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<document id="80B09D1E53216D2E33EFED042287DB12" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630D652E9EF3BFB5B75800289" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Ontherus aphodioides Burmeister 1874" docType="treatment" docVersion="6" lastPageNumber="142" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="141" updateTime="1731448046962" updateUser="carolina" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="A3310DDF84446F8247C07B7268C01C3D">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="D228E6485804BDBDC7C50C8D90695AEF">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="D41B36D8B6913D1B061BB9F156406BF7">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="6698963748B82590E64C29D0C27D16AA">Cupello, Mario</mods:namePart>
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<mods:affiliation id="50861276E068B7B6BC40498EAC22F7A9">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="323C469244FDE197206805AD8FBDCB23" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="AC55176087ADA82187CF85C88D999B5E">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="40133CD0C1F19D668A841FFFBB07A36F">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<subSubSection id="DDE394FB30D652E8EF3BFB5B73040428" box="[556,1032,1276,1303]" pageId="140" pageNumber="141" type="nomenclature">
<paragraph id="9546C77030D652E8EF3BFB5B73040428" blockId="140.[556,1032,1276,1339]" box="[556,1032,1276,1303]" pageId="140" pageNumber="141">
<heading id="CE0E701C30D652E8EF3BFB5B73040428" box="[556,1032,1276,1303]" centered="true" fontSize="11" level="2" pageId="140" pageNumber="141" reason="2">
<taxonomicName id="52F9BCF330D652E8EF3BFB5B73040428" ID-CoL="7ZQ3C" authority="Burmeister, 1874" authorityName="Burmeister" authorityYear="1874" box="[556,1032,1276,1303]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="aphodioides">
<emphasis id="A78D1B6230D652E8EF3BFB5B743A0428" bold="true" box="[556,822,1276,1303]" italics="true" pageId="140" pageNumber="141">Ontherus aphodioides</emphasis>
<bibRefCitation id="F168BA8130D652E8EE2AFB5A73040428" author="Burmeister H." box="[829,1032,1277,1303]" pageId="140" pageNumber="141" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister, 1874</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DDE394FB30D652E8EFF1FA8774430404" box="[742,847,1312,1339]" pageId="140" pageNumber="141" type="description">
<paragraph id="9546C77030D652E8EFF1FA8774430404" blockId="140.[556,1032,1276,1339]" box="[742,847,1312,1339]" pageId="140" pageNumber="141">
<figureCitation id="0DC2DBF530D652E8EFF1FA8774430404" box="[742,847,1312,1339]" captionStart="Fig" captionStartId="142.[188,231,1959,1985]" captionTargetBox="[271,1306,278,1899]" captionTargetId="figure-18@142.[259,1327,265,1917]" captionTargetPageId="142" captionText="Fig. 55. Species distribution. A. Ontherus aphodioides Burmeister, 1874. B. Ontherus appendiculatus (Mannerheim, 1829). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945617" httpUri="https://zenodo.org/record/13945617/files/figure.png" pageId="140" pageNumber="141">Fig. 55A</figureCitation>
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<subSubSection id="DDE394FB30D652E9EDAAFAC6756800C9" lastPageId="141" lastPageNumber="142" pageId="140" pageNumber="141" type="reference_group">
<paragraph id="9546C77030D652E8EDAAFAC6768C0737" blockId="140.[189,1399,1377,1686]" pageId="140" pageNumber="141">
<treatmentCitationGroup id="B5E9E05E30D652E8EDAAFAC6768C0737" pageId="140" pageNumber="141">
<treatmentCitation id="1458E16130D652E8EDAAFAC675C80444" author="Burmeister H." box="[189,708,1377,1404]" page="126" pageId="140" pageNumber="141" year="1874">
<taxonomicName id="52F9BCF330D652E8EDAAFAC675C80444" ID-CoL="7ZQ3C" authority="Burmeister, 1874: 126" authorityName="Burmeister" authorityPageNumber="126" authorityYear="1874" box="[189,708,1377,1404]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="aphodioides">
<emphasis id="A78D1B6230D652E8EDAAFAC676B20444" box="[189,446,1377,1403]" italics="true" pageId="140" pageNumber="141">Ontherus aphodioides</emphasis>
<bibRefCitation id="F168BA8130D652E8ECD4FAC675C80444" author="Burmeister H." box="[451,708,1377,1404]" pageId="140" pageNumber="141" pagination="120 - 133" refId="ref113171" refString="Burmeister H. 1874. Lamellicornia Argentina [Fortsetzung]. Entomologische Zeitung Stettin 35: 120 - 133. Available from https: // www. biodiversitylibrary. org / page / 9236010 [accessed 14 Feb. 2024]." type="journal article" year="1874">Burmeister, 1874: 126</bibRefCitation>
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(original description). Type locality:
<collectingCountry id="EDEE87E030D652E8E979FAC573D50443" box="[1134,1241,1378,1404]" name="Uruguay" pageId="140" pageNumber="141">Uruguay</collectingCountry>
:
<collectingRegion id="573D099230D652E8E9F6FAC6727F0444" box="[1249,1395,1377,1403]" country="Uruguay" name="Montevideo" pageId="140" pageNumber="141">Montevideo</collectingRegion>
. Name-bearing type:
<typeStatus id="4A4279D230D652E8ECF6FA22754704A0" box="[481,587,1413,1439]" pageId="140" pageNumber="141" type="syntype">syntypes</typeStatus>
, unknown whereabouts (
<treatmentCitation id="1458E16130D652E8EE6DFA22731A04A1" author="Genier F." box="[890,1046,1412,1439]" pageId="140" pageNumber="141" year="1996">
<bibRefCitation id="F168BA8130D652E8EE6DFA22731A04A1" author="Genier F." box="[890,1046,1412,1439]" pageId="140" pageNumber="141" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996</bibRefCitation>
</treatmentCitation>
). Described after Burmeister moved to
<collectingCountry id="EDEE87E030D652E8EC4AFA0F76DE04FD" box="[349,466,1448,1474]" name="Argentina" pageId="140" pageNumber="141">Argentina</collectingCountry>
in 1861 (
<treatmentCitation id="1458E16130D652E8EF57FA0F75DE04FD" author="Ulrich W." box="[576,722,1448,1474]" pageId="140" pageNumber="141" year="1972">
<bibRefCitation id="F168BA8130D652E8EF57FA0F75DE04FD" author="Ulrich W." box="[576,722,1448,1474]" pageId="140" pageNumber="141" pagination="1 - 20" refId="ref140112" refString="Ulrich W. 1972. Hermann Burmeister, 1807 to 1892. Annual Review of Entomology 17: 1 - 20. https: // doi. org / 10.1146 / annurev. en. 17.010172.000245" type="journal article" year="1972">Ulrich 1972</bibRefCitation>
</treatmentCitation>
), the type specimens would now be expected to be among the material of the Burmeister collection preserved in the MACN, but they were not found there by MC in 2014.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D652E8EDAAF9B6752F07A9" blockId="140.[189,1399,1377,1686]" pageId="140" pageNumber="141">
<treatmentCitationGroup id="B5E9E05E30D652E8EDAAF9B6752F07A9" pageId="140" pageNumber="141">
<treatmentCitation id="1458E16130D652E8EDAAF9B675B70713" author="Luederwaldt H." box="[189,699,1553,1580]" page="105" pageId="140" pageNumber="141" year="1930">
<taxonomicName id="52F9BCF330D652E8EDAAF9B675B70713" ID-CoL="7XM8B" authority="Luederwaldt, 1930: 105" authorityName="Luederwaldt" authorityPageNumber="105" authorityYear="1930" box="[189,699,1553,1580]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="A78D1B6230D652E8EDAAF9B676910714" box="[189,413,1553,1579]" italics="true" pageId="140" pageNumber="141">Ontherus convexus</emphasis>
<bibRefCitation id="F168BA8130D652E8ECB4F9B675B70713" author="Luederwaldt H." box="[419,699,1553,1580]" pageId="140" pageNumber="141" pagination="105 - 108" refId="ref128717" refString="Luederwaldt H. 1930. Especies novas de Ontherus. Archivos do Instituto Biologico 3: 105 - 108." type="journal article" year="1930">Luederwaldt, 1930: 105</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E030D652E8E967F9B673B00714" box="[1136,1212,1553,1579]" name="Brazil" pageId="140" pageNumber="141">Brazil</collectingCountry>
:
<collectingRegion id="573D099230D652E8E9D0F9B6727E0713" box="[1223,1394,1553,1580]" country="Brazil" name="Espirito Santo" pageId="140" pageNumber="141">Espírito Santo</collectingRegion>
: “Guandú” (possibly the river of this name crossing the state). Name-bearing type:
<typeStatus id="4A4279D230D652E8E9B0F99272020770" box="[1191,1294,1589,1615]" pageId="140" pageNumber="141" type="holotype">holotype</typeStatus>
(MNRJ) (
<treatmentCitation id="1458E16130D652E8EDE6F9FF7681074D" author="Genier F." box="[241,397,1624,1650]" pageId="140" pageNumber="141" year="1996">
<bibRefCitation id="F168BA8130D652E8EDE6F9FF7681074D" author="Genier F." box="[241,397,1624,1650]" pageId="140" pageNumber="141" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996</bibRefCitation>
</treatmentCitation>
), presumably destroyed in the 2018 fire that consumed the museums
<taxonomicName id="52F9BCF330D652E8E9E1F9FF727B074D" authorityName="Linnaeus" authorityYear="1758" box="[1270,1399,1624,1650]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
holdings (see
<treatmentCitation id="1458E16130D652E8EC9AF9DB751F07AA" author="Mega E. R." box="[397,531,1659,1686]" pageId="140" pageNumber="141" year="2019">
<bibRefCitation id="F168BA8130D652E8EC9AF9DB751F07AA" author="Mega E. R." box="[397,531,1659,1686]" pageId="140" pageNumber="141" pagination="312 - 315" refId="ref130951" refString="Mega E. R. 2019. The battle to rebuild centuries of science after an epic inferno. Nature 571: 312 - 315. https: // doi. org / 10.1038 / d 41586 - 019 - 02141 - 2" type="journal article" year="2019">Mega 2019</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D652E8EDAAF91B727B06B9" blockId="140.[189,1399,1724,1927]" pageId="140" pageNumber="141">
<treatmentCitationGroup id="B5E9E05E30D652E8EDAAF91B727B06B9" pageId="140" pageNumber="141">
<emphasis id="A78D1B6230D652E8EDAAF91B76E107E9" box="[189,493,1724,1750]" italics="true" pageId="140" pageNumber="141">
<taxonomicName id="52F9BCF330D652E8EDAAF91B76C707E9" ID-CoL="7ZQ3C" authorityName="Burmeister" authorityYear="1874" box="[189,459,1724,1750]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="140" pageNumber="141" phylum="Arthropoda" rank="species" species="aphodioides">Ontherus aphodioides</taxonomicName>
</emphasis>
<treatmentCitation id="1458E16130D652E8ECE9F91B75DE07E8" author="Gillet J. J. E." box="[510,722,1724,1751]" page="57" pageId="140" pageNumber="141" year="1911">
<bibRefCitation id="F168BA8130D652E8ECE9F91B75DE07E8" author="Gillet J. J. E." box="[510,722,1724,1751]" pageId="140" pageNumber="141" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 57</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E16130D652E8EEA6F91B73E507E8" author="Luederwaldt H." box="[945,1257,1724,1751]" page="371" pageId="140" pageNumber="141" year="1931">
<bibRefCitation id="F168BA8130D652E8EEA6F91B73E507E8" author="Luederwaldt H." box="[945,1257,1724,1751]" pageId="140" pageNumber="141" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 371</bibRefCitation>
</treatmentCitation>
, 384 (key, redescription). —
<treatmentCitation id="1458E16130D652E8ECDAF947741807C5" author="Pessoa S. B. &amp; Lane F." box="[461,788,1759,1786]" page="456" pageId="140" pageNumber="141" year="1941">
<bibRefCitation id="F168BA8130D652E8ECDAF947741807C5" author="Pessoa S. B. &amp; Lane F." box="[461,788,1759,1786]" pageId="140" pageNumber="141" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 456</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130D652E8EE88F94773B107C5" author="Blackwelder R. E." box="[927,1213,1760,1786]" page="206" pageId="140" pageNumber="141" year="1944">
<bibRefCitation id="F168BA8130D652E8EE88F94773B107C5" author="Blackwelder R. E." box="[927,1213,1760,1786]" pageId="140" pageNumber="141" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E030D652E8EDFDF8A4765A0622" box="[234,342,1795,1821]" name="Uruguay" pageId="140" pageNumber="141">Uruguay</collectingCountry>
). —
<treatmentCitation id="1458E16130D652E8EC9CF8A4755C0622" author="Lange R. B." box="[395,592,1795,1821]" page="310" pageId="140" pageNumber="141" year="1947">
<bibRefCitation id="F168BA8130D652E8EC9CF8A4755C0622" author="Lange R. B." box="[395,592,1795,1821]" pageId="140" pageNumber="141" pagination="305 - 315" refId="ref126831" refString="Lange R. B. 1947. Ensaio da zoogeografia dos Scarabaeidae do Parana com algumas notas etnoecologicas. Arquivos Museu Paranaense 6: 305 - 315." type="journal article" year="1947">Lange 1947: 310</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E16130D652E8EFA9F8A4749F0622" author="Martinez A." box="[702,915,1795,1822]" page="69" pageId="140" pageNumber="141" year="1959">
<bibRefCitation id="F168BA8130D652E8EFA9F8A4749F0622" author="Martinez A." box="[702,915,1795,1822]" pageId="140" pageNumber="141" pagination="1 - 126" refId="ref129799" refString="Martinez A. 1959. Catalogo de los Scarabaeidae Argentinos. Revista del Museo Argentino de Ciencias Naturales &quot; Bernardino Rivadavia &quot; 5: 1 - 126." type="journal article" year="1959">Martínez 1959: 69</bibRefCitation>
</treatmentCitation>
(catalogue for
<collectingCountry id="EDEE87E030D652E8E956F8A473B70622" box="[1089,1211,1795,1821]" name="Argentina" pageId="140" pageNumber="141">Argentina</collectingCountry>
). —
<treatmentCitation id="1458E16130D652E8E9E7F8A476A6067F" author="Martinez A. &amp; Pereira F. S." page="79" pageId="140" pageNumber="141" year="1960">
<bibRefCitation id="F168BA8130D652E8E9E7F8A476A6067F" author="Martinez A. &amp; Pereira F. S." pageId="140" pageNumber="141" pagination="77 - 84" refId="ref130211" refString="Martinez A. &amp; Pereira F. S. 1960. Algunos interesantes Coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina 22: 77 - 84." type="journal article" year="1960">Martínez &amp; Pereira 1960: 79</bibRefCitation>
</treatmentCitation>
, 80 (contribution). —
<treatmentCitation id="1458E16130D652E8EFBBF88174B3067F" author="Forsyth A. B. &amp; Spector S. &amp; Gill B. &amp; Guerra F. &amp; Ayzama S." box="[684,959,1830,1856]" page="370" pageId="140" pageNumber="141" year="1998">
<bibRefCitation id="F168BA8130D652E8EFBBF88174B3067F" author="Forsyth A. B. &amp; Spector S. &amp; Gill B. &amp; Guerra F. &amp; Ayzama S." box="[684,959,1830,1856]" pageId="140" pageNumber="141" refId="ref119575" refString="Forsyth A. B., Spector S., Gill B., Guerra F. &amp; Ayzama S. 1998. Dung beetles (Coleoptera: Scarabaeidae: Scarabaeinae) of Parque Nacinal Noel Kempff Mercado / Escarabajos (Coleoptera: Scarabaeidae: Scarabaeinae) del Parque Nacional Noel Kempff Mercado. In: Killeen T. J. &amp; Schulenberg T. S. (eds) Rapid Assessment Program. A Biological Assessment of Parque Nacional Noel Kempff Mercado, Bolivia: 181 - 200 + appendix 6. RAP working papers, Conservation International, Washington, D. C." type="journal volume" year="1998">
Forsyth
<emphasis id="A78D1B6230D652E8EE1BF8807449067F" box="[780,837,1830,1856]" italics="true" pageId="140" pageNumber="141">et al.</emphasis>
1998: 370
</bibRefCitation>
</treatmentCitation>
(conservation, list). —
<treatmentCitation id="1458E16130D652E8E9D8F8817619065C" author="Escobar F." page="208" pageId="140" pageNumber="141" year="2000">
<bibRefCitation id="F168BA8130D652E8E9D8F8817619065C" author="Escobar F." pageId="140" pageNumber="141" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 208</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030D652E8ECD3F8EE7531065C" box="[452,573,1865,1891]" name="Colombia" pageId="140" pageNumber="141">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130D652E8EF6FF8EE7496065C" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[632,922,1865,1891]" page="138" pageId="140" pageNumber="141" year="2001">
<bibRefCitation id="F168BA8130D652E8EF6FF8EE7496065C" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[632,922,1865,1891]" pageId="140" pageNumber="141" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B6230D652E8EFC9F8ED7417065C" box="[734,795,1865,1891]" italics="true" pageId="140" pageNumber="141">et al.</emphasis>
2001: 138
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030D652E8E95EF8EE73CE065C" box="[1097,1218,1865,1891]" name="Colombia" pageId="140" pageNumber="141">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E16130D652E8E9EAF8EE750406B9" author="Medina C. A. &amp; Pulido-Herrera L. A." page="60" pageId="140" pageNumber="141" year="2009">
<bibRefCitation id="F168BA8130D652E8E9EAF8EE750406B9" author="Medina C. A. &amp; Pulido-Herrera L. A." pageId="140" pageNumber="141" pagination="55 - 62" refId="ref130761" refString="Medina C. A. &amp; Pulido-Herrera L. A. 2009. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de la Orinoquia Colombiana. Biota Colombiana 10 (1 - 2): 55 - 62." type="journal article" year="2009">Medina &amp; Pulido-Herrera 2009: 60</bibRefCitation>
</treatmentCitation>
(diversity). —
<treatmentCitation id="1458E16130D652E8EFADF8CB732006B8" ID-DOI="http://doi.org/10.5281/zenodo.3684685" ID-Zenodo-Dep="3684685" author="Hielkema A. J. &amp; Hielkema M. A." box="[698,1068,1900,1927]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFF2FFD4FF48FEA134DDF985" page="56" pageId="140" pageNumber="141" year="2019">
<bibRefCitation id="F168BA8130D652E8EFADF8CB732006B8" author="Hielkema A. J. &amp; Hielkema M. A." box="[698,1068,1900,1927]" pageId="140" pageNumber="141" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 56</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFEAE76720055" blockId="141.[189,1399,265,502]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAFEAE76720055" pageId="141" pageNumber="142">
<emphasis id="A78D1B6230D752E9EDAAFEAE76BA001C" box="[189,438,265,291]" italics="true" pageId="141" pageNumber="142">
<taxonomicName id="52F9BCF330D752E9EDAAFEAE7693001C" ID-CoL="7XM8B" authorityName="Luederwaldt" authorityYear="1930" box="[189,415,265,291]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="convexus">Ontherus convexus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E16130D752E9ECA8FEAE7412001C" author="Luederwaldt H." box="[447,798,265,291]" page="371" pageId="141" pageNumber="142" year="1931">
<bibRefCitation id="F168BA8130D752E9ECA8FEAE7412001C" author="Luederwaldt H." box="[447,798,265,291]" pageId="141" pageNumber="142" pagination="363 - 422" refId="ref128737" refString="Luederwaldt H. 1931 a. O genero Ontherus (Coleop.) (Lamellic-Coprid. - Pinot.) com uma chave, para a determinacao dos pinotides americanos. Revista do Museu Paulista 17: 363 - 422 + 1 pl." type="journal article" year="1931">Luederwaldt 1931a: 371372</bibRefCitation>
</treatmentCitation>
, 386 (key, redescription). —
<treatmentCitation id="1458E16130D752E9E96EFEAE76140078" author="Pessoa S. B. &amp; Lane F." page="457" pageId="141" pageNumber="142" year="1941">
<bibRefCitation id="F168BA8130D752E9E96EFEAE76140078" author="Pessoa S. B. &amp; Lane F." pageId="141" pageNumber="142" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 457</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E16130D752E9EC8FFE8B75A30079" author="Blackwelder R. E." box="[408,687,300,326]" page="206" pageId="141" pageNumber="142" year="1944">
<bibRefCitation id="F168BA8130D752E9EC8FFE8B75A30079" author="Blackwelder R. E." box="[408,687,300,326]" pageId="141" pageNumber="142" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 206</bibRefCitation>
</treatmentCitation>
(list, cited for
<collectingCountry id="EDEE87E030D752E9EE71FE8B74BF0079" box="[870,947,300,326]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E16130D752E9EEF8FE8B727B0079" author="Martinez A. &amp; Pereira F. S." box="[1007,1399,300,326]" page="79" pageId="141" pageNumber="142" year="1960">
<bibRefCitation id="F168BA8130D752E9EEF8FE8B727B0079" author="Martinez A. &amp; Pereira F. S." box="[1007,1399,300,326]" pageId="141" pageNumber="142" pagination="77 - 84" refId="ref130211" refString="Martinez A. &amp; Pereira F. S. 1960. Algunos interesantes Coprini neotropicales (Col. Scarabaeidae). Revista de la Sociedad Entomologica Argentina 22: 77 - 84." type="journal article" year="1960">Martínez &amp; Pereira 1960: 7980</bibRefCitation>
</treatmentCitation>
(comments).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFED57661008F" blockId="141.[189,1399,265,502]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAFED57661008F" pageId="141" pageNumber="142">
<taxonomicName id="52F9BCF330D752E9EDAAFED5754400B3" ID-CoL="7ZQ3C" authorityName="Burmeister" authorityYear="1874" box="[189,584,370,397]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="aphodioides" subGenus="Ontherus">
<emphasis id="A78D1B6230D752E9EDAAFED5762600B3" box="[189,298,370,396]" italics="true" pageId="141" pageNumber="142">Ontherus</emphasis>
(
<emphasis id="A78D1B6230D752E9EC2CFED576A400B3" box="[315,424,370,396]" italics="true" pageId="141" pageNumber="142">Ontherus</emphasis>
)
<emphasis id="A78D1B6230D752E9ECAEFED5754400B3" box="[441,584,370,396]" italics="true" pageId="141" pageNumber="142">aphodioides</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130D752E9EF71FED5743E00B2" author="Genier F." box="[614,818,370,397]" page="117" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9EF71FED5743E00B2" author="Genier F." box="[614,818,370,397]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 117</bibRefCitation>
</treatmentCitation>
(revision). —
<treatmentCitation id="1458E16130D752E9EEC9FED573F100B2" author="Vaz-de-Mello F. Z." box="[990,1277,370,397]" page="194" pageId="141" pageNumber="142" year="2000">
<bibRefCitation id="F168BA8130D752E9EEC9FED573F100B2" author="Vaz-de-Mello F. Z." box="[990,1277,370,397]" pageId="141" pageNumber="142" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030D752E9EC03FE31766D008F" box="[276,353,406,432]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFE1E756800C9" blockId="141.[189,1399,265,502]" pageId="141" pageNumber="142">
<treatmentCitationGroup id="B5E9E05E30D752E9EDAAFE1E756800C9" pageId="141" pageNumber="142">
<taxonomicName id="52F9BCF330D752E9EDAAFE1E754200EC" ID-CoL="7ZQ3C" authorityName="Burmeister" authorityYear="1874" box="[189,590,441,467]" class="Insecta" family="Scarabaeidae" genus="Ontherus" kingdom="Animalia" order="Coleoptera" pageId="141" pageNumber="142" phylum="Arthropoda" rank="species" species="aphodioides" subGenus="Ontherus">
<emphasis id="A78D1B6230D752E9EDAAFE1E762600EC" box="[189,298,441,467]" italics="true" pageId="141" pageNumber="142">Ontherus</emphasis>
(
<emphasis id="A78D1B6230D752E9EC29FE1E76A700EC" box="[318,427,441,467]" italics="true" pageId="141" pageNumber="142">Ontherus</emphasis>
)
<emphasis id="A78D1B6230D752E9ECA8FE1E754200EC" box="[447,590,441,467]" italics="true" pageId="141" pageNumber="142">aphodioides</emphasis>
</taxonomicName>
<treatmentCitation id="1458E16130D752E9EF66FE1E744E00EC" author="Genier F." box="[625,834,441,467]" page="117" pageId="141" pageNumber="142" year="1996">
<bibRefCitation id="F168BA8130D752E9EF66FE1E744E00EC" author="Genier F." box="[625,834,441,467]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 117</bibRefCitation>
</treatmentCitation>
(revision, new combination). —
<treatmentCitation id="1458E16130D752E9E9C2FE1E766C00C9" author="Vaz-de-Mello F. Z." page="194" pageId="141" pageNumber="142" year="2000">
<bibRefCitation id="F168BA8130D752E9E9C2FE1E766C00C9" author="Vaz-de-Mello F. Z." pageId="141" pageNumber="142" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 194</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E030D752E9EF1CFE7B755400C9" box="[523,600,476,502]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30D752E9EDAAFDBA73770359" pageId="141" pageNumber="142" type="materials_examined">
<paragraph id="9546C77030D752E9EDAAFDBA76A60308" blockId="141.[189,426,541,567]" box="[189,426,541,567]" pageId="141" pageNumber="142">
<heading id="CE0E701C30D752E9EDAAFDBA76A60308" bold="true" box="[189,426,541,567]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="A78D1B6230D752E9EDAAFDBA76A60308" bold="true" box="[189,426,541,567]" pageId="141" pageNumber="142">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFDEB73770359" blockId="141.[189,1147,587,615]" box="[189,1147,587,615]" pageId="141" pageNumber="142">
<materialsCitation id="2591CD2D30D752E9EDAAFDEB737B0359" box="[189,1143,587,615]" collectingDate="1992-08-15" collectionCode="CEMT" collectorName="O. Hillert" country="Venezuela" location="Juruani" pageId="141" pageNumber="142" specimenCount="1" stateProvince="Bolivar">
<collectingCountry id="EDEE87E030D752E9EDAAFDEB76670359" box="[189,363,588,614]" name="Venezuela" pageId="141" pageNumber="142">VENEZUELA</collectingCountry>
<emphasis id="A78D1B6230D752E9EC90FDEC76EF0359" bold="true" box="[391,483,587,614]" pageId="141" pageNumber="142">
<collectingRegion id="573D099230D752E9EC90FDEC76EF0359" box="[391,483,587,614]" country="Venezuela" name="Bolivar" pageId="141" pageNumber="142">Bolívar</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF930D752E9ECEBFDEB755D0358" box="[508,593,588,615]" count="1" pageId="141" pageNumber="142" type="generic">1 spec.</specimenCount>
;
<location id="902691AB30D752E9EF4AFDEB75BB0359" LSID="urn:lsid:plazi:treatment:1D50766630D652E9EF3BFB5B75800289:902691AB30D752E9EF4AFDEB75BB0359" box="[605,695,588,614]" country="Venezuela" name="Juruani" pageId="141" pageNumber="142" stateProvince="Bolivar">Juruani</location>
;
<date id="E147E1B030D752E9EFD4FDEB74690359" box="[707,869,588,615]" pageId="141" pageNumber="142" value="1992-08-15">
<collectingDate id="F103185830D752E9EFD4FDEB74690359" box="[707,869,588,615]" pageId="141" pageNumber="142" value="1992-08-15">15 Aug. 1992</collectingDate>
</date>
;
<collectorName id="380CA2A630D752E9EE66FDEB74EE0359" box="[881,994,588,614]" pageId="141" pageNumber="142">O. Hillert</collectorName>
leg.;
<collectionCode id="F3E85FB530D752E9E933FDEB737B0359" box="[1060,1143,588,614]" pageId="141" pageNumber="142">CEMT</collectionCode>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30D752E9EDAAFD2A74960279" pageId="141" pageNumber="142" type="distribution">
<paragraph id="9546C77030D752E9EDAAFD2A765A0398" blockId="141.[189,342,653,679]" box="[189,342,653,679]" pageId="141" pageNumber="142">
<heading id="CE0E701C30D752E9EDAAFD2A765A0398" bold="true" box="[189,342,653,679]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="A78D1B6230D752E9EDAAFD2A765A0398" bold="true" box="[189,342,653,679]" pageId="141" pageNumber="142">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFD1B74E003E9" blockId="141.[189,1004,700,727]" box="[189,1004,700,727]" pageId="141" pageNumber="142">
<collectingCountry id="EDEE87E030D752E9EDAAFD1B763803E9" box="[189,308,700,726]" name="Colombia" pageId="141" pageNumber="142">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E030D752E9EC28FD1B76B703E9" box="[319,443,700,726]" name="Venezuela" pageId="141" pageNumber="142">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E030D752E9ECD1FD1B751D03E9" box="[454,529,700,726]" name="Brazil" pageId="141" pageNumber="142">Brazil</collectingCountry>
,
<collectingCountry id="EDEE87E030D752E9EF0BFD1A755903E8" box="[540,597,701,727]" name="Peru" pageId="141" pageNumber="142">Peru</collectingCountry>
,
<collectingCountry id="EDEE87E030D752E9EF76FD1B75B603E9" box="[609,698,700,726]" name="Bolivia" pageId="141" pageNumber="142">Bolivia</collectingCountry>
,
<collectingCountry id="EDEE87E030D752E9EFD1FD1A743803E8" box="[710,820,701,727]" name="Paraguay" pageId="141" pageNumber="142">Paraguay</collectingCountry>
, and
<collectingCountry id="EDEE87E030D752E9EE67FD1B74E503E9" box="[880,1001,700,726]" name="Argentina" pageId="141" pageNumber="142">Argentina</collectingCountry>
.
</paragraph>
<paragraph id="9546C77030D752E9EDAAFD5A76E10227" blockId="141.[189,493,765,792]" box="[189,493,765,792]" pageId="141" pageNumber="142">
<heading id="CE0E701C30D752E9EDAAFD5A76E10227" bold="true" box="[189,493,765,792]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="A78D1B6230D752E9EDAAFD5A76E10227" bold="true" box="[189,493,765,792]" pageId="141" pageNumber="142">
Subregions of
<collectingCountry id="EDEE87E030D752E9EC78FD5976E10227" box="[367,493,766,792]" name="Venezuela" pageId="141" pageNumber="142">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFC8B74960279" blockId="141.[189,922,812,839]" box="[189,922,812,839]" pageId="141" pageNumber="142">Peneplain of the Caura and Paragua rivers, and Guiana Shield.</paragraph>
</subSubSection>
<subSubSection id="DDE394FB30D752E9EDAAFCCA75800289" pageId="141" pageNumber="142" type="reference_group">
<paragraph id="9546C77030D752E9EDAAFCCA769502B7" blockId="141.[189,409,877,904]" box="[189,409,877,904]" pageId="141" pageNumber="142">
<heading id="CE0E701C30D752E9EDAAFCCA769502B7" bold="true" box="[189,409,877,904]" fontSize="11" level="3" pageId="141" pageNumber="142" reason="3">
<emphasis id="A78D1B6230D752E9EDAAFCCA769502B7" bold="true" box="[189,409,877,904]" pageId="141" pageNumber="142">Literature record</emphasis>
</heading>
</paragraph>
<paragraph id="9546C77030D752E9EDAAFC3B75800289" blockId="141.[189,652,924,950]" box="[189,652,924,950]" pageId="141" pageNumber="142">
<bibRefCitation id="F168BA8130D752E9EDAAFC3B76850289" author="Genier F." box="[189,393,924,950]" pageId="141" pageNumber="142" pagination="1 - 169" refId="ref120431" refString="Genier F. 1996. A revision of the Neotropical genus Ontherus Erichson (Coleoptera: Scarabaeidae, Scarabaeinae). Memoirs of the Entomological Society of Canada 170: 1 - 169. https: // doi. org / 10.4039 / entm 128170 fv" type="journal article" year="1996">Génier 1996: 117</bibRefCitation>
(
<collectingCountry id="EDEE87E030D752E9EC8FFC3B751B0289" box="[408,535,924,950]" name="Venezuela" pageId="141" pageNumber="142">Venezuela</collectingCountry>
:
<collectingRegion id="573D099230D752E9EF35FC3B75730289" box="[546,639,924,950]" country="Venezuela" name="Bolivar" pageId="141" pageNumber="142">Bolívar</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>