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<document id="D2E7F552EEE37FB8CC4EDA83FD291845" ID-CLB-Dataset="7730" ID-DOI="10.11646/zootaxa.4938.4.6" ID-GBIF-Dataset="2d86f0e3-df0c-46a8-9c7e-4a1b11b109a5" ID-ISSN="1175-5326" ID-Zenodo-Dep="4574962" ID-ZooBank="0A410EF8-285F-43BF-B690-6DFFA7DF2BF7" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1614752980813" checkinUser="plazi" docAuthor="Silva, Jaqueline Santana Tavares Ferreira Da, Grazia, Jocelia &amp; Barão, Kim Ribeiro" docDate="2021" docId="3660577CFFA5FF86E5B29792FF07F885" docLanguage="en" docName="zootaxa.4938.4.6.pdf" docOrigin="Zootaxa 4938 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Steleocoris Mayr 1864" docType="treatment" docVersion="7" lastPageNumber="476" masterDocId="CA592F04FFA4FF87E5259314FFF6FF96" masterDocTitle="Redescription of Steleocoris Mayr, 1864 and Theloris Thunberg, 1783 (Hemiptera Pentatomidae), two poorly known African stink bugs" masterLastPageNumber="486" masterPageNumber="475" pageNumber="476" updateTime="1698931916002" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C4235679080BA5BA90115D89C998860C">Redescription of Steleocoris Mayr, 1864 and Theloris Thunberg, 1783 (Hemiptera Pentatomidae), two poorly known African stink bugs</mods:title>
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<mods:namePart id="59804280A19F33C90ECA084277CC7CB3">Grazia, Jocelia</mods:namePart>
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<mods:affiliation id="3D726821A4881173C03C69737CED641F">Departamento de Zoologia, Instituto de Biociências, Universidade Federal do Rio Grande do Sul, Porto Alegre, RS, Brazil. jocelia @ ufrgs. br; https: // orcid. org / 0000 - 0002 - 7027 - 5638</mods:affiliation>
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<mods:namePart id="1149ECFA755E3BEFB4A377D34862A09C">Barão, Kim Ribeiro</mods:namePart>
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<mods:affiliation id="B8FDD06AAD2C8FBA72929FADA9F5345B">Unidade Educacional Penedo, Campus Arapiraca, Universidade Federal de Alagoas, Penedo, AL, Brazil. kim. barao @ penedo. ufal. br; https: // orcid. org / 0000 - 0003 - 1432 - 1756</mods:affiliation>
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<heading id="E53E5106FFA5FF86E5B29792FE46FB36" bold="true" box="[151,432,1158,1185]" fontSize="11" level="1" pageId="1" pageNumber="476" reason="1">
<taxonomicName id="79C99DE9FFA5FF86E5B29792FE46FB36" authority="Mayr, 1864" authorityName="Mayr" authorityYear="1864" box="[151,432,1158,1185]" class="Insecta" family="Pentatomidae" genus="Steleocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E5B29792FE46FB36" bold="true" box="[151,432,1158,1185]" pageId="1" pageNumber="476">
<emphasis id="8CBD3A78FFA5FF86E5B29792FEEDFB37" bold="true" box="[151,283,1158,1185]" italics="true" pageId="1" pageNumber="476">Steleocoris</emphasis>
<bibRefCitation id="DA589B9BFFA5FF86E4069793FE46FB36" author="Mayr, G. L." box="[291,432,1158,1185]" pageId="1" pageNumber="476" pagination="903 - 914" refId="ref7199" refString="Mayr, G. L. (1864) Diagnosen neuer Hemipteren. Verhandlungen des Zoologisch-Botanischen Gesellschaft in Wien, 14, 903 - 914." type="journal article" year="1864">Mayr, 1864</bibRefCitation>
</emphasis>
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<paragraph id="BE76E66AFFA5FF86E5B297DFFB35FB75" blockId="1.[151,1219,1227,1282]" box="[151,1219,1227,1252]" pageId="1" pageNumber="476">
<treatmentCitationGroup id="9ED9C144FFA5FF86E5B297DFFB35FB75" box="[151,1219,1227,1252]" pageId="1" pageNumber="476">
<treatmentCitation id="3F68C07BFFA5FF86E5B297DFFE1AFB75" author="Mayr, G. L." box="[151,492,1227,1252]" page="910" pageId="1" pageNumber="476" year="1864">
<taxonomicName id="79C99DE9FFA5FF86E5B297DFFE1AFB75" authority="Mayr, 1864: 909 - 910" authorityName="Mayr" authorityPageNumber="909 - 910" authorityYear="1864" box="[151,492,1227,1252]" class="Insecta" family="Pentatomidae" genus="Steleocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E5B297DFFEFEFB75" box="[151,264,1227,1251]" italics="true" pageId="1" pageNumber="476">Steleocoris</emphasis>
<bibRefCitation id="DA589B9BFFA5FF86E42B97D8FE1AFB75" author="Mayr, G. L." box="[270,492,1227,1252]" pageId="1" pageNumber="476" pagination="903 - 914" refId="ref7199" refString="Mayr, G. L. (1864) Diagnosen neuer Hemipteren. Verhandlungen des Zoologisch-Botanischen Gesellschaft in Wien, 14, 903 - 914." type="journal article" year="1864">Mayr, 1864: 909–910</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="3F68C07BFFA5FF86E4D297DFFD70FB75" author="Stal, C." box="[503,646,1227,1251]" page="58" pageId="1" pageNumber="476" year="1876">
<bibRefCitation id="DA589B9BFFA5FF86E4D297DFFD70FB75" author="Stal, C." box="[503,646,1227,1251]" pageId="1" pageNumber="476" pagination="1 - 162" refId="ref7819" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en Forteckning ofver alla hittills kanda Hemiptera, Jemte Systematiska Meddelanden. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål, 1876: 58</bibRefCitation>
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, 79;
<treatmentCitation id="3F68C07BFFA5FF86E79D97DFFC0FFB75" author="Lethierry, L. &amp; Severin, G." box="[696,1017,1227,1251]" page="134" pageId="1" pageNumber="476" year="1893">
<bibRefCitation id="DA589B9BFFA5FF86E79D97DFFC0FFB75" author="Lethierry, L. &amp; Severin, G." box="[696,1017,1227,1251]" pageId="1" pageNumber="476" refId="ref7417" refString="Lethierry, L. &amp; Severin, G. (1893) Catalogue general des Hemipteres. Vol. 1. Pentatomidae, Bruxelles, x + 286 pp." type="book" year="1893">Lethierry &amp; Severin, 1893: 134</bibRefCitation>
</treatmentCitation>
;
<bibRefCitation id="DA589B9BFFA5FF86E12197DFFB49FB75" author="Kirkaldy, G. W." box="[1028,1215,1227,1251]" pageId="1" pageNumber="476" refId="ref7225" refString="Kirkaldy, G. W. (1909) Catalogue of the Hemiptera (Heteroptera) with biological and anatomical references, lists of foodplants and parasites, etc. Vol. I. Cimicidae. F. L. Dames, Berlin, xl + 392 pp." type="book" year="1909">Kirkaldy, 1909:74</bibRefCitation>
.
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</paragraph>
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<taxonomicName id="79C99DE9FFA5FF86E5B297FEFE8EFA94" ID-CoL="XSY" authority="Stal, 1865: 78" authorityName="Stal" authorityPageNumber="78" authorityYear="1865" box="[151,376,1257,1282]" class="Insecta" family="Bombyliidae" genus="Antonia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E5B297FEFF11FA97" box="[151,231,1258,1281]" italics="true" pageId="1" pageNumber="476">Antonia</emphasis>
<bibRefCitation id="DA589B9BFFA5FF86E5CB97FDFE8EFA94" author="Stal, C." box="[238,376,1257,1282]" pageId="1" pageNumber="476" refId="ref7796" refString="Stal, C. (1865) Hemiptera Africana. Vol. 1. Norstedtiana, Stockholm, iv + 256 pp." type="book" year="1865">Stål, 1865:78</bibRefCitation>
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, 125–126.
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<paragraph id="BE76E66AFFA5FF86E5B29633FD0AFAD6" blockId="1.[151,764,1319,1344]" box="[151,764,1319,1344]" pageId="1" pageNumber="476">
<emphasis id="8CBD3A78FFA5FF86E5B29633FED0FAA9" bold="true" box="[151,294,1319,1343]" pageId="1" pageNumber="476">
<typeStatus id="617258C8FFA5FF86E5B29633FF3AFAA9" box="[151,204,1319,1343]" pageId="1" pageNumber="476">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="79C99DE9FFA5FF86E409963CFD90FAA9" authority="Thunberg, 1783" authorityName="Thunberg" authorityYear="1783" box="[300,614,1319,1343]" class="Insecta" family="Cimicidae" genus="Cimex" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="species" species="comma">
<emphasis id="8CBD3A78FFA5FF86E409963CFE4AFAA9" box="[300,444,1320,1343]" italics="true" pageId="1" pageNumber="476">Cimex comma</emphasis>
<bibRefCitation id="DA589B9BFFA5FF86E4E79633FD90FAA9" author="Thunberg, C. P." box="[450,614,1319,1343]" pageId="1" pageNumber="476" pagination="29 - 52" refId="ref7857" refString="Thunberg, C. P. (1783) Dissertatio entomologica novas insectorum species, sistens, cujus partem secundam, cons. exper. facult, med. upsal., publice ventilandam exhibent Johan. Edman, Upsala, pp. 29 - 52, pl. 2." type="book chapter" year="1783">Thunberg, 1783</bibRefCitation>
</taxonomicName>
, by monotypy.
</paragraph>
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<subSubSection id="F6D3B5E1FFA5FF86E5B29679FD40F93E" pageId="1" pageNumber="476" type="diagnosis">
<paragraph id="BE76E66AFFA5FF86E5B29679FD40F93E" blockId="1.[151,1437,1389,1811]" pageId="1" pageNumber="476">
<emphasis id="8CBD3A78FFA5FF86E5B29679FEE4FA11" bold="true" box="[151,274,1389,1415]" pageId="1" pageNumber="476">Diagnosis.</emphasis>
Head longer than wide, mandibular plates longer than clypeus. Antenniferous tubercles with lateral and ventral tubercles. Antennomere II longer than other antennomeres. Anterior margin of bucculae lobate, posterior margin lobate tapering towards base of head. Labiomere I contained between bucculae; labium reaching posterior margin of mesocoxae. Scutellum longer than wide; frenal lobe as long as post-frenal lobe. Pro-, meso- and metasternum sulcate. Mesosternum with slightly elevated carina. Peritreme of metasternal gland opening spout. Meso- and metacoxae closer to each other than to procoxae. Trichobothria posterior to spiracles, one mesial and other laterad to an imaginary line connecting spiracles. Genital cup of pygophore striated. Paramere crown falcate, with scale-like sculptures laterally. Segment X up to two times longer than wide. Gonocoxites VIII as long as wide, swollen, facing posteriorly; posterior and mesial margins setose.
</paragraph>
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<subSubSection id="F6D3B5E1FFA5FF86E5E295A5FF07F885" pageId="1" pageNumber="476" type="discussion">
<paragraph id="BE76E66AFFA5FF86E5E295A5FF07F885" blockId="1.[151,1437,1389,1811]" pageId="1" pageNumber="476">
<emphasis id="8CBD3A78FFA5FF86E5E295A5FECDF95D" bold="true" box="[199,315,1713,1739]" pageId="1" pageNumber="476">Remarks.</emphasis>
<bibRefCitation id="DA589B9BFFA5FF86E46195A5FE32F95A" author="Stal, C." box="[324,452,1713,1740]" pageId="1" pageNumber="476" pagination="1 - 162" refId="ref7819" refString="Stal, C. (1876) Enumeratio Hemipterorum. Bidrag till en Forteckning ofver alla hittills kanda Hemiptera, Jemte Systematiska Meddelanden. Kongliga Svenska Vetenskaps-Akademiens Handlingar, 14 (4), 1 - 162." type="journal article" year="1876">Stål (1876)</bibRefCitation>
included
<taxonomicName id="79C99DE9FFA5FF86E71295A5FD45F95D" authorityName="Mayr" authorityYear="1864" box="[567,691,1713,1739]" class="Insecta" genus="Steleocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E71295A5FD45F95D" box="[567,691,1713,1739]" italics="true" pageId="1" pageNumber="476">Steleocoris</emphasis>
</taxonomicName>
in the Diploxyini, along with
<taxonomicName id="79C99DE9FFA5FF86E13695A5FB7EF95A" authority="Stal" authorityName="Stal" authorityYear="1876" box="[1043,1160,1713,1740]" class="Insecta" family="Pentatomidae" genus="Adria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E13695A5FBA5F95D" box="[1043,1107,1713,1739]" italics="true" pageId="1" pageNumber="476">Adria</emphasis>
Stål
</taxonomicName>
and
<taxonomicName id="79C99DE9FFA5FF86E1E695A5FECCF966" authority="Amyot &amp; Serville. More" authorityName="Amyot &amp; Serville. More" authorityYear="1843" class="Insecta" family="Pentatomidae" genus="Diploxys" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E1E695A5FAD1F95D" box="[1219,1319,1713,1739]" italics="true" pageId="1" pageNumber="476">Diploxys</emphasis>
Amyot &amp; Serville. More
</taxonomicName>
recently, the
<taxonomicName id="79C99DE9FFA5FF86E4F495C1FDBBF979" authorityName="Mayr" authorityYear="1864" box="[465,589,1749,1775]" class="Insecta" genus="Steleocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="genus">
<emphasis id="8CBD3A78FFA5FF86E4F495C1FDBBF979" box="[465,589,1749,1775]" italics="true" pageId="1" pageNumber="476">Steleocoris</emphasis>
</taxonomicName>
has been placed in
<taxonomicName id="79C99DE9FFA5FF86E60E95C1FC47F979" authorityName="Mulsant &amp; Rey" authorityYear="1866" box="[811,945,1749,1775]" class="Insecta" family="Pentatomidae" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="476" phylum="Arthropoda" rank="tribe" tribe="Carpocorini">Carpocorini</taxonomicName>
(
<bibRefCitation id="DA589B9BFFA5FF86E69A95C1FB77F979" author="Rider, D. A. &amp; Schwertner, C. F. &amp; Vilimova, J. &amp; Redei, D. &amp; Kment, P. &amp; Thomas, D. B." box="[959,1153,1749,1776]" pageId="1" pageNumber="476" pagination="25 - 201" refId="ref7513" refString="Rider, D. A., Schwertner, C. F., Vilimova, J., Redei, D., Kment, P. &amp; Thomas, D. B. (2018) Higher systematics of the Pentatomoidea. In: McPherson, J. E. (Ed.), Invasive stink bugs and related species (Pentatomoidea): biology, higher systematics, semiochemistry, and management. CRC Press, Boca Raton, pp. 25 - 201. https: // doi. org / 10.1201 / 9781315371221 - 2" type="book chapter" year="2018">
Rider
<emphasis id="8CBD3A78FFA5FF86E12295C2FBCEF979" box="[1031,1080,1749,1775]" italics="true" pageId="1" pageNumber="476">et al</emphasis>
. 2018
</bibRefCitation>
;
<bibRefCitation id="DA589B9BFFA5FF86E1A895C1FAFBF979" author="Rider, D. A." box="[1165,1293,1749,1775]" pageId="1" pageNumber="476" refId="ref7614" refString="Rider, D. A. (2020) Pentatomoidea Home page, The Department of Entomology. Available from: https: // www. ndsu. edu / pubweb / ~ rider / Pentatomoidea / Genus _ Index / genus _ index. htm (accessed 31 July 2020)" type="url" year="2020">Rider 2020</bibRefCitation>
) though tentatively.
</paragraph>
</subSubSection>
</treatment>
</document>
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