195 lines
21 KiB
XML
195 lines
21 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1148.98271" ID-Pensoft-Pub="1313-2970-1148-119" ID-Pensoft-UUID="46E30F13F8295A1F8A4D4DF3DA05E02F" ID-ZooBank="D2805F995B104CB2B2D7F2A7343628D9" ModsDocID="1313-2970-1148-119" checkinTime="1676637833858" checkinUser="pensoft" docAuthor="Wang, Cheng, Li, Shuqiang & Pham, Dinh-Sac" docDate="2023" docId="24CC7EE10B925B8383E28B3B0A0634A4" docLanguage="en" docName="ZooKeys 1148: 119-165" docOrigin="ZooKeys 1148" docPubDate="2023-02-16" docSource="http://dx.doi.org/10.3897/zookeys.1148.98271" docTitle="Eupoa maidinhyeni Wang & Li & Pham 2023, sp. nov." docType="treatment" docUuid="26326AF6-A19C-45DD-A65D-F61D976C5309" docUuidSource="ZooBank" docVersion="1" id="46E30F13F8295A1F8A4D4DF3DA05E02F" lastPageNumber="119" masterDocId="46E30F13F8295A1F8A4D4DF3DA05E02F" masterDocTitle="Thirteen species of jumping spiders from northern Vietnam (Araneae, Salticidae)" masterLastPageNumber="165" masterPageNumber="119" pageNumber="119" updateTime="1676637833858" updateUser="pensoft">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Thirteen species of jumping spiders from northern Vietnam (Araneae, Salticidae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Wang, Cheng</mods:namePart>
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<mods:affiliation>College of Agriculture and Forestry Engineering and Planning, Guizhou Provincial Key Laboratory of Biodiversity Conservation and Utilization in the Fanjing Mountain Region, Tongren University, Tongren 554300, Guizhou, China</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Li, Shuqiang</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-3290-5416</mods:nameIdentifier>
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<mods:affiliation>Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China</mods:affiliation>
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<mods:nameIdentifier type="email">lisq@ioz.ac.cn</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Pham, Dinh-Sac</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8594-5270</mods:nameIdentifier>
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<mods:affiliation>Vietnam National Museum of Nature (VNMN), Vietnam Academy of Science and Technology (VAST), 18 Hoang Quoc Viet, Cau Giay, Hanoi, Vietnam</mods:affiliation>
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<mods:nameIdentifier type="email">phamdinhsac@gmail.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-16</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1148</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>119</mods:start>
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<mods:end>165</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1148.98271</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1148.98271</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1148-119</mods:identifier>
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<mods:identifier type="ZooBank">D2805F995B104CB2B2D7F2A7343628D9</mods:identifier>
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<mods:identifier type="Pensoft-UUID">46E30F13F8295A1F8A4D4DF3DA05E02F</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:26326AF6-A19C-45DD-A65D-F61D976C5309" httpUri="http://treatment.plazi.org/id/24CC7EE10B925B8383E28B3B0A0634A4" lastPageNumber="119" pageId="0" pageNumber="119">
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<subSubSection pageId="0" pageNumber="119" type="nomenclature">
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<paragraph pageId="0" pageNumber="119">
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<taxonomicName LSID="https://zoobank.org/26326AF6-A19C-45DD-A65D-F61D976C5309" authority="Wang & Li & Pham, 2023" authorityName="Wang & Li & Pham" authorityYear="2023" class="Arachnida" family="Salticidae" genus="Eupoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupoa maidinhyeni" order="Araneae" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="maidinhyeni" status="sp. nov.">Eupoa maidinhyeni</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="119">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Eupoa maidinhyeni sp. nov., holotype and paratype A holotype, prolateral B ditto, retrolateral C ditto, ventral D paratype bulb, apical. Scale bars: 0.1 mm. Abbreviations: E - embolus; CTA - compound terminal apophysis; MA - median apophysis; RFA - retrolateral femoral apophysis; RPA - retrolateral patellar apophysis; RTA - retrolateral tibial apophysis; TA - terminal apophysis." figureDoi="10.3897/zookeys.1148.98271.figure5" httpUri="https://binary.pensoft.net/fig/811035" pageId="0" pageNumber="119">Figs 5</figureCitation>
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eupoa maidinhyeni sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca." figureDoi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119">, 6</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="type material">
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<paragraph pageId="0" pageNumber="119">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="119">Holotype</emphasis>
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♂ (IZCAS-Ar44192), Vietnam: Ninh Binh Province: Cuc Phuong National Park, Disturbed Forest (
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<geoCoordinate degrees="20" direction="north" minutes="16.07" orientation="latitude" precision="9" value="20.267834">20°16.07'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="42.04" orientation="longitude" precision="9" value="105.70067">105°42.04'E</geoCoordinate>
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, ca. 250 m), 18.VIII.2007, D.S. Pham leg.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="119">Paratypes</emphasis>
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1♀ (IZCAS-Ar44193), same data as holotype; 2♀ (IZCAS-Ar44194-44195), Cuc Phuong National Park (
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<geoCoordinate degrees="20" direction="north" minutes="16.16" orientation="latitude" precision="9" value="20.269333">20°16.16'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="41.32" orientation="longitude" precision="9" value="105.68867">105°41.32'E</geoCoordinate>
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, ca. 250 m), 9.V.2007, same collector; 1♀ (IZCAS-Ar44196), Cuc Phuong National Park (
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<geoCoordinate degrees="20" direction="north" minutes="23.12" orientation="latitude" precision="9" value="20.385334">20°23.12'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="33.34" orientation="longitude" precision="9" value="105.555664">105°33.34'E</geoCoordinate>
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, ca. 200 m), 7.VI.2007, same collector; 1♀ (IZCAS-Ar44197), Disturbed Forest (
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<geoCoordinate degrees="20" direction="north" minutes="22.27" orientation="latitude" precision="9" value="20.371166">20°22.27'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="33.05" orientation="longitude" precision="9" value="105.550835">105°33.05'E</geoCoordinate>
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, ca. 340 m), 6.XI.2007, same collector; 1♂ (IZCAS-Ar44198), Cuc Phuong National Park (
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<geoCoordinate degrees="20" direction="north" minutes="22.27" orientation="latitude" precision="9" value="20.371166">20°22.27'N</geoCoordinate>
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,
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<geoCoordinate degrees="105" direction="east" minutes="33.05" orientation="longitude" precision="9" value="105.550835">105°33.05'E</geoCoordinate>
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, ca. 200 m), 6.II.2008, same collector.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="etymology">
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<paragraph pageId="0" pageNumber="119">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="119">The specific name is after ichthyologist Mai Dinh Yen, born in 1933 in Ba Vi, Hanoi; noun (name) in genitive case.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="diagnosis">
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<paragraph pageId="0" pageNumber="119">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<taxonomicName authorityName="Wang & Li & Pham" authorityYear="2023" class="Arachnida" family="Salticidae" genus="Eupoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupoa maidinhyeni" order="Araneae" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="maidinhyeni">
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<emphasis italics="true" pageId="0" pageNumber="119">Eupoa maidinhyeni</emphasis>
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</taxonomicName>
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sp. nov. can be easily distinguished from other congeners by the well-developed CTA, which extends beyond the cymbial retromargin in ventral view, the thick, sickle-shaped embolus, and the circled atrial ridges (Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Eupoa maidinhyeni sp. nov., holotype and paratype A holotype, prolateral B ditto, retrolateral C ditto, ventral D paratype bulb, apical. Scale bars: 0.1 mm. Abbreviations: E - embolus; CTA - compound terminal apophysis; MA - median apophysis; RFA - retrolateral femoral apophysis; RPA - retrolateral patellar apophysis; RTA - retrolateral tibial apophysis; TA - terminal apophysis." figureDoi="10.3897/zookeys.1148.98271.figure5" httpUri="https://binary.pensoft.net/fig/811035" pageId="0" pageNumber="119">5C</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eupoa maidinhyeni sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca." figureDoi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119">6A</figureCitation>
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), whereas the CTA does not extend beyond the cymbial margin, the embolus is slender, flagelliform, and lacks similar circled atrial ridges in the other species (
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<bibRefCitation author="Metzner, H" journalOrPublisher="ZooKeys" pageId="0" pageNumber="119" publicationUrl="https://www.jumping-spiders.com" refId="B16" refString="Metzner, H, 2023. Jumping spiders (Arachnida: Araneae: Salticidae) of the world. https://www.jumping-spiders.com" title="Jumping spiders (Arachnida: Araneae: Salticidae) of the world." url="https://www.jumping-spiders.com" year="2023">Metzner 2023</bibRefCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.1148.98271.figure5" httpUri="https://binary.pensoft.net/fig/811035" pageId="0" pageNumber="119" start="Figure 5" startId="F5">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 5.</emphasis>
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Male palp of
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<taxonomicName authorityName="Wang & Li & Pham" authorityYear="2023" class="Arachnida" family="Salticidae" genus="Eupoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupoa maidinhyeni" order="Araneae" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="maidinhyeni">
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<emphasis italics="true" pageId="0" pageNumber="119">Eupoa maidinhyeni</emphasis>
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</taxonomicName>
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sp. nov., holotype and paratype
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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holotype, prolateral
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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ditto, retrolateral
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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ditto, ventral
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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paratype bulb, apical. Scale bars: 0.1 mm. Abbreviations: E - embolus; CTA - compound terminal apophysis; MA - median apophysis; RFA - retrolateral femoral apophysis; RPA - retrolateral patellar apophysis; RTA - retrolateral tibial apophysis; TA - terminal apophysis.
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</paragraph>
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</caption>
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<caption doi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119" start="Figure 6" startId="F6">
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Figure 6.</emphasis>
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<taxonomicName authorityName="Wang & Li & Pham" authorityYear="2023" class="Arachnida" family="Salticidae" genus="Eupoa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Eupoa maidinhyeni" order="Araneae" pageId="0" pageNumber="119" phylum="Arthropoda" rank="species" species="maidinhyeni">
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<emphasis italics="true" pageId="0" pageNumber="119">Eupoa maidinhyeni</emphasis>
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</taxonomicName>
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sp. nov., male holotype and female paratype
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<emphasis bold="true" pageId="0" pageNumber="119">A</emphasis>
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epigyne, ventral
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<emphasis bold="true" pageId="0" pageNumber="119">B</emphasis>
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vulva, dorsal
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<emphasis bold="true" pageId="0" pageNumber="119">C</emphasis>
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holotype habitus, dorsal
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<emphasis bold="true" pageId="0" pageNumber="119">D</emphasis>
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ditto, ventral
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<emphasis bold="true" pageId="0" pageNumber="119">E</emphasis>
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female paratype habitus, dorsal
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<emphasis bold="true" pageId="0" pageNumber="119">F</emphasis>
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holotype carapace, frontal
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<emphasis bold="true" pageId="0" pageNumber="119">G</emphasis>
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holotype chelicera, posterior. Scale bars: 0.1 mm (
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<emphasis bold="true" pageId="0" pageNumber="119">A, B, G</emphasis>
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); 0.5 mm (
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<emphasis bold="true" pageId="0" pageNumber="119">C-F</emphasis>
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). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="description">
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<paragraph pageId="0" pageNumber="119">Description.</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Male</emphasis>
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(Figs
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Eupoa maidinhyeni sp. nov., holotype and paratype A holotype, prolateral B ditto, retrolateral C ditto, ventral D paratype bulb, apical. Scale bars: 0.1 mm. Abbreviations: E - embolus; CTA - compound terminal apophysis; MA - median apophysis; RFA - retrolateral femoral apophysis; RPA - retrolateral patellar apophysis; RTA - retrolateral tibial apophysis; TA - terminal apophysis." figureDoi="10.3897/zookeys.1148.98271.figure5" httpUri="https://binary.pensoft.net/fig/811035" pageId="0" pageNumber="119">5</figureCitation>
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,
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eupoa maidinhyeni sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca." figureDoi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119">6C, D, F, G</figureCitation>
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). Total length 2.13. Carapace 1.05 long, 0.93 wide. Abdomen 1.09 long, 0.72 wide. Clypeus 0.05 high. Eye sizes and inter-distances: AME 0.30, ALE 0.21, PLE 0.18, AERW 0.99, PERW 0.85, EFL 0.57. Legs: I 1.81 (0.55, 0.28, 0.40, 0.38, 0.20), II 1.58 (0.50, 0.25, 0.33, 0.30, 0.20), III 1.61 (0.48, 0.25, 0.30, 0.38, 0.20), IV 2.14 (0.70, 0.28, 0.53, 0.43, 0.20). Carapace longer than wide, pale to dark yellow except the sides of eye field and posterior eyes bases dark, with pair of irregular dark patches medially on eye field, and longitudinal, central, yellow patch extending across thorax; fovea indistinct. Chelicerae pale, with two promarginal and four retromarginal teeth. Endites, sternum colored as chelicerae. Labium slightly darker than endites. Legs pale to yellow, with one ventral spine on tibiae I, and three pairs of ventral spines on metatarsi I. Abdomen elongated, dorsum pale yellow to green-brown, covered wholly by scutum, with pair of longitudinal, irregular, pale yellow stripes on ~ 3/4 of length, and separated by longitudinal, irregular, green brown central stripes and followed by transverse, pale yellow, posterior stripe; venter pale, without markings. Palp (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Male palp of Eupoa maidinhyeni sp. nov., holotype and paratype A holotype, prolateral B ditto, retrolateral C ditto, ventral D paratype bulb, apical. Scale bars: 0.1 mm. Abbreviations: E - embolus; CTA - compound terminal apophysis; MA - median apophysis; RFA - retrolateral femoral apophysis; RPA - retrolateral patellar apophysis; RTA - retrolateral tibial apophysis; TA - terminal apophysis." figureDoi="10.3897/zookeys.1148.98271.figure5" httpUri="https://binary.pensoft.net/fig/811035" pageId="0" pageNumber="119">5A-D</figureCitation>
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): femur longer than wide, with L-shaped disto-retrolateral apophysis; patella slightly longer than wide, with slender, curved, apically pointed retrolateral apophysis broadened proximally and more than half cymbial length in retrolateral view; tibia very short, with flat, broad retrolateral apophysis pointed apically; cymbium acutely tapered, strongly curved ventrally at distal 1/4; bulb almost round in ventral view; MA extending retrolaterally, with membranous base, forming hook at distal end; TA flat, slightly curved medially in ventral view, tapered to rather pointed tip distally; CTA strongly sclerotized, originates from antero-prolateral portion of bulb, extending retrolaterally and beyond retrolateral cymbial margin distally; embolus long, partly invisible, circled and sickle-shaped.
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</paragraph>
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<paragraph pageId="0" pageNumber="119">
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<emphasis bold="true" pageId="0" pageNumber="119">Female</emphasis>
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(Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eupoa maidinhyeni sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca." figureDoi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119">6A, B, E</figureCitation>
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). Total length 2.44. Carapace 1.06 long, 0.96 wide. Abdomen 1.38 long, 0.96 wide. Clypeus 0.05 high. Eye sizes and inter-distances: AME 0.31, ALE 0.21, PLE 0.18, AERW 1.00, PERW 0.95, EFL 0.58. Legs: I 1.86 (0.55, 0.28, 0.45, 0.38, 0.20), II 1.63 (0.50, 0.25, 0.35, 0.33, 0.20), III 1.66 (0.50, 0.25, 0.33, 0.38, 0.20), IV 2.34 (0.75, 0.33, 0.58, 0.48, 0.20). Habitus (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eupoa maidinhyeni sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca." figureDoi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119">6E</figureCitation>
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) similar to that of male except paler, without dorsal abdominal scutum, and with three pairs of ventral spines on tibiae I. Epigyne (Fig.
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<figureCitation captionStart="Figure 6" captionStartId="F6" captionText="Figure 6. Eupoa maidinhyeni sp. nov., male holotype and female paratype A epigyne, ventral B vulva, dorsal C holotype habitus, dorsal D ditto, ventral E female paratype habitus, dorsal F holotype carapace, frontal G holotype chelicera, posterior. Scale bars: 0.1 mm (A, B, G); 0.5 mm (C-F). Abbreviations: AR - atrial ridge; CD - copulatory duct; CO - copulatory opening; S - spermatheca." figureDoi="10.3897/zookeys.1148.98271.figure6" httpUri="https://binary.pensoft.net/fig/811036" pageId="0" pageNumber="119">6A, B</figureCitation>
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): wider than long, with pair of round atria with circled, and arc-shaped ridges; copulatory openings posteriorly located, separated from each other by ~ 2
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<normalizedToken originalValue="×">x</normalizedToken>
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their width; copulatory ducts short, bent medially, connected to mediolateral margins of spermathecae; spermathecae elongate-oval.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="119" type="distribution">
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<paragraph pageId="0" pageNumber="119">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="119">Known only from the type locality in Ninh Binh Province, Vietnam.</paragraph>
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</subSubSection>
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</treatment>
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</document> |