201 lines
21 KiB
XML
201 lines
21 KiB
XML
<document id="5EBCA5EB2913B989EBD2CF46AD355451" ID-CLB-Dataset="30654" ID-DOI="10.11646/zootaxa.4387.1.2" ID-GBIF-Dataset="4958a805-21f8-430b-a1c3-d848dfe5d90e" ID-ISSN="1175-5326" ID-Zenodo-Dep="1186533" ID-ZooBank="C679BD1D-6F0A-4347-B7A7-9C527A8BA675" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1519897358538" checkinUser="plazi" docAuthor="Fišer, Cene, Zagmajster, Maja & Dethier, Michel" docDate="2018" docId="241C87DE2D2CB05BFF10FECFC22AFB53" docLanguage="en" docName="zootaxa.4387.1.2.pdf" docOrigin="Zootaxa 4387 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Niphargus pachypus Schellenberg 1933" docType="treatment" docVersion="7" lastPageNumber="53" masterDocId="D825FFA62D2AB05DFF87FFE8C008FFCD" masterDocTitle="Overview of Niphargidae (Crustacea: Amphipoda) in Belgium: distribution, taxonomic notes and conservation issues" masterLastPageNumber="74" masterPageNumber="47" pageNumber="53" updateTime="1698524409052" updateUser="ExternalLinkService">
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<mods:mods id="49D9942E4BDB5D70D53EE9C212482CE4" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="CE2767550FABFCBB288E875CDEF6CE2F">
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<mods:title id="BF0DF8EFD9261A2AFD9A595F490BE7E9">Overview of Niphargidae (Crustacea: Amphipoda) in Belgium: distribution, taxonomic notes and conservation issues</mods:title>
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<mods:name id="C153EF0DB03886078F0EB570B6D0D1E7" type="personal">
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<mods:namePart id="CBA910F8FC70DB0B9AFF45603A083AF1">Fišer, Cene</mods:namePart>
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<mods:namePart id="B7AF42C29D2711BD18B6AC1ADFC39E3F">Zagmajster, Maja</mods:namePart>
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<mods:roleTerm id="19B1CC146BE4B1431B6381D4B5687A0B">Author</mods:roleTerm>
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<mods:namePart id="4BFA043927F32950373B1A167B7732E5">Dethier, Michel</mods:namePart>
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<mods:typeOfResource id="4C219F90C1CB30F982892C2A6939CF08">text</mods:typeOfResource>
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<mods:title id="9923839F7422152A8E900F8E7A3F572F">Zootaxa</mods:title>
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<mods:part id="19C2025058375C68C6A7BC0E0EC929BA">
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<mods:date id="6525982C70DE8EDC13F40F7D043D8DB8">2018</mods:date>
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<mods:number id="756C802F8CA97F81C0C8812B93B30A39">2018-02-26</mods:number>
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<mods:number id="F46EF63A43DE0496836FF4993B4FDEB4">4387</mods:number>
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<mods:detail id="99ECEBC0AD6AF9A57D842C2E1C72682A" type="issue">
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<mods:number id="37AC1BC17E1818C5B991E1C53D808079">1</mods:number>
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<mods:identifier id="022A1ED83A10228921CF813465F8624E" type="DOI">10.11646/zootaxa.4387.1.2</mods:identifier>
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<treatment id="241C87DE2D2CB05BFF10FECFC22AFB53" ID-DOI="http://doi.org/10.5281/zenodo.5989012" ID-GBIF-Taxon="141251076" ID-Zenodo-Dep="5989012" LSID="urn:lsid:plazi:treatment:241C87DE2D2CB05BFF10FECFC22AFB53" httpUri="http://treatment.plazi.org/id/241C87DE2D2CB05BFF10FECFC22AFB53" lastPageNumber="53" pageId="6" pageNumber="53">
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<subSubSection id="E4AF65432D2CB05BFF10FECFC13AFDF6" pageId="6" pageNumber="53" type="nomenclature">
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<paragraph id="AC0A36C82D2CB05BFF10FECFC28BFE8F" blockId="6.[151,643,295,322]" box="[151,643,295,322]" pageId="6" pageNumber="53">
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<heading id="F74281A42D2CB05BFF10FECFC28BFE8F" bold="true" box="[151,643,295,322]" fontSize="11" level="1" pageId="6" pageNumber="53" reason="1">
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<taxonomicName id="6BB54D4B2D2CB05BFF10FECFC28BFE8F" ID-CoL="47G63" authority="Schellenberg, 1933" authorityName="Schellenberg" authorityYear="1933" box="[151,643,295,322]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="pachypus">
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<emphasis id="9EC1EADA2D2CB05BFF10FECFC28BFE8F" bold="true" box="[151,643,295,322]" pageId="6" pageNumber="53">
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<emphasis id="9EC1EADA2D2CB05BFF10FECFC199FE8C" bold="true" box="[151,401,295,321]" italics="true" pageId="6" pageNumber="53">Niphargus pachypus</emphasis>
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<bibRefCitation id="C8244B392D2CB05BFE1FFEC0C28BFE8F" box="[408,643,296,322]" pageId="6" pageNumber="53" refId="ref13983" refString="Schellenberg, A. (1933) Die Niphargiden der Brusseler Naturhistorischen Museum. Bulletin du Musee royal d'Histoire naturelle de Belgique, 9 (50), 1 - 8." type="journal article">Schellenberg, 1933</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="AC0A36C82D2CB05BFF10FE86C13AFDF6" blockId="6.[151,1436,366,1183]" pageId="6" pageNumber="53">
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<emphasis id="9EC1EADA2D2CB05BFF10FE86C2C9FE4A" bold="true" box="[151,705,366,391]" pageId="6" pageNumber="53">Published records: L.112—Sainte Anne cave</emphasis>
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(
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<bibRefCitation id="C8244B392D2CB05BFD52FE86C37DFE4A" box="[725,885,366,391]" pageId="6" pageNumber="53" refId="ref13248" refString="Leruth, R. (1934 a) Exploration biologique des cavernes de la Belgique et du Limbourg hollandais. XVIIIeme contribution. Crustaces aquatiques. Natuurhistorisch Maandblad, 23 (11), 146 - 149." type="journal article">Leruth 1934a</bibRefCitation>
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,
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<bibRefCitation id="C8244B392D2CB05BFC01FE86C3C7FE4A" box="[902,975,366,391]" pageId="6" pageNumber="53" refId="ref13286" refString="Leruth, R. (1934 b) Exploration biologique des cavernes de la Belgique et du Limbourg hollandais. XVIIIeme contribution. Crustaces aquatiques (suite). Natuurhistorisch Maandblad, 23 (12), 156 - 159." type="journal article">1934b</bibRefCitation>
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,
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<bibRefCitation id="C8244B392D2CB05BFC67FE86C421FE4A" box="[992,1065,366,391]" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">1939b</bibRefCitation>
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,
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<bibRefCitation id="C8244B392D2CB05BFBBEFE86C513FE4A" box="[1081,1307,366,391]" pageId="6" pageNumber="53" refId="ref11187" refString="Delhez, F., Dethier, M. & Hubart, J. - M. (1999) Contribution a la connaissance de la faune des grottes de la Wallonie. Bulletin des Chercheurs de la Wallonie, 39, 27 - 54." type="journal article">
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Delhez
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<emphasis id="9EC1EADA2D2CB05BFB12FE87C4DCFE4B" box="[1173,1236,366,390]" italics="true" pageId="6" pageNumber="53">et al.</emphasis>
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1999
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</bibRefCitation>
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, Hubart & Dethier 1999, ZMB Berlin);
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<emphasis id="9EC1EADA2D2CB05BFE6EFE7AC318FE66" bold="true" box="[489,784,402,427]" pageId="6" pageNumber="53">L.132—Trou des Sottais</emphasis>
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cave (
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<bibRefCitation id="C8244B392D2CB05BFCE6FE7AC409FE66" box="[865,1025,402,427]" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">Leruth 1939b</bibRefCitation>
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, Hubart & Dethier 1999);
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<emphasis id="9EC1EADA2D2CB05BFABBFE7AC1CAFE02" bold="true" pageId="6" pageNumber="53">L.146— Spring S.1 of La Xhavée</emphasis>
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(
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<bibRefCitation id="C8244B392D2CB05BFE55FE5EC278FE02" box="[466,624,438,463]" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">Leruth 1939b</bibRefCitation>
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, Hubart & Dethier 1999);
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<emphasis id="9EC1EADA2D2CB05BFC24FE5EC4D7FE02" bold="true" box="[931,1247,438,463]" pageId="6" pageNumber="53">L.149—Ouffet, Spring S.1</emphasis>
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(
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<bibRefCitation id="C8244B392D2CB05BFB68FE5EC587FE02" box="[1263,1423,438,463]" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">Leruth 1939b</bibRefCitation>
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);
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<emphasis id="9EC1EADA2D2CB05BFF10FE32C283FE3E" bold="true" box="[151,651,474,499]" pageId="6" pageNumber="53">L.152—Gravel of Meuse river, phreatic</emphasis>
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(
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<bibRefCitation id="C8244B392D2CB05BFD25FE32C34FFE3E" box="[674,839,474,499]" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">Leruth 1939b</bibRefCitation>
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);
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<emphasis id="9EC1EADA2D2CB05BFCE4FE32C594FE3E" bold="true" box="[867,1436,474,499]" pageId="6" pageNumber="53">L.153—Hermalle sous Argenteau, pump well</emphasis>
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(Hubart & Dethier 1999);
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<emphasis id="9EC1EADA2D2CB05BFE47FE16C2CDFDDA" bold="true" box="[448,709,510,535]" pageId="6" pageNumber="53">L.157—Waha springs</emphasis>
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(
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<bibRefCitation id="C8244B392D2CB05BFD52FE16C406FDDA" box="[725,1038,510,535]" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">Leruth 1939b, ZMB Berlin</bibRefCitation>
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);
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<emphasis id="9EC1EADA2D2CB05BFBA5FE16C4E3FDDA" bold="true" box="[1058,1259,510,535]" pageId="6" pageNumber="53">L.169—Nys cave</emphasis>
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(
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<bibRefCitation id="C8244B392D2CB05BFB7DFE16C12EFDF6" pageId="6" pageNumber="53" refId="ref13411" refString="Leruth, R. (1939 b) La biologie du domaine souterrain et la faune cavernicole de la Belgique. Memoires du Musee royal d'Histoire naturelle, 87, 1 - 506." type="journal article">Leruth 1939b, ZMB Berlin</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="E4AF65432D2CB05BFF40FDAEC22AFB53" pageId="6" pageNumber="53" type="description">
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<paragraph id="AC0A36C82D2CB05BFF40FDAEC22BFD22" blockId="6.[151,1436,366,1183]" pageId="6" pageNumber="53">
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<emphasis id="9EC1EADA2D2CB05BFF40FDAEC2E0FD92" bold="true" box="[199,744,582,607]" pageId="6" pageNumber="53">New records: L.110—Ru de Targnon, brook</emphasis>
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(
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<date id="D80B10082D2CB05BFD7DFDAFC367FD92" box="[762,879,583,607]" pageId="6" pageNumber="53" value="2000-08-23">23.8.2000</date>
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, MD;
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<date id="D80B10082D2CB05BFC39FDAEC438FD92" box="[958,1072,582,607]" pageId="6" pageNumber="53" value="2002-06-11">6.11.2002</date>
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, MD & JMH);
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<emphasis id="9EC1EADA2D2CB05BFB6AFDAEC1B8FD4E" bold="true" pageId="6" pageNumber="53">L.113—Spring close to Monceau cave</emphasis>
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(
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<date id="D80B10082D2CB05BFE44FD82C24EFD4E" box="[451,582,618,643]" pageId="6" pageNumber="53" value="2001-04-06">04.06.2001</date>
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, MD & JMH; 06-
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<date id="D80B10082D2CB05BFCA6FD82C3ACFD4E" box="[801,932,618,643]" pageId="6" pageNumber="53" value="2002-12-06">12.06.2002</date>
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, MD);
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<emphasis id="9EC1EADA2D2CB05BFC79FD82C0E0FD6A" bold="true" pageId="6" pageNumber="53">L.123—Blanc Gravier, brook, G2 station</emphasis>
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(
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<date id="D80B10082D2CB05BFF79FD67C17BFD6A" box="[254,371,655,679]" pageId="6" pageNumber="53" value="2003-04-29">29.4.2003</date>
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, MD);
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<emphasis id="9EC1EADA2D2CB05BFE54FD66C456FD6A" bold="true" box="[467,1118,654,679]" pageId="6" pageNumber="53">L.141—Petites Soeurs des Pauvres, draining gallery</emphasis>
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(
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<date id="D80B10082D2CB05BFBF3FD67C4E1FD6A" box="[1140,1257,655,679]" pageId="6" pageNumber="53" value="2002-03-23">23.3.2002</date>
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, MD & JMH;
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<date id="D80B10082D2CB05BFF10FD5AC101FD06" box="[151,265,690,715]" pageId="6" pageNumber="53" value="2003-05-12">5.12.2003</date>
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, MD & JMH);
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<emphasis id="9EC1EADA2D2CB05BFE45FD5AC370FD06" bold="true" box="[450,888,690,715]" pageId="6" pageNumber="53">L.148—Rhieux, underground brook</emphasis>
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(
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<date id="D80B10082D2CB05BFC0DFD5AC3F7FD06" box="[906,1023,690,715]" pageId="6" pageNumber="53" value="2006-03-23">23.3.2006</date>
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, MD & JMH);
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<emphasis id="9EC1EADA2D2CB05BFB30FD5AC15FFD22" bold="true" pageId="6" pageNumber="53">L.150—Argenteau, draining gallery</emphasis>
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(
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<date id="D80B10082D2CB05BFEE2FD3EC1D1FD22" box="[357,473,726,751]" pageId="6" pageNumber="53" value="2006-06-15">15.6.2006</date>
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, MD).
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</paragraph>
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<paragraph id="AC0A36C82D2CB05BFF40FD12C36AFC0A" blockId="6.[151,1436,366,1183]" pageId="6" pageNumber="53">
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The species was described as
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<taxonomicName id="6BB54D4B2D2CB05BFDBAFD13C358FCDE" authorityName="Schellenberg" authorityYear="1933" box="[573,848,763,787]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="kochianus" subSpecies="pachypus">
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<emphasis id="9EC1EADA2D2CB05BFDBAFD13C358FCDE" box="[573,848,763,787]" italics="true" pageId="6" pageNumber="53">N. kochianus pachypus</emphasis>
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</taxonomicName>
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from
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<collectingCountry id="D4A276582D2CB05BFC23FD13C3F8FCDF" box="[932,1008,763,786]" name="France" pageId="6" pageNumber="53">France</collectingCountry>
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(
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<bibRefCitation id="C8244B392D2CB05BFB80FD12C4EEFCDE" box="[1031,1254,762,787]" pageId="6" pageNumber="53" refId="ref13983" refString="Schellenberg, A. (1933) Die Niphargiden der Brusseler Naturhistorischen Museum. Bulletin du Musee royal d'Histoire naturelle de Belgique, 9 (50), 1 - 8." type="journal article">Schellenberg 1933</bibRefCitation>
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). The taxon is distributed in
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<collectingCountry id="D4A276582D2CB05BFEB3FCF7C18BFCFB" box="[308,387,799,822]" name="France" pageId="6" pageNumber="53">France</collectingCountry>
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,
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<collectingCountry id="D4A276582D2CB05BFE09FCF6C1FAFCFA" box="[398,498,798,823]" name="Belgium" pageId="6" pageNumber="53">Belgium</collectingCountry>
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,
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<collectingRegion id="6E71F82A2D2CB05BFE7AFCF6C285FCFA" box="[509,653,798,823]" country="Luxembourg" name="Luxembourg" pageId="6" pageNumber="53">
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<collectingCountry id="D4A276582D2CB05BFE7AFCF6C285FCFA" box="[509,653,798,823]" name="Luxembourg" pageId="6" pageNumber="53">Luxembourg</collectingCountry>
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</collectingRegion>
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and
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<collectingCountry id="D4A276582D2CB05BFD42FCF6C344FCFA" box="[709,844,798,823]" name="Netherlands" pageId="6" pageNumber="53">Netherlands</collectingCountry>
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(
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<bibRefCitation id="C8244B392D2CB05BFCDCFCF6C469FCFA" box="[859,1121,798,823]" pageId="6" pageNumber="53" refId="ref14385" refString="Stock, J. & Gledhill T. (1977) The Niphargus kochianus - group in North-Western Europe. Crustaceana, 4 (Supplement), 212 - 243." type="journal article">Stock & Gledhill 1977</bibRefCitation>
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). In
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<collectingCountry id="D4A276582D2CB05BFB14FCF6C4FFFCFA" box="[1171,1271,798,823]" name="Belgium" pageId="6" pageNumber="53">Belgium</collectingCountry>
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, it is restricted to karst areas, where it was collected along rivers. In newly collected material we found the species only in the localities around Liège in
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<collectingRegion id="6E71F82A2D2CB05BFE38FC8FC21BFCB3" box="[447,531,871,894]" country="American Samoa" name="Eastern" pageId="6" pageNumber="53">Eastern</collectingRegion>
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part of the country (
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<figureCitation id="348E2A4D2D2CB05BFD78FC8EC336FCB2" box="[767,830,870,895]" captionStart="FIGURE 4" captionStartId="6.[151,250,1773,1795]" captionTargetBox="[151,1436,1238,1752]" captionTargetId="figure@6.[151,1436,1238,1752]" captionTargetPageId="6" captionText="FIGURE 4. Distribution of N.pachypus (left) and N.puteanus (right) in Belgium. Blue circles refer to data from published sources, while red circles refer to new records published for the first time in this study." httpUri="https://zenodo.org/record/1186541/files/figure.png" pageId="6" pageNumber="53">Fig.4</figureCitation>
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). We collected the species mainly in springs, but also caves (often in “gours”
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<emphasis id="9EC1EADA2D2CB05BFE2DFC62C1CEFC6E" bold="true" box="[426,454,906,931]" pageId="6" pageNumber="53">—</emphasis>
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little pools in the calcite reservoirs; see the list, App. 1). The species seems to be generally bound to small pore habitats (
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<bibRefCitation id="C8244B392D2CB05BFDD5FC46C35DFC0A" box="[594,853,942,967]" pageId="6" pageNumber="53" refId="ref14385" refString="Stock, J. & Gledhill T. (1977) The Niphargus kochianus - group in North-Western Europe. Crustaceana, 4 (Supplement), 212 - 243." type="journal article">Stock & Gledhill 1977</bibRefCitation>
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).
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</paragraph>
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<paragraph id="AC0A36C82D2CB05BFF40FC3AC22AFB53" blockId="6.[151,1436,366,1183]" pageId="6" pageNumber="53">
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Already
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<bibRefCitation id="C8244B392D2CB05BFEABFC3AC24FFC26" box="[300,583,978,1003]" pageId="6" pageNumber="53" refId="ref14385" refString="Stock, J. & Gledhill T. (1977) The Niphargus kochianus - group in North-Western Europe. Crustaceana, 4 (Supplement), 212 - 243." type="journal article">Stock & Gledhill (1977)</bibRefCitation>
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erected
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<taxonomicName id="6BB54D4B2D2CB05BFD2AFC3BC352FC26" authorityName="Schellenberg" authorityYear="1933" box="[685,858,979,1003]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="subSpecies" species="kochianus" subSpecies="pachypus">
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<emphasis id="9EC1EADA2D2CB05BFD2AFC3BC352FC26" box="[685,858,979,1003]" italics="true" pageId="6" pageNumber="53">N. k. pachypus</emphasis>
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</taxonomicName>
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from subspecies to species level, and subsequent molecular analyses suggest that the species is not related to
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<taxonomicName id="6BB54D4B2D2CB05BFCC4FC1FC3DEFBC3" authority="Fišer & Zagmajster & Dethier, 2018" authorityName="Fišer & Zagmajster & Dethier" authorityYear="2018" box="[835,982,1014,1038]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" sensu="stricto" species="kochianus">
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<emphasis id="9EC1EADA2D2CB05BFCC4FC1FC3DEFBC3" box="[835,982,1014,1038]" italics="true" pageId="6" pageNumber="53">N. kochianus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="85F257A12D2CB05BFC58FC1FC413FBC3" box="[991,1051,1015,1038]" pageId="6" pageNumber="53" sensu="stricto">s. str.</taxonomicNameLabel>
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(McInnerney
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<emphasis id="9EC1EADA2D2CB05BFB3AFC1FC4FFFBC3" box="[1213,1271,1014,1038]" italics="true" pageId="6" pageNumber="53">et al.</emphasis>
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2014). Unlike previous species, this species shows relatedness towards niphargids from continental Europe. Phylogeography and population structure of this species was never studied, but indirect evidence from
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<taxonomicName id="6BB54D4B2D2CB05BFBCDFBD7C4A8FB9B" authorityName="Chevreux" authorityYear="2001" box="[1098,1184,1087,1110]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="virei">
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<emphasis id="9EC1EADA2D2CB05BFBCDFBD7C4A8FB9B" box="[1098,1184,1087,1110]" italics="true" pageId="6" pageNumber="53">N. virei</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="6BB54D4B2D2CB05BFB5BFBD7C59DFB9B" authorityName="S. Karaman" authorityYear="1932" box="[1244,1429,1086,1110]" class="Malacostraca" family="Niphargidae" genus="Niphargus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Amphipoda" pageId="6" pageNumber="53" phylum="Arthropoda" rank="species" species="schellenbergi">
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<emphasis id="9EC1EADA2D2CB05BFB5BFBD7C59DFB9B" box="[1244,1429,1086,1110]" italics="true" pageId="6" pageNumber="53">N. schellenbergi</emphasis>
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</taxonomicName>
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, derived from similar distributional patterns (see below), suggests that some phylogeographic structure and perhaps cryptic species cannot be ruled out.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |