217 lines
22 KiB
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217 lines
22 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zse.99.99030" ID-Pensoft-Pub="1860-0743-1-195" ID-Pensoft-UUID="C59B2291F71D52B89499FD9433B4B57E" ID-ZooBank="E28DD35A30EA4C53B5A6F0A80CA0CFC0" ModsDocID="1860-0743-99-1-195" checkinTime="1678295867757" checkinUser="pensoft" docAuthor="Boonmachai, Tuangthong, Bergey, Elizabeth A. & Nantarat, Nattawadee" docDate="2023" docId="246F60194E505262B837F5A71BAFBC4A" docLanguage="en" docName="ZoosystEvol 99(1): 195-207" docOrigin="Zoosystematics and Evolution 99 (1)" docPubDate="2023-03-07" docSource="http://dx.doi.org/10.3897/zse.99.99030" docTitle="Diplommatina laemsonensis Boonmachai & Nantarat 2023, sp. nov." docType="treatment" docUuid="A1F06C5D-F1EC-41FC-8345-D9BA7693D8E2" docUuidSource="ZooBank" docVersion="2" id="C59B2291F71D52B89499FD9433B4B57E" lastPageNumber="195" masterDocId="C59B2291F71D52B89499FD9433B4B57E" masterDocTitle="First record and description of three new species in the land snail genus Diplommatina Benson, 1849 (Caenogastropoda, Diplommatinidae) from Satun Province, Thailand" masterLastPageNumber="207" masterPageNumber="195" pageNumber="195" updateTime="1678296169138" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>First record and description of three new species in the land snail genus Diplommatina Benson, 1849 (Caenogastropoda, Diplommatinidae) from Satun Province, Thailand</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Boonmachai, Tuangthong</mods:namePart>
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<mods:affiliation>Ph. D. ' s Degree Program in Biology (International Program), Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Bergey, Elizabeth A.</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-7160-6590</mods:nameIdentifier>
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<mods:affiliation>Department of Biology, Faculty of Science, Chiang Mai University 50200, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Nantarat, Nattawadee</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0003-3317-0041</mods:nameIdentifier>
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<mods:affiliation>Ph. D. ' s Degree Program in Biology (International Program), Faculty of Science, Chiang Mai University, Chiang Mai 50200, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">n_nantarat@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zoosystematics and Evolution</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-03-07</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>99</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>1</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>195</mods:start>
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<mods:end>207</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zse.99.99030</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zse.99.99030</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1860-0743-1-195</mods:identifier>
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<mods:identifier type="ZooBank">E28DD35A30EA4C53B5A6F0A80CA0CFC0</mods:identifier>
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<mods:identifier type="Pensoft-UUID">C59B2291F71D52B89499FD9433B4B57E</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:zoobank.org:act:A1F06C5D-F1EC-41FC-8345-D9BA7693D8E2" httpUri="http://treatment.plazi.org/id/246F60194E505262B837F5A71BAFBC4A" lastPageNumber="195" pageId="0" pageNumber="195">
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<subSubSection pageId="0" pageNumber="195" type="nomenclature">
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<paragraph pageId="0" pageNumber="195">
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<taxonomicName LSID="https://zoobank.org/A1F06C5D-F1EC-41FC-8345-D9BA7693D8E2" authority="Boonmachai & Nantarat" authorityName="Boonmachai & Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Diplommatina laemsonensis" order="Mesogastropoda" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="laemsonensis" status="sp. nov.">Diplommatina laemsonensis Boonmachai & Nantarat</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="195">sp. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="description">
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<paragraph pageId="0" pageNumber="195">
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of Diplommatina from Satun Province. A. D. bulonensis sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); B. D. laemsonensis sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); C. D. prakaiphetensis sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); D. D. naiyanetri (CMUZ 905090; Tarutao Island); E. D. canaliculata (CMUZ 9050158; Tharn Pliew Waterfall)." figureDoi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195">Figs 2B</figureCitation>
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">, 3D-F</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="type material">
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<paragraph pageId="0" pageNumber="195">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="195">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="195">Holotype</emphasis>
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CMUZ 9050019 (Fig.
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of Diplommatina from Satun Province. A. D. bulonensis sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); B. D. laemsonensis sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); C. D. prakaiphetensis sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); D. D. naiyanetri (CMUZ 905090; Tarutao Island); E. D. canaliculata (CMUZ 9050158; Tharn Pliew Waterfall)." figureDoi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195">2B</figureCitation>
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); Shell measurements: SH = 2.26 mm, SW = 1.17 mm, AH = 0.77 mm, AW = 0.83 mm, W = 7.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="195">Paratypes</emphasis>
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CMUZ 9050020-9050024 (5 shells); Shell measurements: SH = 2.14-2.49 mm, SW = 1.08-1.21 mm, AH = 0.76-0.91 mm, AW = 0.79-0.86 mm, W = 6 1/2-7.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="type locality">
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<paragraph pageId="0" pageNumber="195">Type locality.</paragraph>
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<paragraph pageId="0" pageNumber="195">
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Thailand, Satun Province, La-ngu District, Limestone Hill,
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<geoCoordinate degrees="6" direction="north" minutes="54" orientation="latitude" precision="1" seconds="43.7" value="6.912139">6°54'43.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="99" direction="east" minutes="41" orientation="longitude" precision="1" seconds="59.0" value="99.69972">99°41'59.0"E</geoCoordinate>
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, 25 December 2020, coll. T. Booonmachai.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="materials_examined">
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<paragraph pageId="0" pageNumber="195">Other material examined.</paragraph>
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<paragraph pageId="0" pageNumber="195">
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<materialsCitation collectingDate="2020-12-25" collectingDateMax="2022-12-21" collectingDateMin="2020-12-25" country="Thailand" county="La-ngu District" latitude="6.912139" location="La-ngu District" longLatPrecision="1" longitude="99.69972" municipality="Limestone Hill" specimenCount="1" stateProvince="Satun Province">
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<collectingCountry name="Thailand">Thailand</collectingCountry>
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,
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<collectingRegion country="Thailand" name="Satun">Satun Province</collectingRegion>
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;
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<collectingCounty>La-ngu District</collectingCounty>
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,
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<collectingMunicipality>Limestone Hill</collectingMunicipality>
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,
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<geoCoordinate degrees="6" direction="north" minutes="54" orientation="latitude" precision="1" seconds="43.7" value="6.912139">6°54'43.7"N</geoCoordinate>
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,
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<geoCoordinate degrees="99" direction="east" minutes="41" orientation="longitude" precision="1" seconds="59.0" value="99.69972">99°41'59.0"E</geoCoordinate>
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,
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<collectingDate value="2020-12-25">25 December 2020</collectingDate>
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: CMUZ 9050025-9050047 (23 shells);
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<location LSID="urn:lsid:plazi:treatment:246F60194E505262B837F5A71BAFBC4A:E341DD72EDEECE920B6CD9227CA9B2D7" country="Thailand" county="La-ngu District" latitude="6.912139" longLatPrecision="1" longitude="99.69972" municipality="Limestone Hill" name="La-ngu District" stateProvince="Satun Province">La-ngu District</location>
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,
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<location LSID="urn:lsid:plazi:treatment:246F60194E505262B837F5A71BAFBC4A:34519609DD72ECBFEACDA3923F1EBB9F" country="Thailand" county="La-ngu District" latitude="6.912139" longLatPrecision="1" longitude="99.69972" municipality="Limestone Hill" name="Khoa Yai Island" stateProvince="Satun Province">Khoa Yai Island</location>
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, 6°83'08.6"N, 99°69'70.1"E,
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<collectingDate value="2022-12-21">21 December 2022</collectingDate>
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: CMUZ 9050048-9050065 (18 shells)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="etymology">
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<paragraph pageId="0" pageNumber="195">Etymology.</paragraph>
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<paragraph pageId="0" pageNumber="195">
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The specific epithet
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<emphasis italics="true" pageId="0" pageNumber="195">laemsonensis</emphasis>
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is an adjective referring to the type locality (Laem Son subdistrict, Lan-gu district, Satun Province, Thailand).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="195">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="195">
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Among the sinistral species of
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<taxonomicName authorityName="Benson" authorityYear="1849" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina</emphasis>
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</taxonomicName>
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,
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<taxonomicName authorityName="Boonmachai & Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina laemsonensis" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="laemsonensis">
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<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina laemsonensis</emphasis>
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</taxonomicName>
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sp. nov. is most similar to
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<taxonomicName lsidName="D. diminuta" pageId="0" pageNumber="195" rank="species" species="diminuta">
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<emphasis italics="true" pageId="0" pageNumber="195">D. diminuta</emphasis>
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</taxonomicName>
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<normalizedToken originalValue="Möllendorff">Moellendorff</normalizedToken>
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, 1891,
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<taxonomicName lsidName="D. sinistra" pageId="0" pageNumber="195" rank="species" species="sinistra">
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<emphasis italics="true" pageId="0" pageNumber="195">D. sinistra</emphasis>
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</taxonomicName>
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Tomlin, 1938 and
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<taxonomicName lsidName="D. acme" pageId="0" pageNumber="195" rank="species" species="acme">
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<emphasis italics="true" pageId="0" pageNumber="195">D. acme</emphasis>
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</taxonomicName>
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Laidlaw, 1949 (Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P <0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
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). All these species share a moderately slender fusiform shape, rounded periphery, shell height range and number of whorls (Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P <0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
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). However,
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<taxonomicName authorityName="Boonmachai & Nantarat" authorityYear="2023" class="Gastropoda" family="Diplommatinidae" genus="Diplommatina" kingdom="Animalia" lsidName="Diplommatina laemsonensis" order="Stylommatophora" pageId="0" pageNumber="195" phylum="Mollusca" rank="species" species="laemsonensis">
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<emphasis italics="true" pageId="0" pageNumber="195">Diplommatina laemsonensis</emphasis>
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</taxonomicName>
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differs from
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<taxonomicName lsidName="D. acme" pageId="0" pageNumber="195" rank="species" species="acme">
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<emphasis italics="true" pageId="0" pageNumber="195">D. acme</emphasis>
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</taxonomicName>
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,
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<taxonomicName lsidName="D. diminuta" pageId="0" pageNumber="195" rank="species" species="diminuta">
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<emphasis italics="true" pageId="0" pageNumber="195">D. diminuta</emphasis>
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</taxonomicName>
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and
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<taxonomicName lsidName="D. sinistra" pageId="0" pageNumber="195" rank="species" species="sinistra">
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<emphasis italics="true" pageId="0" pageNumber="195">D. sinistra</emphasis>
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</taxonomicName>
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in the number of radial ribs in the penultimate whorl (Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P <0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
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). It differs from
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<taxonomicName lsidName="D. diminuta" pageId="0" pageNumber="195" rank="species" species="diminuta">
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<emphasis italics="true" pageId="0" pageNumber="195">D. diminuta</emphasis>
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</taxonomicName>
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in shell width and the ratio of AH/AW (Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P <0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
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). Moreover, it differs from
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<taxonomicName lsidName="D. diminuta" pageId="0" pageNumber="195" rank="species" species="diminuta">
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<emphasis italics="true" pageId="0" pageNumber="195">D. diminuta</emphasis>
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</taxonomicName>
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by the absence of teeth on the basal margin of the aperture and differs from
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<taxonomicName lsidName="D. sinistra" pageId="0" pageNumber="195" rank="species" species="sinistra">
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<emphasis italics="true" pageId="0" pageNumber="195">D. sinistra</emphasis>
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</taxonomicName>
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in having a distinct columellar denticle in the aperture.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="195" type="description">
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<paragraph pageId="0" pageNumber="195">Description.</paragraph>
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<paragraph pageId="0" pageNumber="195">
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Shell sinistral, fusiform, convex and honey brown in color (Figs
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<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Shell morphology of Diplommatina from Satun Province. A. D. bulonensis sp. nov. (Holotype CMUZ 9050001; Bulon Pai Island); B. D. laemsonensis sp. nov. (Holotype CMUZ 9050019; Laem Son Hill); C. D. prakaiphetensis sp. nov. (Holotype CMUZ 9050066; Prakaiphet Hill); D. D. naiyanetri (CMUZ 905090; Tarutao Island); E. D. canaliculata (CMUZ 9050158; Tharn Pliew Waterfall)." figureDoi="10.3897/zse.99.99030.figure2" httpUri="https://binary.pensoft.net/fig/820985" pageId="0" pageNumber="195">2B</figureCitation>
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,
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">3D</figureCitation>
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). The penultimate whorl widest. Spire conical with slightly convex sides. Suture impressed. Constriction level with the edge between the parietal and columellar side of the peristome, with three lamellae: one distinct parietalis which starts at the constriction, one transversal palatalis, one distinct columellaris which continues into the tuba and visible in the aperture (Fig.
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Shell internal terminology of new species genus Diplommatina from Satun Province under the stereo microscope and the drawing. A, B. D. bulonensis sp. nov. (Bulon Pai Island); C, D. D. laemsonensis sp. nov. (Laem Son Hill); E, F. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure4" httpUri="https://binary.pensoft.net/fig/820987" pageId="0" pageNumber="195">4C, D</figureCitation>
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). Protoconch sculpture smooth with fine pitting and about 1 whorl (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">3E</figureCitation>
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). Radial ribs straight, distinct, rather dense on top of the teleoconch, suddenly changing to a moderately spaced about 8-9 ribs/0.5 mm on the penultimate whorl (Table
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<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of shell morphological characters of closely related Diplommatina species with new species. Shell morphological characters were tested using one-way ANOVA (P <0.05)." httpUri="http://table.plazi.org/id/EF8D3F539259B811645CFEF5AF2E8FAE" pageId="0" pageNumber="195" tableUuid="EF8D3F539259B811645CFEF5AF2E8FAE">1</tableCitation>
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). The aperture tilted up to 30 °C in relation to the coiling axis, columellar denticle visible in the aperture, deflected downwards. Peristome double, expanding, palatal side rounded with slightly edge, basal side with edge slightly protruding. Outer peristome expands beyond the inner peristome with about four layers, inner peristome with distinct a palatal, basal and columellar lips. Umbilicus closed. Operculum multispiral, flat, corneous, transparent, slightly concave, outer surface smooth with small pits distributed over whole surface, inner surface smooth with raised peripheral circular margin (Fig.
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<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Scanning electron microscope of shell morphology, protoconch and operculum of Diplommatina new species. A-C. D. bulonensis sp. nov. (Bulon Pai Island); D-F. D. laemsonensis sp. nov. (Laem Son Hill); G-I. D. prakaiphetensis sp. nov. (Prakaiphet Hill)." figureDoi="10.3897/zse.99.99030.figure3" httpUri="https://binary.pensoft.net/fig/820986" pageId="0" pageNumber="195">3F</figureCitation>
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). The radula of the taenioglossate type (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Radular morphology of Diplommatina. A. Diplommatina laemsonensis sp. nov. (Laem Son Hill); B. Diplommatina bulonensis sp. nov. (Bulon Pai Island); C. Diplommatina prakaiphetensis sp. nov. (Prakaiphet Hill). Colours show the position of the teeth of the radula; blue: central tooth, yellow: the basal plate of the central tooth, green: lateral teeth, pink: inner marginal teeth, and red: outer marginal teeth." figureDoi="10.3897/zse.99.99030.figure5" httpUri="https://binary.pensoft.net/fig/820988" pageId="0" pageNumber="195">5A</figureCitation>
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). Central tooth strong with a large central cusp, with 2 pairs of developed lateral cusps. Basal plate of central tooth prominent with 2 small cusps. The lateral teeth with six cusps, the third cusp longest. In the marginal teeth, the inner teeth larger than the outer, with six cusps, the third cusp longest. The outer tooth with four cusps (Fig.
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<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Radular morphology of Diplommatina. A. Diplommatina laemsonensis sp. nov. (Laem Son Hill); B. Diplommatina bulonensis sp. nov. (Bulon Pai Island); C. Diplommatina prakaiphetensis sp. nov. (Prakaiphet Hill). Colours show the position of the teeth of the radula; blue: central tooth, yellow: the basal plate of the central tooth, green: lateral teeth, pink: inner marginal teeth, and red: outer marginal teeth." figureDoi="10.3897/zse.99.99030.figure5" httpUri="https://binary.pensoft.net/fig/820988" pageId="0" pageNumber="195">5A</figureCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |