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<document id="D700BB26D6A8E003935D89D38C10133D" ID-CLB-Dataset="8641" ID-DOI="10.11646/zootaxa.4912.1.1" ID-GBIF-Dataset="8a5efe86-cabc-4981-afb4-163791f2530c" ID-ISSN="1175-5326" ID-Zenodo-Dep="4450930" ID-ZooBank="9536C1CF-4AEF-47F8-959B-48CD7A5392D8" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1611086167060" checkinUser="plazi" docAuthor="Van, Rob W. M., Aryasari, Ratih &amp; De, Nicole J." docDate="2021" docId="361087A7FF5CFF3B55ABFC7755F9CAE4" docLanguage="en" docName="zootaxa.4912.1.1.pdf" docOrigin="Zootaxa 4912 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Mycale (Naviculina) mascarenensis Van &amp; Aryasari &amp; De 2021, sp.nov." docType="treatment" docVersion="12" lastPageNumber="156" masterDocId="CA29FFDFFFC5FFA0553CFFA5506ACD15" masterDocTitle="Mycale species of the tropical Indo-West Pacific (Porifera, Demospongiae, Poecilosclerida)" masterLastPageNumber="212" masterPageNumber="1" pageNumber="154" updateTime="1698924900638" updateUser="ExternalLinkService">
<mods:mods id="1F771372285EC45FC22476F28110BA0C" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="052ABF4F1D17378D7E1D0C317BEE75EC">
<mods:title id="27454ECDA7C5C243B47FE876FBD6E5A3">Mycale species of the tropical Indo-West Pacific (Porifera, Demospongiae, Poecilosclerida)</mods:title>
</mods:titleInfo>
<mods:name id="E7E71CAAB19B9428C9D92C6FE2173C32" type="personal">
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<mods:namePart id="02C2D9C57701DC97C35C77E5235A34B2">Van, Rob W. M.</mods:namePart>
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<mods:role id="B799D272EC2B972822CECA4B1BFEDE35">
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<mods:namePart id="023B251B6B369A7379C6A7384E311332">Aryasari, Ratih</mods:namePart>
</mods:name>
<mods:name id="109BEE52CEEF726CC75F83E0D4C9103C" type="personal">
<mods:role id="2C7F1C0307591B0FCB845D78CE7ABA9A">
<mods:roleTerm id="24178BCAE8ECC9FE4F83F34906D38F4E">Author</mods:roleTerm>
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<mods:namePart id="17CCB68169FECB7232BF82494AA515DF">De, Nicole J.</mods:namePart>
<mods:nameIdentifier id="1532663655F5AC309A3CEDBA90F8EA66" type="ORCID">0000-0002-7985-5604</mods:nameIdentifier>
<mods:nameIdentifier id="AE78D058D7EE4604AC597FE604C96822" type="email">rob.vansoest@naturalis.nl</mods:nameIdentifier>
</mods:name>
<mods:typeOfResource id="EA69407F00EA69E3AB53B163FB550EDA">text</mods:typeOfResource>
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<mods:title id="9F61E824EFF9632483463F340B1526EE">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="97E570A6A7F40E2AC43759A0EF58BEF0">
<mods:date id="91EB4B9F4D9657F9D5AAE6661848D138">2021</mods:date>
<mods:detail id="DFE0687D0B8D6EC89D90B39B2B86C7E3" type="pubDate">
<mods:number id="6E89CD222F27F31CB01CA7318BA6F982">2021-01-19</mods:number>
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<mods:detail id="20F37C2C44528B92169F9E055EEE219A" type="volume">
<mods:number id="B794D35DF249C51EF76BE02D30760EF5">4912</mods:number>
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<mods:classification id="A122E156B763A1768504FAC80AAE621E">journal article</mods:classification>
<mods:identifier id="6CB8ABDD5536EED837B56FDBCC68DB26" type="CLB-Dataset">8641</mods:identifier>
<mods:identifier id="B345069F6E36D159962BB4C70D152813" type="DOI">10.11646/zootaxa.4912.1.1</mods:identifier>
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<treatment id="361087A7FF5CFF3B55ABFC7755F9CAE4" ID-DOI="http://doi.org/10.5281/zenodo.4451076" ID-GBIF-Taxon="177132463" ID-Zenodo-Dep="4451076" LSID="urn:lsid:plazi:treatment:361087A7FF5CFF3B55ABFC7755F9CAE4" httpUri="http://treatment.plazi.org/id/361087A7FF5CFF3B55ABFC7755F9CAE4" lastPageId="155" lastPageNumber="156" pageId="153" pageNumber="154">
<subSubSection id="F6A3653AFF5CFF3955ABFC7752CCCEF8" box="[151,678,978,1005]" pageId="153" pageNumber="154" type="nomenclature">
<paragraph id="BE0636B1FF5CFF3955ABFC7752CCCEF8" blockId="153.[151,678,978,1041]" box="[151,678,978,1005]" pageId="153" pageNumber="154">
<heading id="E54E81DDFF5CFF3955ABFC7752CCCEF8" bold="true" box="[151,678,978,1005]" fontSize="11" level="2" pageId="153" pageNumber="154" reason="4">
<emphasis id="8CCDEAA3FF5CFF3955ABFC7752CCCEF8" bold="true" box="[151,678,978,1005]" pageId="153" pageNumber="154">
<taxonomicName id="79B94D32FF5CFF3955ABFC775223CEF8" ID-CoL="86X5C" authority="Van &amp; Aryasari &amp; De, 2021" authorityName="Van &amp; Aryasari &amp; De" authorityYear="2021" box="[151,585,978,1005]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="153" pageNumber="154" phylum="Porifera" rank="species" species="mascarenensis" status="sp. nov." subGenus="Naviculina">
<emphasis id="8CCDEAA3FF5CFF3955ABFC775223CEF8" bold="true" box="[151,585,978,1005]" italics="true" pageId="153" pageNumber="154">Mycale (Naviculina) mascarenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97FE57D8FF5CFF39576CFC7652CCCEF8" box="[592,678,979,1005]" pageId="153" pageNumber="154" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="F6A3653AFF5CFF3955ABFC5251BAC904" box="[151,464,1015,1041]" pageId="153" pageNumber="154" type="description">
<paragraph id="BE0636B1FF5CFF3955ABFC5251BAC904" blockId="153.[151,678,978,1041]" box="[151,464,1015,1041]" pageId="153" pageNumber="154">
<figureCitation id="26822A34FF5CFF3955ABFC525081C904" box="[151,235,1015,1041]" captionStart="FIGURE 99" captionStartId="154.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-26@154.[151,1435,181,1914]" captionTargetPageId="154" captionText="FIGURE 99. Mycale (Naviculina) mascarenensis sp.nov., habitus in situ, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd)." figureDoi="http://doi.org/10.5281/zenodo.4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="153" pageNumber="154">Figs 99</figureCitation>
a–c, 100a–d, 101a–e
</paragraph>
</subSubSection>
<subSubSection id="F6A3653AFF5CFF3955ABFB9B532BC9D3" pageId="153" pageNumber="154" type="materials_examined">
<paragraph id="BE0636B1FF5CFF3955ABFB9B54AFC96B" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
<emphasis id="8CCDEAA3FF5CFF3955ABFB9B511DC94D" bold="true" box="[151,375,1086,1112]" pageId="153" pageNumber="154">Material examined</emphasis>
.
<materialsCitation id="0ED13CECFF5CFF3954BFFB9A54AFC96B" ID-GBIF-Occurrence="3026466714" collectingDate="2016-10-19" collectionCode="RMNH, SCUBA" collectorName="N. J. de Voogd" elevation="18" latitude="-19.7648" location="Castel Rock" longLatPrecision="7" longitude="63.4626" municipality="Mourouk Ebony" pageId="153" pageNumber="154" specimenCount="1" stateProvince="Rodrigues" typeStatus="holotype">
<typeStatus id="61028813FF5CFF3954BFFB9A5181C94C" box="[387,491,1087,1113]" pageId="153" pageNumber="154" type="holotype">Holotype</typeStatus>
<collectionCode id="D8A8AE74FF5CFF3954CCFB9A522EC94C" box="[496,580,1087,1113]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="153" pageNumber="154" type="Museum">RMNH</collectionCode>
Por. 11689,
<collectingRegion id="7C7DF853FF5CFF3957F6FB9A532AC94C" box="[714,832,1087,1113]" country="Mauritius" name="Rodrigues" pageId="153" pageNumber="154">Rodrigues</collectingRegion>
,
<collectingMunicipality id="5E62ACCBFF5CFF395675FB9A546AC94C" box="[841,1024,1087,1113]" pageId="153" pageNumber="154">Mourouk Ebony</collectingMunicipality>
,
<location id="BB66606AFF5CFF395135FB9A54F8C94C" LSID="urn:lsid:plazi:treatment:361087A7FF5CFF3B55ABFC7755F9CAE4:BB66606AFF5CFF395135FB9A54F8C94C" box="[1033,1170,1087,1113]" latitude="-19.7648" longLatPrecision="7" longitude="63.4626" municipality="Mourouk Ebony" name="Castel Rock" pageId="153" pageNumber="154" stateProvince="Rodrigues">Castel Rock</location>
,
<geoCoordinate id="DB8D5076FF5CFF3951A6FB9A557AC94C" box="[1178,1296,1087,1113]" degrees="19.7648" direction="south" orientation="latitude" pageId="153" pageNumber="154" precision="5" value="-19.7648">19.7648°S</geoCoordinate>
S
<geoCoordinate id="DB8D5076FF5CFF395015FB9A55F2C94C" box="[1321,1432,1087,1113]" degrees="63.4626" direction="east" orientation="longitude" pageId="153" pageNumber="154" precision="5" value="63.4626">63.4626E</geoCoordinate>
,
<elevation id="3594D182FF5CFF3955ABFBC65172C968" box="[151,280,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="153" pageNumber="154" unit="m" value="18.0">
<quantity id="79419B54FF5CFF3955ABFBC650BAC968" box="[151,208,1123,1149]" metricMagnitude="1" metricUnit="m" metricValue="1.8" pageId="153" pageNumber="154" unit="m" value="18.0">18 m</quantity>
depth
</elevation>
,
<collectionCode id="D8A8AE74FF5CFF39541FFBC651E9C968" box="[291,387,1123,1149]" pageId="153" pageNumber="154">SCUBA</collectionCode>
, coll.
<collectorName id="134C5367FF5CFF3954FBFBC65202C968" box="[455,616,1123,1149]" pageId="153" pageNumber="154">N.J. de Voogd</collectorName>
, field nr. ROG 103,
<date id="CA071071FF5CFF395676FBC6546DC968" box="[842,1031,1123,1149]" pageId="153" pageNumber="154" value="2016-10-19">
<collectingDate id="DA43E999FF5CFF395676FBC6546DC968" box="[842,1031,1123,1149]" pageId="153" pageNumber="154" value="2016-10-19">19 October 2016</collectingDate>
</date>
(orange-brown).
</materialsCitation>
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBFB22532BC9D3" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
<materialsCitation id="0ED13CECFF5CFF3955FBFB2254F6C9B4" ID-GBIF-Occurrence="3026466830" box="[199,1180,1158,1185]" collectionCode="RMNH" country="Grenada" elevation="8" latitude="-19.675" location="Rodrigues" longLatPrecision="55" longitude="63.4842" municipality="Passe" pageId="153" pageNumber="154" specimenCount="1" typeStatus="paratype">
<typeStatus id="61028813FF5CFF3955FBFB225143C9B4" box="[199,297,1159,1185]" pageId="153" pageNumber="154" type="paratype">Paratype</typeStatus>
<collectionCode id="D8A8AE74FF5CFF39540DFB2251EFC9B4" box="[305,389,1159,1185]" country="Netherlands" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34992" lsid="urn:lsid:biocol.org:col:34992" name="National Museum of Natural History, Naturalis" pageId="153" pageNumber="154" type="Museum">RMNH</collectionCode>
Por. 11723,
<location id="BB66606AFF5CFF395728FB2252E0C9B4" LSID="urn:lsid:plazi:treatment:361087A7FF5CFF3B55ABFC7755F9CAE4:BB66606AFF5CFF395728FB2252E0C9B4" box="[532,650,1159,1185]" country="Grenada" latitude="-19.675" longLatPrecision="55" longitude="63.4842" municipality="Passe" name="Rodrigues" pageId="153" pageNumber="154">Rodrigues</location>
,
<collectingMunicipality id="5E62ACCBFF5CFF3957AAFB2252BEC9B4" box="[662,724,1159,1185]" pageId="153" pageNumber="154">Passe</collectingMunicipality>
<collectingCountry id="C6AE7621FF5CFF3957E0FB225354C9B4" box="[732,830,1159,1185]" name="Grenada" pageId="153" pageNumber="154">Grenade</collectingCountry>
,
<geoCoordinate id="DB8D5076FF5CFF395675FB2353CCC9B4" box="[841,934,1158,1185]" degrees="19.675" direction="south" orientation="latitude" pageId="153" pageNumber="154" precision="55" value="-19.675">19.675S</geoCoordinate>
<geoCoordinate id="DB8D5076FF5CFF395692FB225476C9B4" box="[942,1052,1159,1185]" degrees="63.4842" direction="east" orientation="longitude" pageId="153" pageNumber="154" precision="5" value="63.4842">63.4842E</geoCoordinate>
,
<elevation id="3594D182FF5CFF395114FB2254F6C9B4" box="[1064,1180,1159,1185]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="153" pageNumber="154" unit="m" value="8.0">
<quantity id="79419B54FF5CFF395114FB22543EC9B4" box="[1064,1108,1159,1185]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="153" pageNumber="154" unit="m" value="8.0">8 m</quantity>
depth
</elevation>
</materialsCitation>
,
<materialsCitation id="0ED13CECFF5CFF395194FB225357C9D3" ID-GBIF-Occurrence="3026466890" collectingDate="2016-10-21" collectionCode="SCUBA" collectorName="N. J. de Voogd" country="Grenada" pageId="153" pageNumber="154" specimenCount="1" typeStatus="paratype">
<collectionCode id="D8A8AE74FF5CFF395194FB225562C9B4" box="[1192,1288,1159,1185]" pageId="153" pageNumber="154">SCUBA</collectionCode>
, coll.
<collectorName id="134C5367FF5CFF395071FB225089C9D0" pageId="153" pageNumber="154">N.J. de Voogd</collectorName>
, field nr. ROG 140,
<date id="CA071071FF5CFF3954FAFB0E52E8C9D0" box="[454,642,1195,1221]" pageId="153" pageNumber="154" value="2016-10-21">
<collectingDate id="DA43E999FF5CFF3954FAFB0E52E8C9D0" box="[454,642,1195,1221]" pageId="153" pageNumber="154" value="2016-10-21">21 October 2016</collectingDate>
</date>
(orange-brown)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="F6A3653AFF5CFF3B55FBFB6B512FCA0C" lastPageId="155" lastPageNumber="156" pageId="153" pageNumber="154" type="description">
<paragraph id="BE0636B1FF5CFF3955FBFB6B534BC86C" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
<emphasis id="8CCDEAA3FF5CFF3955FBFB6B513BC9FD" bold="true" box="[199,337,1230,1256]" pageId="153" pageNumber="154">Description</emphasis>
(
<figureCitation id="26822A34FF5CFF39545DFB6A51D2C9FC" box="[353,440,1231,1257]" captionStart="FIGURE 99" captionStartId="154.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-26@154.[151,1435,181,1914]" captionTargetPageId="154" captionText="FIGURE 99. Mycale (Naviculina) mascarenensis sp.nov., habitus in situ, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd)." figureDoi="http://doi.org/10.5281/zenodo.4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="153" pageNumber="154">Figs 99</figureCitation>
a–c, 100a–b). Massively encrusting on dead corals and rocks (
<figureCitation id="26822A34FF5CFF395144FB6A54A5C9FC" box="[1144,1231,1231,1257]" captionStart="FIGURE 99" captionStartId="154.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-26@154.[151,1435,181,1914]" captionTargetPageId="154" captionText="FIGURE 99. Mycale (Naviculina) mascarenensis sp.nov., habitus in situ, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd)." figureDoi="http://doi.org/10.5281/zenodo.4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="153" pageNumber="154">Figs 99</figureCitation>
a–c), also consolidating coral rubble (
<figureCitation id="26822A34FF5CFF395442FB56518EC81B" box="[382,484,1267,1294]" captionStart="FIGURE 100" captionStartId="155.[151,250,1540,1564]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-19@155.[151,1435,181,1517]" captionTargetPageId="155" captionText="FIGURE 100. Mycale (Naviculina) mascarenensis sp.nov., habitus and skeleton of holotype RMNH Por. 11689, from Rodrigues, a, on deck, b, detail of preserved specimen (scale bar = 1 cm), c, light microscopic image of ectosomal skeleton showing intercrossing spicule tracts and rosettes of anisochelae I, d, details of skeleton with flagellated sigma, rosettes of anisochelae and scattered naviculichelae." figureDoi="http://doi.org/10.5281/zenodo.4451175" httpUri="https://zenodo.org/record/4451175/files/figure.png" pageId="153" pageNumber="154">Figs 100</figureCitation>
a–b). Color orange-brown, faintly punctate (
<figureCitation id="26822A34FF5CFF3956E7FB565457C818" box="[987,1085,1267,1293]" captionStart="FIGURE 99" captionStartId="154.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-26@154.[151,1435,181,1914]" captionTargetPageId="154" captionText="FIGURE 99. Mycale (Naviculina) mascarenensis sp.nov., habitus in situ, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd)." figureDoi="http://doi.org/10.5281/zenodo.4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="153" pageNumber="154">Fig. 99c</figureCitation>
), surface not clearly reticulate. On deck, color turns more reddish orange. In preserved condition the specimens are collapsed and fragmented, and beige colored. The detachable skin is transparent and becomes largely loose from the pulpy choanosomal tisse. Size up to 5 x 3 x
<quantity id="79419B54FF5CFF395417FAFA5108C86C" box="[299,354,1375,1401]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="153" pageNumber="154" unit="cm" value="2.0">2 cm</quantity>
(preserved
<typeStatus id="61028813FF5CFF3954DAFAFA5227C86F" box="[486,589,1375,1402]" pageId="153" pageNumber="154" type="holotype">holotype</typeStatus>
). Consistency soft.
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBFA2752E7CB60" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
<emphasis id="8CCDEAA3FF5CFF3955FBFA275144C889" bold="true" box="[199,302,1410,1436]" pageId="153" pageNumber="154">Skeleton</emphasis>
(
<figureCitation id="26822A34FF5CFF395402FA2651C8C88B" box="[318,418,1411,1438]" captionStart="FIGURE 100" captionStartId="155.[151,250,1540,1564]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-19@155.[151,1435,181,1517]" captionTargetPageId="155" captionText="FIGURE 100. Mycale (Naviculina) mascarenensis sp.nov., habitus and skeleton of holotype RMNH Por. 11689, from Rodrigues, a, on deck, b, detail of preserved specimen (scale bar = 1 cm), c, light microscopic image of ectosomal skeleton showing intercrossing spicule tracts and rosettes of anisochelae I, d, details of skeleton with flagellated sigma, rosettes of anisochelae and scattered naviculichelae." figureDoi="http://doi.org/10.5281/zenodo.4451175" httpUri="https://zenodo.org/record/4451175/files/figure.png" pageId="153" pageNumber="154">Figs 100</figureCitation>
c-d). Choanosomal skeleton cavernous, with a basal spongin-rich spicule mass enveloping coral debris, from which issue comparatively thin spicule tracts,
<quantity id="79419B54FF5CFF39564DFA035399C8D4" box="[881,1011,1446,1473]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" metricValueMax="1.5" metricValueMin="0.5" pageId="153" pageNumber="154" unit="mm" value="100.0" valueMax="150.0" valueMin="50.0">50–150 µm</quantity>
in diameter, further thinning out near the surface to carry the ectosomal skeleton. The detachable tangential skeleton is parchment-like and consists of the usual aegogropila-type reticulation of intercrossing spicule tracts (
<figureCitation id="26822A34FF5CFF395643FA4A5384CB1C" box="[895,1006,1519,1545]" captionStart="FIGURE 100" captionStartId="155.[151,250,1540,1564]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-19@155.[151,1435,181,1517]" captionTargetPageId="155" captionText="FIGURE 100. Mycale (Naviculina) mascarenensis sp.nov., habitus and skeleton of holotype RMNH Por. 11689, from Rodrigues, a, on deck, b, detail of preserved specimen (scale bar = 1 cm), c, light microscopic image of ectosomal skeleton showing intercrossing spicule tracts and rosettes of anisochelae I, d, details of skeleton with flagellated sigma, rosettes of anisochelae and scattered naviculichelae." figureDoi="http://doi.org/10.5281/zenodo.4451175" httpUri="https://zenodo.org/record/4451175/files/figure.png" pageId="153" pageNumber="154">Fig. 100c</figureCitation>
). These are comparatively thin,
<quantity id="79419B54FF5CFF39506AFA4B50D5CB38" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="153" pageNumber="154" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15–35 µm</quantity>
, consisting of up to 4 spicules in cross section, meshes up to
<quantity id="79419B54FF5CFF395658F9B653D0CB38" box="[868,954,1555,1581]" metricMagnitude="-1" metricUnit="m" metricValue="3.0" pageId="153" pageNumber="154" unit="mm" value="300.0">300 µm</quantity>
in widest dimension. Rosettes of anisochelae,
<quantity id="79419B54FF5CFF3955F9F992512FCB44" box="[197,325,1591,1617]" metricMagnitude="-2" metricUnit="m" metricValue="8.5" metricValueMax="10.0" metricValueMin="7.0" pageId="153" pageNumber="154" unit="mm" value="85.0" valueMax="100.0" valueMin="70.0">70–100 µm</quantity>
in diameter (
<figureCitation id="26822A34FF5CFF3954EBF9925250CB47" box="[471,570,1591,1618]" captionStart="FIGURE 100" captionStartId="155.[151,250,1540,1564]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-19@155.[151,1435,181,1517]" captionTargetPageId="155" captionText="FIGURE 100. Mycale (Naviculina) mascarenensis sp.nov., habitus and skeleton of holotype RMNH Por. 11689, from Rodrigues, a, on deck, b, detail of preserved specimen (scale bar = 1 cm), c, light microscopic image of ectosomal skeleton showing intercrossing spicule tracts and rosettes of anisochelae I, d, details of skeleton with flagellated sigma, rosettes of anisochelae and scattered naviculichelae." figureDoi="http://doi.org/10.5281/zenodo.4451175" httpUri="https://zenodo.org/record/4451175/files/figure.png" pageId="153" pageNumber="154">Figs 100</figureCitation>
c–d), as well as single anisochelae II and less frequent flagellated sigmas (
<figureCitation id="26822A34FF5CFF395053F99250A8CB63" captionStart="FIGURE 99" captionStartId="154.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-26@154.[151,1435,181,1914]" captionTargetPageId="154" captionText="FIGURE 99. Mycale (Naviculina) mascarenensis sp.nov., habitus in situ, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd)." figureDoi="http://doi.org/10.5281/zenodo.4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="153" pageNumber="154">Fig. 99d</figureCitation>
) are scattered between the sutface tracts.
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBF9DB516CCBA8" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
<emphasis id="8CCDEAA3FF5CFF3955FBF9DB5143CB8D" bold="true" box="[199,297,1662,1688]" pageId="153" pageNumber="154">Spicules</emphasis>
(
<figureCitation id="26822A34FF5CFF395404F9DA51F1CB8C" box="[312,411,1663,1689]" captionStart="FIGURE 101" captionStartId="156.[151,250,1554,1578]" captionTargetBox="[151,1437,181,1530]" captionTargetId="figure-26@156.[151,1437,181,1530]" captionTargetPageId="156" captionText="FIGURE 101. Mycale (Naviculina) mascarenensis sp.nov., SEM images of spicules of holotype RMNH Por. 11723, from Rodrigues, a, mycalostyle, a1, details of mycalostyle, b, anisochela I, c, anisochelae II, d, anisochela III (anisochela II and III same scale), e, flagellated sigmas." figureDoi="http://doi.org/10.5281/zenodo.4451179" httpUri="https://zenodo.org/record/4451179/files/figure.png" pageId="153" pageNumber="154">Figs 101</figureCitation>
a–e). Mycalostyles, three categories of anisochelae, flagellated sigmas (no ‘normal’ sigmas, no toxas).
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBF962519BCA10" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
Mycalostyles (
<figureCitation id="26822A34FF5CFF395457F962519ECBF4" box="[363,500,1735,1761]" captionStart="FIGURE 101" captionStartId="156.[151,250,1554,1578]" captionTargetBox="[151,1437,181,1530]" captionTargetId="figure-26@156.[151,1437,181,1530]" captionTargetPageId="156" captionText="FIGURE 101. Mycale (Naviculina) mascarenensis sp.nov., SEM images of spicules of holotype RMNH Por. 11723, from Rodrigues, a, mycalostyle, a1, details of mycalostyle, b, anisochela I, c, anisochelae II, d, anisochela III (anisochela II and III same scale), e, flagellated sigmas." figureDoi="http://doi.org/10.5281/zenodo.4451179" httpUri="https://zenodo.org/record/4451179/files/figure.png" pageId="153" pageNumber="154">Figs 101a,a</figureCitation>
<figureCitation id="26822A34FF5CFF3954CFF9625197CBF4" box="[499,509,1735,1761]" captionStart="FIGURE 1" captionStartId="6.[151,250,1177,1201]" captionTargetBox="[151,1436,381,1154]" captionTargetId="figure-111@6.[151,1436,381,1154]" captionTargetPageId="6" captionText="FIGURE 1. A. Numbers of available specimens for the present study from the collections of the Naturalis Biodiversity Center distributed over Indo-West Pacific Marine Ecoregions (MEOWS, cf. Spalding et al. 2007). B. Ditto for type specimens and additional specimens from other institutions consulted for this study." figureDoi="http://doi.org/10.5281/zenodo.4450932" httpUri="https://zenodo.org/record/4450932/files/figure.png" pageId="153" pageNumber="154">1</figureCitation>
), straight, slim, with elongated, barely developed heads and pointed opposite ends, 318–
<emphasis id="8CCDEAA3FF5CFF3955F3F94E5164CA10" box="[207,270,1771,1797]" italics="true" pageId="153" pageNumber="154">355.8</emphasis>
–399 x 3–
<emphasis id="8CCDEAA3FF5CFF395442F94E51CBCA10" box="[382,417,1771,1797]" italics="true" pageId="153" pageNumber="154">5.2</emphasis>
–
<quantity id="79419B54FF5CFF39548CF94E5187CA10" box="[432,493,1771,1797]" metricMagnitude="-3" metricUnit="m" metricValue="7.0" pageId="153" pageNumber="154" unit="mm" value="7.0">7 µm</quantity>
.
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBF8AA53E9CA58" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
Anisochelae I (
<figureCitation id="26822A34FF5CFF395449F8AA518DCA3F" box="[373,487,1807,1834]" captionStart="FIGURE 101" captionStartId="156.[151,250,1554,1578]" captionTargetBox="[151,1437,181,1530]" captionTargetId="figure-26@156.[151,1437,181,1530]" captionTargetPageId="156" captionText="FIGURE 101. Mycale (Naviculina) mascarenensis sp.nov., SEM images of spicules of holotype RMNH Por. 11723, from Rodrigues, a, mycalostyle, a1, details of mycalostyle, b, anisochela I, c, anisochelae II, d, anisochela III (anisochela II and III same scale), e, flagellated sigmas." figureDoi="http://doi.org/10.5281/zenodo.4451179" httpUri="https://zenodo.org/record/4451179/files/figure.png" pageId="153" pageNumber="154">Fig. 101b</figureCitation>
), ‘normal’ shaped, with well-developed alae, upper median alae extended outward, free part of the shaft about 40% of spicule length, 33–
<emphasis id="8CCDEAA3FF5CFF3957CFF897534ECA59" box="[755,804,1842,1868]" italics="true" pageId="153" pageNumber="154">36.3</emphasis>
–
<quantity id="79419B54FF5CFF39560FF8965314CA58" box="[819,894,1843,1869]" metricMagnitude="-2" metricUnit="m" metricValue="3.9" pageId="153" pageNumber="154" unit="mm" value="39.0">39 µm</quantity>
.
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBF8F25273CAAC" blockId="153.[151,1437,1086,2014]" pageId="153" pageNumber="154">
Anisochelae II (
<figureCitation id="26822A34FF5CFF3954BCF8F2519ACA64" box="[384,496,1879,1905]" captionStart="FIGURE 101" captionStartId="156.[151,250,1554,1578]" captionTargetBox="[151,1437,181,1530]" captionTargetId="figure-26@156.[151,1437,181,1530]" captionTargetPageId="156" captionText="FIGURE 101. Mycale (Naviculina) mascarenensis sp.nov., SEM images of spicules of holotype RMNH Por. 11723, from Rodrigues, a, mycalostyle, a1, details of mycalostyle, b, anisochela I, c, anisochelae II, d, anisochela III (anisochela II and III same scale), e, flagellated sigmas." figureDoi="http://doi.org/10.5281/zenodo.4451179" httpUri="https://zenodo.org/record/4451179/files/figure.png" pageId="153" pageNumber="154">Fig. 101c</figureCitation>
), naviculichelae, squarish in outline, central plate leaving only small open spaces between the upper and lower alae, rims smooth or with slight upper bump, but not provided with spines, both sides slightly different, 24–
<emphasis id="8CCDEAA3FF5CFF3954B6F83A51D1CAAC" box="[394,443,1951,1977]" italics="true" pageId="153" pageNumber="154">27.3</emphasis>
–
<quantity id="79419B54FF5CFF3954F6F83A527FCAAC" box="[458,533,1951,1977]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="153" pageNumber="154" unit="mm" value="30.0">30 µm</quantity>
.
</paragraph>
<paragraph id="BE0636B1FF5CFF3955FBF866544ACAC8" blockId="153.[151,1437,1086,2014]" box="[199,1056,1986,2014]" pageId="153" pageNumber="154">
Anisochelae III (
<figureCitation id="26822A34FF5CFF3954B8F866519ECACB" box="[388,500,1987,2014]" captionStart="FIGURE 101" captionStartId="156.[151,250,1554,1578]" captionTargetBox="[151,1437,181,1530]" captionTargetId="figure-26@156.[151,1437,181,1530]" captionTargetPageId="156" captionText="FIGURE 101. Mycale (Naviculina) mascarenensis sp.nov., SEM images of spicules of holotype RMNH Por. 11723, from Rodrigues, a, mycalostyle, a1, details of mycalostyle, b, anisochela I, c, anisochelae II, d, anisochela III (anisochela II and III same scale), e, flagellated sigmas." figureDoi="http://doi.org/10.5281/zenodo.4451179" httpUri="https://zenodo.org/record/4451179/files/figure.png" pageId="153" pageNumber="154">Fig. 101d</figureCitation>
), naviculichelae, oval in outline, 15–
<emphasis id="8CCDEAA3FF5CFF3956ADF86653A8CAC8" box="[913,962,1987,2013]" italics="true" pageId="153" pageNumber="154">17.1</emphasis>
–
<quantity id="79419B54FF5CFF3956EEF8665476CAC8" box="[978,1052,1987,2013]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="153" pageNumber="154" unit="mm" value="19.0">19 µm</quantity>
.
</paragraph>
<caption id="EAC66639FF5FFF3A55ABF837550CCADB" ID-DOI="http://doi.org/10.5281/zenodo.4451173" ID-Zenodo-Dep="4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="154" pageNumber="155" startId="154.[151,250,1938,1962]" targetBox="[151,1435,181,1914]" targetPageId="154">
<paragraph id="BE0636B1FF5FFF3A55ABF837550CCADB" blockId="154.[151,1436,1938,1999]" pageId="154" pageNumber="155">
<emphasis id="8CCDEAA3FF5FFF3A55ABF8375170CABF" bold="true" box="[151,282,1938,1962]" pageId="154" pageNumber="155">FIGURE 99</emphasis>
.
<taxonomicName id="79B94D32FF5FFF3A5415F83752FDCABF" authority="Van &amp; Aryasari &amp; De, 2021" authorityName="Van &amp; Aryasari &amp; De" authorityYear="2021" box="[297,663,1938,1962]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="154" pageNumber="155" phylum="Porifera" rank="species" species="mascarenensis" status="sp. nov." subGenus="Naviculina">
<emphasis id="8CCDEAA3FF5FFF3A5415F83752FDCABF" box="[297,663,1938,1962]" italics="true" pageId="154" pageNumber="155">Mycale (Naviculina) mascarenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97FE57D8FF5FFF3A57A2F8375280CABF" box="[670,746,1938,1962]" pageId="154" pageNumber="155" rank="species">
<emphasis id="8CCDEAA3FF5FFF3A57A2F837528ECABF" bold="true" box="[670,740,1938,1962]" pageId="154" pageNumber="155">sp.nov</emphasis>
.
</taxonomicNameLabel>
, habitus
<taxonomicName id="79B94D32FF5FFF3A5675F83653E0CABF" box="[841,906,1939,1962]" class="Demospongiae" family="Irciniidae" genus="Ircinia" kingdom="Animalia" order="Dictyoceratida" pageId="154" pageNumber="47" phylum="Porifera" rank="species" species="situ">
<emphasis id="8CCDEAA3FF5FFF3A5675F83653E0CABF" box="[841,906,1939,1962]" italics="true" pageId="154" pageNumber="155">in situ</emphasis>
</taxonomicName>
, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd).
</paragraph>
</caption>
<caption id="EAC66639FF5EFF3B55ABF9A151D6CB9C" ID-DOI="http://doi.org/10.5281/zenodo.4451175" ID-Zenodo-Dep="4451175" httpUri="https://zenodo.org/record/4451175/files/figure.png" pageId="155" pageNumber="156" startId="155.[151,250,1540,1564]" targetBox="[151,1435,181,1517]" targetPageId="155">
<paragraph id="BE0636B1FF5EFF3B55ABF9A151D6CB9C" blockId="155.[151,1436,1540,1673]" pageId="155" pageNumber="156">
<emphasis id="8CCDEAA3FF5EFF3B55ABF9A15143CB08" bold="true" box="[151,297,1540,1565]" pageId="155" pageNumber="156">FIGURE 100</emphasis>
.
<taxonomicName id="79B94D32FF5EFF3B5404F9A052C2CB09" authority="Van &amp; Aryasari &amp; De, 2021" authorityName="Van &amp; Aryasari &amp; De" authorityYear="2021" box="[312,680,1541,1565]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="155" pageNumber="156" phylum="Porifera" rank="species" species="mascarenensis" status="sp. nov." subGenus="Naviculina">
<emphasis id="8CCDEAA3FF5EFF3B5404F9A052C2CB09" box="[312,680,1541,1565]" italics="true" pageId="155" pageNumber="156">Mycale (Naviculina) mascarenensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="97FE57D8FF5EFF3B578DF9A05297CB09" box="[689,765,1541,1565]" pageId="155" pageNumber="156" rank="species">
<emphasis id="8CCDEAA3FF5EFF3B578DF9A0529DCB08" bold="true" box="[689,759,1541,1565]" pageId="155" pageNumber="156">sp.nov</emphasis>
.
</taxonomicNameLabel>
, habitus and skeleton of holotype RMNH Por. 11689, from Rodrigues, a, on deck, b, detail of preserved specimen (scale bar = 1 cm), c, light microscopic image of ectosomal skeleton showing intercrossing spicule tracts and rosettes of anisochelae I, d, details of skeleton with flagellated sigma, rosettes of anisochelae and scattered naviculichelae.
</paragraph>
</caption>
<paragraph id="BE0636B1FF5EFF3B55FBF912512FCA0C" blockId="155.[151,1437,1719,2033]" pageId="155" pageNumber="156">
Flagellated sigmas (
<figureCitation id="26822A34FF5EFF3B5499F912527DCBC4" box="[421,535,1719,1745]" captionStart="FIGURE 101" captionStartId="156.[151,250,1554,1578]" captionTargetBox="[151,1437,181,1530]" captionTargetId="figure-26@156.[151,1437,181,1530]" captionTargetPageId="156" captionText="FIGURE 101. Mycale (Naviculina) mascarenensis sp.nov., SEM images of spicules of holotype RMNH Por. 11723, from Rodrigues, a, mycalostyle, a1, details of mycalostyle, b, anisochela I, c, anisochelae II, d, anisochela III (anisochela II and III same scale), e, flagellated sigmas." figureDoi="http://doi.org/10.5281/zenodo.4451179" httpUri="https://zenodo.org/record/4451179/files/figure.png" pageId="155" pageNumber="156">Figs 101e</figureCitation>
), strongly curved, asymmetrical endings (reminding strongly of
<taxonomicName id="79B94D32FF5EFF3B51DEF9125259CBE0" authority="Van Soest, 1984" authorityName="Van Soest" authorityYear="1984" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="155" pageNumber="156" phylum="Porifera" rank="species" species="diversisigmata" subGenus="Naviculina">
<emphasis id="8CCDEAA3FF5EFF3B51DEF9125112CBE0" italics="true" pageId="155" pageNumber="156">Mycale (Naviculina) diversisigmata</emphasis>
<bibRefCitation id="DA284B40FF5EFF3B5442F97E5259CBE0" author="Van Soest, R. W. M." box="[382,563,1755,1781]" pageId="155" pageNumber="156" pagination="1 - 167" refId="ref137151" refString="Van Soest, R. W. M. (1984) Marine sponges from Curacao and other Caribbean localities. Part III. Poecilosclerida. In: Hummelinck, P. W. &amp; Van der Steen, L. J. (Eds.), Uitgaven van de Natuurwetenschappelijke Studiekring voor Suriname en de Nederlandse Antillen. No. 112. Studies on the Fauna of Curacao and other Caribbean Islands, 66 (199), 1 - 167." type="journal article" year="1984">Van Soest, 1984</bibRefCitation>
</taxonomicName>
) (cf.
<bibRefCitation id="DA284B40FF5EFF3B5752F97E534ACBE0" author="Van Soest, R. W. M." box="[622,800,1755,1781]" pageId="155" pageNumber="156" pagination="1 - 48" refId="ref137479" refString="Van Soest, R. W. M. (2017) Flagellia, a new subgenus of Haliclona (Porifera, Haplosclerida). European Journal of Taxonomy, 351, 1 - 48. https: // doi. org / 10.5852 / ejt. 2017.351" type="journal article" year="2017">Van Soest 2017</bibRefCitation>
: fig. 2), 47–
<emphasis id="8CCDEAA3FF5EFF3B569AF97E538FCBE0" box="[934,997,1755,1781]" italics="true" pageId="155" pageNumber="156">122.3</emphasis>
–201 x 36–
<emphasis id="8CCDEAA3FF5EFF3B515EF97E54CBCBE0" box="[1122,1185,1755,1781]" italics="true" pageId="155" pageNumber="156">101.2</emphasis>
–
<quantity id="79419B54FF5EFF3B518CF97E5563CBE0" box="[1200,1289,1755,1782]" metricMagnitude="-1" metricUnit="m" metricValue="1.56" pageId="155" pageNumber="156" unit="mm" value="156.0">156 µm</quantity>
. No ‘normal’ sigmas I and II.
</paragraph>
</subSubSection>
<subSubSection id="F6A3653AFF5EFF3B55FBF88655FFCA28" box="[199,1429,1827,1854]" pageId="155" pageNumber="156" type="distribution">
<paragraph id="BE0636B1FF5EFF3B55FBF88655FFCA28" blockId="155.[151,1437,1719,2033]" box="[199,1429,1827,1854]" pageId="155" pageNumber="156">
<emphasis id="8CCDEAA3FF5EFF3B55FBF8865184CA28" bold="true" box="[199,494,1827,1853]" pageId="155" pageNumber="156">Distribution and ecology</emphasis>
.
<collectingRegion id="7C7DF853FF5EFF3B54C7F886521BCA28" box="[507,625,1827,1853]" country="Mauritius" name="Rodrigues" pageId="155" pageNumber="156">Rodrigues</collectingRegion>
, Mascarene Islands (dependency of
<collectingCountry id="C6AE7621FF5EFF3B513BF8865412CA2B" box="[1031,1144,1827,1854]" name="Mauritius" pageId="155" pageNumber="156">Mauritius</collectingCountry>
), on reefs,
<quantity id="79419B54FF5EFF3B51CFF8865522CA28" box="[1267,1352,1827,1853]" metricMagnitude="1" metricUnit="m" metricValue="1.3" metricValueMax="1.8" metricValueMin="0.8" pageId="155" pageNumber="156" unit="m" value="13.0" valueMax="18.0" valueMin="8.0">8–18 m</quantity>
depth.
</paragraph>
</subSubSection>
<subSubSection id="F6A3653AFF5EFF3B55FBF8E25519CA74" box="[199,1395,1863,1889]" pageId="155" pageNumber="156" type="etymology">
<paragraph id="BE0636B1FF5EFF3B55FBF8E25519CA74" blockId="155.[151,1437,1719,2033]" box="[199,1395,1863,1889]" pageId="155" pageNumber="156">
<emphasis id="8CCDEAA3FF5EFF3B55FBF8E25122CA74" bold="true" box="[199,328,1863,1889]" pageId="155" pageNumber="156">Etymology</emphasis>
. Named after the island group of the Mascarenes to which the
<typeStatus id="61028813FF5EFF3B5134F8E25452CA74" box="[1032,1080,1863,1889]" pageId="155" pageNumber="156">type</typeStatus>
locality
<collectingRegion id="7C7DF853FF5EFF3B51A6F8E25567CA74" box="[1178,1293,1863,1889]" country="Mauritius" name="Rodrigues" pageId="155" pageNumber="156">Rodrigues</collectingRegion>
belongs.
</paragraph>
</subSubSection>
<subSubSection id="F6A3653AFF5EFF3B55FBF8CE55F9CAE4" pageId="155" pageNumber="156" type="discussion">
<paragraph id="BE0636B1FF5EFF3B55FBF8CE55F9CAE4" blockId="155.[151,1437,1719,2033]" pageId="155" pageNumber="156">
<emphasis id="8CCDEAA3FF5EFF3B55FBF8CE515ECA90" bold="true" box="[199,308,1899,1925]" pageId="155" pageNumber="156">Remarks</emphasis>
. So far, the new species appears to be endemic to the island of
<collectingRegion id="7C7DF853FF5EFF3B56C1F8CE541ECA90" box="[1021,1140,1899,1925]" country="Mauritius" name="Rodrigues" pageId="155" pageNumber="156">Rodrigues</collectingRegion>
. It is obviously closely related to
<taxonomicName id="79B94D32FF5EFF3B55CEF83551B8CABC" authorityName="Vacelet &amp; Vasseur" authorityYear="1971" box="[242,466,1935,1961]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="155" pageNumber="156" phylum="Porifera" rank="species" species="cleistochela" subGenus="Naviculina">
<emphasis id="8CCDEAA3FF5EFF3B55CEF83551B8CABC" box="[242,466,1935,1961]" italics="true" pageId="155" pageNumber="156">M. (N.) cleistochela</emphasis>
</taxonomicName>
, which shares the possession of flagellated sigmas as well as the ‘normal’ anisochelae I and naviculichelae II and III. The two specimens described here differ clearly from
<taxonomicName id="79B94D32FF5EFF3B5176F811554DCAD8" authorityName="Vacelet &amp; Vasseur" authorityYear="1971" box="[1098,1319,1971,1997]" class="Demospongiae" family="Mycalidae" genus="Mycale" kingdom="Animalia" order="Poecilosclerida" pageId="155" pageNumber="156" phylum="Porifera" rank="species" species="cleistochela" subGenus="Naviculina">
<emphasis id="8CCDEAA3FF5EFF3B5176F811554DCAD8" box="[1098,1319,1971,1997]" italics="true" pageId="155" pageNumber="156">M. (N.) cleistochela</emphasis>
</taxonomicName>
in lacking toxas and ‘normal’ sigmas. Also the
<taxonomicName id="79B94D32FF5EFF3B5713F87D521ECAE4" box="[559,628,2008,2033]" class="Demospongiae" family="Irciniidae" genus="Ircinia" kingdom="Animalia" order="Dictyoceratida" pageId="155" pageNumber="47" phylum="Porifera" rank="species" species="situ">
<emphasis id="8CCDEAA3FF5EFF3B5713F87D521ECAE4" box="[559,628,2008,2033]" italics="true" pageId="155" pageNumber="156">in situ</emphasis>
</taxonomicName>
shapes and surface characters are clearly different (cf.
<figureCitation id="26822A34FF5EFF3B51E1F872555ACAE4" box="[1245,1328,2007,2033]" captionStart="FIGURE 93" captionStartId="147.[151,250,1919,1943]" captionTargetBox="[151,1435,182,1894]" captionTargetId="figure-19@147.[151,1435,181,1894]" captionTargetPageId="147" captionText="FIGURE 93. Mycale (Naviculina) cleistochela Vacelet &amp; Vasseur, 1971, habitus, a, RMNH Por. 1904 in situ, from Bali, Indonesia (photo N.J. de Voogd), b, ditto preserved, c, ZMA Por. 15126 preserved, from SW Sulawesi, Indonesia (scale bar = 1 cm)." figureDoi="http://doi.org/10.5281/zenodo.4451161" httpUri="https://zenodo.org/record/4451161/files/figure.png" pageId="155" pageNumber="156">Figs 93</figureCitation>
and
<figureCitation id="26822A34FF5EFF3B505BF87255EDCAE4" box="[1383,1415,2007,2033]" captionStart="FIGURE 99" captionStartId="154.[151,250,1938,1962]" captionTargetBox="[151,1435,181,1914]" captionTargetId="figure-26@154.[151,1435,181,1914]" captionTargetPageId="154" captionText="FIGURE 99. Mycale (Naviculina) mascarenensis sp.nov., habitus in situ, a, holotype RMNH Por. 11689 from Rodrigues, b, paratype RMNH Por. 11723, also from Rodrigues, c, detail of holotype showing punctate surface (photos N.J. de Voogd)." figureDoi="http://doi.org/10.5281/zenodo.4451173" httpUri="https://zenodo.org/record/4451173/files/figure.png" pageId="155" pageNumber="156">99</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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