]" pageId="3" pageNumber="4" pagination="1 - 58" refId="ref7804" refString="Engel, M. S., and D. A. Grimaldi. 2007. The neuropterid fauna of Dominican and Mexican amber (Neuropterida: Megaloptera, Neuroptera). American Museum Novitates 3587: 1 - 58." type="journal article" year="2007">Engel and Grimaldi (2007)</bibRefCitation>
. It is a wellpreserved female in a small, clear yellow square of amber, 4 × 4 ×
<quantity id="045D9B2D5744D30C3F8FFCF0FC72D00B" box="[903,997,841,868]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="4" unit="mm" value="1.5">1.5 mm</quantity>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
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