321 lines
29 KiB
XML
321 lines
29 KiB
XML
<document ID-CLB-Dataset="55588" ID-DOI="http://dx.doi.org/10.3897/alpento.6.79598" ID-GBIF-Dataset="310f1b04-d893-41f3-9570-a115045628c1" ID-Pensoft-Pub="2535-0889-6-7" ID-Pensoft-UUID="A48E84FA8FF753CF979C8BAEE7950B6D" ID-ZooBank="75DB5DB6C3E54B3C9B5C15722723ED3B" ModsDocID="2535-0889-6-7" checkinTime="1652223128647" checkinUser="pensoft" docAuthor="Balke, Michael, Bilton, David T., Garcia, Mauricio, Viloria, Angel L., Villastrigo, Adrian & Hendrich, Lars" docDate="2022" docId="4AA9D981254655039E4C77AD3F1D809F" docLanguage="en" docName="AlpEntomol 6: 7-12" docOrigin="Alpine Entomology 6" docPubDate="2022-05-10" docSource="http://dx.doi.org/10.3897/alpento.6.79598" docTitle="Liodessus meridensis Balke & Bilton & García & Viloria & Villastrigo & Hendrich 2022, sp. nov." docType="treatment" docUuid="CB562F9F-95AD-4537-ACA5-305AA84CB53C" docUuidSource="ZooBank" docVersion="5" id="A48E84FA8FF753CF979C8BAEE7950B6D" lastPageNumber="7" masterDocId="A48E84FA8FF753CF979C8BAEE7950B6D" masterDocTitle="Two new high Andean species of Liodessus diving beetles from Venezuela (Coleoptera, Dytiscidae, Bidessini)" masterLastPageNumber="12" masterPageNumber="7" pageNumber="7" updateTime="1732939330256" updateUser="ExternalLinkService">
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<mods:mods id="57B1B8C715E9AE3A3FF154DDC776D5B3" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A61E8E9456E30A6479999FFA0E31A976">
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<mods:title id="0E76A1DD7A78A4B710BE477D82FDA393">Two new high Andean species of Liodessus diving beetles from Venezuela (Coleoptera, Dytiscidae, Bidessini)</mods:title>
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<mods:name id="6782CFAB1EAADDF4DBFFB57FE2C631D4" type="personal">
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<mods:role id="C3728A389FEE52C52E1673CB5A7BB32E">
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<mods:roleTerm id="19E99F76337FC16C6FB3BED234206398">Author</mods:roleTerm>
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<mods:namePart id="ABA8771557F7E3F5BC15BF72F9172200">Balke, Michael</mods:namePart>
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<mods:nameIdentifier id="06DF2347C34841058E0BF8FB483DE16B" type="ORCID">https://orcid.org/0000-0002-3773-6586</mods:nameIdentifier>
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<mods:affiliation id="AA7205CDFE6A6596D642030CFBEACAD1">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany and GeoBioCenter, Ludwig Maximilians University, Munich, Germany</mods:affiliation>
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<mods:nameIdentifier id="05BBB900D0583234470839310109914C" type="email">balke.m@snsb.de</mods:nameIdentifier>
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</mods:name>
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<mods:name id="F99662EA733BE6893D01A4616DBF3EA8" type="personal">
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<mods:roleTerm id="4C9C6E62A4B482E57D768C1A047A21C6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FBB097600F2E4D5A83548C998DE0E369">Bilton, David T.</mods:namePart>
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<mods:affiliation id="A4499DED670BD45418DB17C4ACFEF59C">School of Biological and Marine Sciences, University of Plymouth, Drake Circus, Plymouth, UK & Department of Zoology, University of Johannesburg, PO Box 524, Auckland Park, 2006 Johannesburg, South Africa</mods:affiliation>
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</mods:name>
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<mods:name id="4BADE796BCE6CA4C83F64E3FD55F4A64" type="personal">
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<mods:role id="47586E8B9647803D704BE94632B598CF">
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<mods:roleTerm id="FE078014B2D3F95B32B759FE3AC18F2F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="95657625BF7CDCB17A03B4B60FA9063C">Garcia, Mauricio</mods:namePart>
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<mods:affiliation id="DA50A2465DBF19B35170807820620948">Centro de Investigaciones Biologicas, Facultad de Humanidades y Educacion, Universidad del Zulia, Apartado 526, Maracaibo A- 4001, Estado Zulia, Venezuela</mods:affiliation>
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</mods:name>
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<mods:name id="A90C6656CE59E51285146D91B22E1BFB" type="personal">
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<mods:role id="B9ECC5ED60044B69CB22E7E9A3690242">
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<mods:roleTerm id="FE3991577C0EF78E908D7459C521736F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="A85EA97ACBFEF2AAEAB8CA559E2C8DAF">Viloria, Angel L.</mods:namePart>
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<mods:affiliation id="91AEC20F7C7E144FFB11A59647BE9353">Centro de Ecologia, Instituto Venezolano de Investigaciones Cientificas (IVIC), km 11 carretera Panamericana, Altos de Pipe, edo. Miranda 1204, Venezuela</mods:affiliation>
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</mods:name>
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<mods:name id="EF4476111346913904C05D9C58B27795" type="personal">
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<mods:role id="2EC37845A6CA97798FEB6B37820604D0">
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<mods:roleTerm id="6312283187CD2F283910266E9442F25F">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E04BD9FC9636A6E8368F316422BE1490">Villastrigo, Adrian</mods:namePart>
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<mods:nameIdentifier id="8574ADFB8BBFABD93164372DDF0BED5D" type="ORCID">https://orcid.org/0000-0002-3531-0821</mods:nameIdentifier>
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<mods:affiliation id="B9C0EAED8054CCB3C994DFA213F848E1">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany</mods:affiliation>
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</mods:name>
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<mods:name id="526F6C24CD43904401F11BA45CDD3CBB" type="personal">
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<mods:roleTerm id="CD2E1A812B0D3B7E90EB72B071F4FC07">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="65B07DD0C10A594464B339155A4A9841">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="A18729431EC0828829FB76C4FCE1D435">SNSB-Zoologische Staatssammlung Muenchen, Muenchhausenstrasse 21, D- 81247 Munich, Germany</mods:affiliation>
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<mods:typeOfResource id="535539177CC3DEAABCA8F049E5476135">text</mods:typeOfResource>
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<mods:titleInfo id="6AE600326F1F360B32CC489D23C099D6">
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<mods:title id="93C7A63C04437341B4DA7D293EDA9ABD">Alpine Entomology</mods:title>
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</mods:titleInfo>
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<mods:part id="C126289BD6C1A1143CB6CBCCA10989B9">
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<mods:date id="88352A73032DB46C73E9AF8B369E7967">2022</mods:date>
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<mods:number id="274151F87FC936D20230185FD2F52637">2022-05-10</mods:number>
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</mods:detail>
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<mods:detail id="DDD1A3882CDB0D827083AEB7701DA0C2" type="volume">
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<mods:number id="7179A954704D924FEB22EB2017F6F880">6</mods:number>
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</mods:detail>
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<mods:extent id="2AB21ADCC8E60B54C8C5FF1E9DBB76EE" unit="page">
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<mods:start id="A3F44220B6A30008C7857AC0F3F99AEC">7</mods:start>
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<mods:end id="35EA75787954CCAEB32EC9B267C06CE1">12</mods:end>
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</mods:extent>
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<mods:location id="B6C430BC5F94DD13109866664F010626">
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<mods:url id="F3550DCAB304E36B9771C160EA84DB7F">http://dx.doi.org/10.3897/alpento.6.79598</mods:url>
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</mods:location>
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<mods:classification id="98181AA04FD36866472ABD1C1142A6F9">journal article</mods:classification>
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<mods:identifier id="3758D191DEFCE1C50EE8892A78076181" type="DOI">http://dx.doi.org/10.3897/alpento.6.79598</mods:identifier>
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<mods:identifier id="C86BF10B468EEC061B02700AF17093C6" type="ZooBank">75DB5DB6C3E54B3C9B5C15722723ED3B</mods:identifier>
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</mods:mods>
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<treatment id="4AA9D981254655039E4C77AD3F1D809F" ID-GBIF-Taxon="195298556" LSID="urn:lsid:zoobank.org:act:CB562F9F-95AD-4537-ACA5-305AA84CB53C" httpUri="http://treatment.plazi.org/id/4AA9D981254655039E4C77AD3F1D809F" lastPageNumber="7" pageId="0" pageNumber="7" scope_family="Dytiscidae" scope_order="Coleoptera" scope_tribe="Bidessini">
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<subSubSection id="7BBEA706D004B939DAD9B330BF99B3A8" pageId="0" pageNumber="7" type="nomenclature">
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<paragraph id="E127D0EE7EFFD4CE1CE6239E08065AD1" pageId="0" pageNumber="7">
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<taxonomicName id="6908B311125FE0471D6B57F074F9B7E5" LSID="http://zoobank.org/CB562F9F-95AD-4537-ACA5-305AA84CB53C" authority="Balke & Bilton & García & Viloria & Villastrigo & Hendrich, 2022" authorityName="Balke & Bilton & García & Viloria & Villastrigo & Hendrich" authorityYear="2022" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus meridensis" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="species" species="meridensis" status="sp. nov.">Liodessus meridensis</taxonomicName>
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<taxonomicNameLabel id="5BC989A68694892333A003CD22858462" pageId="0" pageNumber="7">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="3FEB630910EFA1DCB76008F152B0ECB3" pageId="0" pageNumber="7" type="description">
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<paragraph id="693B2A65A3BD72BA4F13C1FBC2EC8410" pageId="0" pageNumber="7">
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<figureCitation id="ABC4486654776C8167E72DB603FDAA27" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">Figs 1A</figureCitation>
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<figureCitation id="989C20F556BC90EFE7AD19F26FDBDEE2" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Liodessus spp. nov. Detail of head in dorsal view. L. meridensis (A, B); L. venezuelanus (C, D)." figureDoi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7">, 2A, B</figureCitation>
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<figureCitation id="5C3B7AA69CB68385FEAADFAD1B8A470F" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Liodessus spp. nov. Median lobe ventral and lateral views; parameres outer and inner surfaces, L. meridensis (A); L. venezuelanus (B). Scale bar: 0.5 mm." figureDoi="10.3897/alpento.6.79598.figure3" httpUri="https://binary.pensoft.net/fig/682026" pageId="0" pageNumber="7">, 3A</figureCitation>
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<figureCitation id="BCEE9A250657A42447C5E3C1F73ECCEA" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Liodessus spp. nov. near Gavidia, Venenzuela, where both new species were collected (collecting event VZ 10 - 0123 - 03 A). Overview (A); detail of shallow water at the edge (B)." figureDoi="10.3897/alpento.6.79598.figure4" httpUri="https://binary.pensoft.net/fig/682027" pageId="0" pageNumber="7">, 4</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="D770C8E6ABDDB0DE4826D5ED647C74AD" pageId="0" pageNumber="7" type="type locality">
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<paragraph id="70F4A80442BFFF4CB435F815FC2A018B" pageId="0" pageNumber="7">Type locality.</paragraph>
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<paragraph id="9CC4B2D5BF218DDE87655CD6CFECF70F" pageId="0" pageNumber="7">
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Laguna de
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<normalizedToken id="7C372B52E04E80B303AC77AD9C94F949" originalValue="Mucabají">Mucabaji</normalizedToken>
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,
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<normalizedToken id="2BD341D9AD14557198B12079BF72876A" originalValue="Mérida">Merida</normalizedToken>
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, Venezuela.
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</paragraph>
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</subSubSection>
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<subSubSection id="5233C86BBE3C249A5D31F5B4EDCBE487" pageId="0" pageNumber="7" type="type material">
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<paragraph id="ACDE8773C52B6B0A2ACB4176203E3A62" pageId="0" pageNumber="7">Type material.</paragraph>
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<paragraph id="166E842486EA0F6D12732B83400B4397" pageId="0" pageNumber="7">
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<emphasis id="8662E43133A1D9528A7C3555F0A13F7E" bold="true" italics="true" pageId="0" pageNumber="7">Holotype</emphasis>
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: Male: Venezuela;
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<normalizedToken id="E5778021B12F4B19CDDAAAE5ACE83FC8" originalValue="Mérida">Merida</normalizedToken>
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, Laguna de
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<normalizedToken id="5E73E6D86CE030AA0E148080FB040AC9" originalValue="Mucabají">Mucabaji</normalizedToken>
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, 3,500 m, i.2004, 8.7964, -70.8255,
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<normalizedToken id="6D15594FBA2A7F7D6F5A2811685587F4" originalValue="García">Garcia</normalizedToken>
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& Balke (MIZA).
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<emphasis id="003D841B865F2CE74994FE85CD75F849" bold="true" italics="true" pageId="0" pageNumber="7">Paratypes</emphasis>
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: 13 exs same label data as holotype (MIZA, ZSM); 47 exs Venezuela:
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<normalizedToken id="EF48AE91F6290E92AC85EC698EE43830" originalValue="Mérida">Merida</normalizedToken>
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, ca. 5 km E Gavidia, 3,600 m, 23.I.2012,
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<geoCoordinate id="2EBC3D6A4BE92051979AED4D3C7DCF05" degrees="8.6895" direction="north" orientation="latitude" precision="5" value="8.6895">8.6895</geoCoordinate>
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,
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<geoCoordinate id="D74A2F2394449B6E89922EA6CCEB62D6" degrees="70.8817" direction="west" orientation="longitude" precision="5" value="-70.8817">-70.8817</geoCoordinate>
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, Short, Arias & Gustafson, lagoon margin, VZ12-0123-03A (MIZA, SEMC).
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</paragraph>
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</subSubSection>
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<subSubSection id="AF662A415781963E2BDC85D07F9EDEF7" pageId="0" pageNumber="7" type="description">
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<paragraph id="BEE9A0B26E5A51B1CC6E4E19053583A0" pageId="0" pageNumber="7">Description.</paragraph>
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<paragraph id="CC9ACA81E4BEE4D962EF2CAFC6056238" pageId="0" pageNumber="7">
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Habitus with only slight discontinuity between pronotum and elytron (Fig.
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<figureCitation id="F9C344173F237040A3DE57A59C182191" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
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), pronotum widest before base and more parallel sided laterally (Fig.
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<figureCitation id="52E187EA8E2D7BF3637229D109D8D678" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
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). Total length (TL) 2.1 mm; length without head (TL-h) 1.9 mm; maximum width (MW) 0.9 mm.
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</paragraph>
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<caption id="CAB71067E19685DF5385957BFD160D35" doi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7" start="Figure 1" startId="F1">
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<paragraph id="D4A75683DDA57803884F42D12EE0620B" pageId="0" pageNumber="7">
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<emphasis id="839CB0E2CAA9BFB2F70BD0AADDB768BF" bold="true" pageId="0" pageNumber="7">Figure 1.</emphasis>
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Dorsal habitus of
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<taxonomicName id="35DB2CFF8C3E24DA330EB0BE488E5AAB" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
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<emphasis id="8AAA44D38206D963BCF43ECA81484383" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
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</taxonomicName>
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spp. nov.
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<taxonomicName id="0F0856BCACEE972708BCA7AE81B8ED14" lsidName="L. meridensis" pageId="0" pageNumber="7" rank="species" species="meridensis">
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<emphasis id="6B2AA0C01CE3BE31460A030D2A39E5B7" italics="true" pageId="0" pageNumber="7">L. meridensis</emphasis>
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</taxonomicName>
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(
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<emphasis id="64E2C81D8745709D283062E219FF61B1" bold="true" pageId="0" pageNumber="7">A</emphasis>
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);
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<taxonomicName id="17BEAFBD3A4696F2FB126598D3637477" lsidName="L. venezuelanus" pageId="0" pageNumber="7" rank="species" species="venezuelanus">
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<emphasis id="B8D052ADF7A44520B14F5C74F2290FD0" italics="true" pageId="0" pageNumber="7">L. venezuelanus</emphasis>
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</taxonomicName>
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(
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<emphasis id="3560AA694F4DC24274F0CCB028FF7FE3" bold="true" pageId="0" pageNumber="7">B</emphasis>
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). Scale bar: 2 mm.
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</paragraph>
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</caption>
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<paragraph id="0E676BF2F90BDF1E211FF83C1405A3A8" pageId="0" pageNumber="7">
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<emphasis id="B439B7BC7559A84B3D2C3E4DB394B9F4" bold="true" pageId="0" pageNumber="7">
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<emphasis id="98B4119B43A0A59BF48E9759B3E318BC" italics="true" pageId="0" pageNumber="7">Color</emphasis>
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.
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</emphasis>
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Very dark brown to blackish dorsally and ventrally, basal antennomeres, base of meso- and metatibia contrastingly of lighter color (Figs
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<figureCitation id="3F998AE0C801ED4BF442F3246D2763E3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
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,
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<figureCitation id="53F19D1E9C5B18AAE24C32275F763A62" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Liodessus spp. nov. Detail of head in dorsal view. L. meridensis (A, B); L. venezuelanus (C, D)." figureDoi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7">2A, B</figureCitation>
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).
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</paragraph>
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<caption id="8BFABEBD48A8234FABCBB33F848A965C" doi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7" start="Figure 2" startId="F2">
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<paragraph id="6242193A35CBAE2E1E19EE2B21C06FC5" pageId="0" pageNumber="7">
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<emphasis id="DDD7DB9ECC51DE9095DB9A4D544908B3" bold="true" pageId="0" pageNumber="7">Figure 2.</emphasis>
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<taxonomicName id="9E0B751A937891106CB36A39389859C1" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="B04D68ED9F4A709161A54EEDFA961351" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
|
|
</taxonomicName>
|
|
spp. nov. Detail of head in dorsal view.
|
|
<taxonomicName id="3A6BA061481B293B73340F7E3477DE75" lsidName="L. meridensis" pageId="0" pageNumber="7" rank="species" species="meridensis">
|
|
<emphasis id="24F0DF41D9D42AB83EE451FC7DEA4E64" italics="true" pageId="0" pageNumber="7">L. meridensis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="0029F4E84AB0AE7E60AE95E12B9A2E17" bold="true" pageId="0" pageNumber="7">A, B</emphasis>
|
|
);
|
|
<taxonomicName id="FA2114203831BDCB9AAEC3EB04206CCC" lsidName="L. venezuelanus" pageId="0" pageNumber="7" rank="species" species="venezuelanus">
|
|
<emphasis id="65707D81590C5C7F02DC60F919C9C13B" italics="true" pageId="0" pageNumber="7">L. venezuelanus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="32F4220FEE1DE6A3F02FD47E2540AFA4" bold="true" pageId="0" pageNumber="7">C, D</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="B24BE24BC89C703E4E21AFFF94E383F9" pageId="0" pageNumber="7">
|
|
<emphasis id="779DBC1011486DF0FB8B4A69EF06D1ED" bold="true" pageId="0" pageNumber="7">
|
|
<emphasis id="231DD0FC220EBDD5353A8AEFDF95EFA1" italics="true" pageId="0" pageNumber="7">Surface sculpture</emphasis>
|
|
.
|
|
</emphasis>
|
|
Head with few larger and smaller setiferous punctures; with distinct microreticulation except on middle of head between the eyes (Figs
|
|
<figureCitation id="0A3B573607DBFA61DF5168016D05FE65" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
|
|
,
|
|
<figureCitation id="6BE7E7C5F016AC672699B64ACACB3B51" captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Liodessus spp. nov. Detail of head in dorsal view. L. meridensis (A, B); L. venezuelanus (C, D)." figureDoi="10.3897/alpento.6.79598.figure2" httpUri="https://binary.pensoft.net/fig/682025" pageId="0" pageNumber="7">2A, B</figureCitation>
|
|
). Pronotum and elytron shiny; with moderately dense and coarse setiferous punctation (Fig.
|
|
<figureCitation id="7619A08F3FE5F2749D53E1E1B760B0B3" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="BF627F345782131DA7DCD80A02AE53FD" pageId="0" pageNumber="7">
|
|
<emphasis id="B46D0E3F432DDDE77EC9201ACD17AE0B" bold="true" pageId="0" pageNumber="7">
|
|
<emphasis id="BEC47A4A8289B4D66549A2442A8745F2" italics="true" pageId="0" pageNumber="7">Structures</emphasis>
|
|
.
|
|
</emphasis>
|
|
Antenna stout. Head without occipital line; with rounded clypeus. Pronotum with distinct lateral beads; with distinct long and deep basal striae (Fig.
|
|
<figureCitation id="973A691C9E6696E9F3EDE370182A28A6" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Dorsal habitus of Liodessus spp. nov. L. meridensis (A); L. venezuelanus (B). Scale bar: 2 mm." figureDoi="10.3897/alpento.6.79598.figure1" httpUri="https://binary.pensoft.net/fig/682024" pageId="0" pageNumber="7">1A</figureCitation>
|
|
). Elytron with short basal stria; without sutural line; without basal epipleural transverse carina. Metathoracic wings not examined in holotype (in paratype: short, about half the length of elytron).
|
|
</paragraph>
|
|
<paragraph id="4BCB78F07C4289FB81328D651EC31227" pageId="0" pageNumber="7">
|
|
<emphasis id="FDD222413E1850D9C25FF2D98EBDEF1B" bold="true" pageId="0" pageNumber="7">
|
|
<emphasis id="42E9D2F4043A707DB8A1DF38FE6BA7E2" italics="true" pageId="0" pageNumber="7">Genitalia</emphasis>
|
|
.
|
|
</emphasis>
|
|
Median lobe of aedeagus of long, simple curvature in lateral view, more or less gradually narrowing towards tip; also gradually narrowing towards a narrow tip in ventral view; parameres of simple "
|
|
<taxonomicName id="5FDE7DAA163E25BEDF656A40962468A2" authorityName="Sharp" authorityYear="1880" lsidName="" pageId="0" pageNumber="7" rank="tribe" tribe="Bidessini">Bidessini</taxonomicName>
|
|
" type, two-segmented (Fig.
|
|
<figureCitation id="7C0ADAD429901B5392F783950CEB5F33" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Liodessus spp. nov. Median lobe ventral and lateral views; parameres outer and inner surfaces, L. meridensis (A); L. venezuelanus (B). Scale bar: 0.5 mm." figureDoi="10.3897/alpento.6.79598.figure3" httpUri="https://binary.pensoft.net/fig/682026" pageId="0" pageNumber="7">3A</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="3738A4624590D0BEC0ACF30E20D1D891" doi="10.3897/alpento.6.79598.figure3" httpUri="https://binary.pensoft.net/fig/682026" pageId="0" pageNumber="7" start="Figure 3" startId="F3">
|
|
<paragraph id="240D7537EAD317EB4D6B1C595E21B573" pageId="0" pageNumber="7">
|
|
<emphasis id="2CD5C27F368EEA37A3683BD8CACFCC01" bold="true" pageId="0" pageNumber="7">Figure 3.</emphasis>
|
|
<taxonomicName id="FA10F6CB2B6EAB9574E461CF53486BA1" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="F94BAFC2083C45EC73692FC2D5AF7F30" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
|
|
</taxonomicName>
|
|
spp. nov. Median lobe ventral and lateral views; parameres outer and inner surfaces,
|
|
<taxonomicName id="0496CDBC1A437AB79A59ECC0C899E24D" lsidName="L. meridensis" pageId="0" pageNumber="7" rank="species" species="meridensis">
|
|
<emphasis id="3E9556036311224E038954C714754C07" italics="true" pageId="0" pageNumber="7">L. meridensis</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="95AC2E4C02E5B7EE0F1341C6EDC8B14A" bold="true" pageId="0" pageNumber="7">A</emphasis>
|
|
);
|
|
<taxonomicName id="F6E1A0DD7B84C5646EB0789F855A9283" lsidName="L. venezuelanus" pageId="0" pageNumber="7" rank="species" species="venezuelanus">
|
|
<emphasis id="26F3DC07F6150F46011876AFB2269645" italics="true" pageId="0" pageNumber="7">L. venezuelanus</emphasis>
|
|
</taxonomicName>
|
|
(
|
|
<emphasis id="3DAF8332CE3737ABE777503FE4294475" bold="true" pageId="0" pageNumber="7">B</emphasis>
|
|
). Scale bar: 0.5 mm.
|
|
</paragraph>
|
|
</caption>
|
|
<paragraph id="CC8CA5B426BC5A3296A702C26F6B4206" pageId="0" pageNumber="7">
|
|
<emphasis id="363DD92250C053C83A2245B140C33212" bold="true" pageId="0" pageNumber="7">Female.</emphasis>
|
|
Pronotum and elytra microreticulate, dull (specimens from Gavidia); shiny in the ones from Laguna de
|
|
<normalizedToken id="B9BDD0A0956362F705F04568BBDEB378" originalValue="Mucabají">Mucabaji</normalizedToken>
|
|
.
|
|
</paragraph>
|
|
<paragraph id="21CE0618C92AB6E1AF0819D3EF60E01D" pageId="0" pageNumber="7">
|
|
<emphasis id="798D34C8640AE27B4CA313844B368C01" bold="true" pageId="0" pageNumber="7">
|
|
<emphasis id="88C079729A8A640F5BC747F15D8F378E" italics="true" pageId="0" pageNumber="7">Variation</emphasis>
|
|
.
|
|
</emphasis>
|
|
Size variation of the paratypes is TL 2.1-2.3 mm; TL-h 1.8-2.0 mm; MW 0.9-1.1 mm. The elytral striae are hardly noticeable in a few paratypes.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="EDD396C957EC1B2C00EAEACFDE672552" pageId="0" pageNumber="7" type="etymology">
|
|
<paragraph id="2BCEC36EAFDBDC2C15546ADDA3745C14" pageId="0" pageNumber="7">Etymology.</paragraph>
|
|
<paragraph id="830AB3FF2798BDCDDD0193C6810C14F2" pageId="0" pageNumber="7">
|
|
Named after the state of
|
|
<normalizedToken id="C7B44656EC24EE6A281DF2AE839108BF" originalValue="Mérida">Merida</normalizedToken>
|
|
, in which the collecting localities are situated. The name is an adjective in the nominative singular.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="FE57076322CD33BCE012BC3930451438" pageId="0" pageNumber="7" type="comparisons">
|
|
<paragraph id="4F19774A941396396FDD4F3A11120F63" pageId="0" pageNumber="7">Comparative notes.</paragraph>
|
|
<paragraph id="2805BB0E486FBD7188FE100F0A2C6CDA" pageId="0" pageNumber="7">
|
|
Distinguished from the other species of
|
|
<taxonomicName id="282DF1C530122467879AF00AF705811C" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="438FDF6B045AE542D50750555BFEDC59" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
|
|
</taxonomicName>
|
|
by the diagnostic combination of the following features: Species from Venezuela; beetle smaller (TL 2.1-2.3 mm); very dark brown to blackish; head without occipital line; pronotal and elytral striae present; wings reduced considerably; shape of median lobe (narrow and
|
|
<normalizedToken id="F5F46FA9F405DB3815EF08B7C7D7F703" originalValue="“pointed”">"pointed"</normalizedToken>
|
|
in lateral and ventral view, with its particular shape).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="DE50293CDF5CCD82823A4C82818857B7" pageId="0" pageNumber="7" type="bold platform">
|
|
<paragraph id="D3B13BB45B07B9015914B8587B603973" pageId="0" pageNumber="7">BOLD platform.</paragraph>
|
|
<paragraph id="E4A000BD009A90F2B199BC48DFE8F6AC" pageId="0" pageNumber="7">
|
|
We provided 1 entry to our BOLD
|
|
<taxonomicName id="E9AA83F9F8341CDEA0CF842E00E30E60" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2A79D984628CDC7FC27890DE8D6A35DD" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
|
|
</taxonomicName>
|
|
project. The species is well delineated from all other
|
|
<taxonomicName id="EE90593A3948B95E8EDFFD33B2278D22" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="F606F2469668922E068DDF614451DD18" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
|
|
</taxonomicName>
|
|
, with an uncorrected p-distance of around 9% or more.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="49435EF458DD65B160724489272E0254" pageId="0" pageNumber="7" type="distribution">
|
|
<paragraph id="C82A2B257A490623DB4F8B5C529985AF" pageId="0" pageNumber="7">Distribution.</paragraph>
|
|
<paragraph id="00C4312C1564DF31F8A432B0CEE01F30" pageId="0" pageNumber="7">
|
|
Known from around
|
|
<normalizedToken id="A50889EA86A838F1F22BFEB06B9DED8D" originalValue="Mérida">Merida</normalizedToken>
|
|
, Venezuela.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="01E23B8A29A949841BB46BD013A30A54" pageId="0" pageNumber="7" type="habitat">
|
|
<paragraph id="B1B3DBCC49B522AD30C2C6A6A4F22B9C" pageId="0" pageNumber="7">Habitat.</paragraph>
|
|
<paragraph id="7AC4C8D9CBDC6051293264EAD4B87D40" pageId="0" pageNumber="7">
|
|
Collected from shallow water at the edge of high-altitude ponds (Fig.
|
|
<figureCitation id="97F3C4E3E1AF461DEBDF1A7AEFD77136" captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Habitat of Liodessus spp. nov. near Gavidia, Venenzuela, where both new species were collected (collecting event VZ 10 - 0123 - 03 A). Overview (A); detail of shallow water at the edge (B)." figureDoi="10.3897/alpento.6.79598.figure4" httpUri="https://binary.pensoft.net/fig/682027" pageId="0" pageNumber="7">4</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<caption id="CF3451A9766A210E09C9EEBC4DAA7EDB" doi="10.3897/alpento.6.79598.figure4" httpUri="https://binary.pensoft.net/fig/682027" pageId="0" pageNumber="7" start="Figure 4" startId="F4">
|
|
<paragraph id="A810FE1B0ECAF21BF3EE2F2E49E27217" pageId="0" pageNumber="7">
|
|
<emphasis id="B0F694901EA0C2EE9B19737C93F0B77F" bold="true" pageId="0" pageNumber="7">Figure 4.</emphasis>
|
|
Habitat of
|
|
<taxonomicName id="14E07E9868EEC31EA38914BA3529615B" authorityName="Guignot" authorityYear="1939" class="Insecta" family="Dytiscidae" genus="Liodessus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Liodessus" order="Coleoptera" pageId="0" pageNumber="7" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="FC070A7C971C35BA37CA212205069305" italics="true" pageId="0" pageNumber="7">Liodessus</emphasis>
|
|
</taxonomicName>
|
|
spp. nov. near Gavidia, Venenzuela, where both new species were collected (collecting event VZ10-0123-03A). Overview (
|
|
<emphasis id="A5C15717A3FAE12E2F91824FD9751365" bold="true" pageId="0" pageNumber="7">A</emphasis>
|
|
); detail of shallow water at the edge (
|
|
<emphasis id="EFB5AAD7DE146D8F9A18E7F20E110DB4" bold="true" pageId="0" pageNumber="7">B</emphasis>
|
|
).
|
|
</paragraph>
|
|
</caption>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |