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<document id="74F74BC97731D1E0D3FB25508E0DA56D" ID-CLB-Dataset="305167" ID-DOI="10.5852/ejt.2024.959.2677" ID-GBIF-Dataset="bf132c8e-dbc7-4f3d-96e9-1a0b153e765a" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666309452ABEF48FA99726B00C9" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Uroxys Westwood 1842" docType="treatment" docVersion="9" lastPageNumber="208" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="207" updateTime="1731534417997" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="68FB50B556483AFB22167E24788B3DC9">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="05EF23DA5B65FC851C24AABF35FBA957">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="AA0C7F96D6434652561E8C8F54EA90D8">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="4CD623A5F189D8DDC8159AF9AF1CDD4B">Cupello, Mario</mods:namePart>
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<mods:affiliation id="AE2E9149ADA9C03914E6AC1C20F7792A">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="81C294842205C59820160675CA0C23EE" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:affiliation id="7E3FA6C03570ED278D53828D6368B916">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<mods:date id="2177D6C67141A54316C6AC2481F7CEA3">2024</mods:date>
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Genus
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<emphasis id="A78D1B62309452AAEFA5FA9974040467" bold="true" box="[690,776,1342,1368]" italics="true" pageId="206" pageNumber="207">Uroxys</emphasis>
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<emphasis id="A78D1B62309452AAEDAAFA27761D04A6" box="[189,273,1408,1433]" italics="true" pageId="206" pageNumber="207">Uroxys</emphasis>
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(original description).
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species:
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<taxonomicName id="52F9BCF3309452AAEED1FA27731604A6" ID-CoL="85CV" authorityName="Westwood" authorityYear="1842" box="[966,1050,1408,1433]" class="Insecta" family="Scarabaeidae" genus="Uroxys" kingdom="Animalia" order="Coleoptera" pageId="206" pageNumber="207" phylum="Arthropoda" rank="genus">Uroxys</taxonomicName>
cuprescens
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<bibRefCitation id="F168BA81309452AAE9BDFAD8727D04A6" author="Westwood J. O." box="[1194,1393,1407,1433]" pageId="206" pageNumber="207" pagination="58 - 59" refId="ref141897" refString="Westwood J. O. 1842. Description of new exotic genera belonging to the family of the sacred beetles. Proceedings of the Entomological Society of London 1842: 58 - 59." type="book chapter" year="1842">Westwood, 1842</bibRefCitation>
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, by original monotypy.
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<treatmentCitationGroup id="B5E9E05E309452AAEDAAFA627534073C" pageId="206" pageNumber="207">
<treatmentCitation id="1458E161309452AAEDAAFA62755E04E0" author="Balthasar V." box="[189,594,1477,1503]" page="210" pageId="206" pageNumber="207" year="1938">
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<emphasis id="A78D1B62309452AAEDAAFA62765E04E0" box="[189,338,1477,1503]" italics="true" pageId="206" pageNumber="207">Pseuduroxys</emphasis>
<bibRefCitation id="F168BA81309452AAEC4CFA62755E04E0" author="Balthasar V." box="[347,594,1477,1503]" pageId="206" pageNumber="207" pagination="210 - 223" refId="ref110121" refString="Balthasar V. 1938. Neue Gattungen und Arten der sudamerikanischen Coprophagen. (4. Beitrag zur Kenntnis der Scarabaeiden der neotropischen Region). Entomologische Blatter 34 (4): 210 - 223." type="journal article" year="1938">Balthasar, 1938: 210</bibRefCitation>
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(original description).
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species:
<treatmentCitation id="1458E161309452AAE903FA62762B073C" author="Balthasar V." pageId="206" pageNumber="207" year="1938">
<taxonomicName id="52F9BCF3309452AAE903FA62762B073C" ID-CoL="4PFHP" authority="Balthasar 1938" authorityName="Balthasar" authorityYear="1938" class="Insecta" family="Scarabaeidae" genus="Pseuduroxys" kingdom="Animalia" order="Coleoptera" pageId="206" pageNumber="207" phylum="Arthropoda" rank="species" species="ohausi">
<emphasis id="A78D1B62309452AAE903FA62720C04E0" box="[1044,1280,1477,1503]" italics="true" pageId="206" pageNumber="207">Pseuduroxys ohausi</emphasis>
<bibRefCitation id="F168BA81309452AAE81DFA62762B073C" author="Balthasar V." pageId="206" pageNumber="207" pagination="210 - 223" refId="ref110121" refString="Balthasar V. 1938. Neue Gattungen und Arten der sudamerikanischen Coprophagen. (4. Beitrag zur Kenntnis der Scarabaeiden der neotropischen Region). Entomologische Blatter 34 (4): 210 - 223." type="journal article" year="1938">Balthasar 1938</bibRefCitation>
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, by original monotypy.
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</paragraph>
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<emphasis id="A78D1B62309452AAEDAAF98C761D077B" box="[189,273,1579,1604]" italics="true" pageId="206" pageNumber="207">Uroxys</emphasis>
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<treatmentCitation id="1458E161309452AAEC2BF98D7539077B" author="Westwood J. O." box="[316,565,1578,1604]" page="61" pageId="206" pageNumber="207" year="1843">
<bibRefCitation id="F168BA81309452AAEC2BF98D7539077B" author="Westwood J. O." box="[316,565,1578,1604]" pageId="206" pageNumber="207" pagination="61 - 62" refId="ref141931" refString="Westwood J. O. 1843. Description of some new exotic genera belonging to the family of the sacred beetles. The Annals and Magazine of Natural History, Zoology, Botany, and Geology 11: 61 - 62. Available from https: // www. biodiversitylibrary. org / page / 2326216 [accessed 14 Feb. 2024]." type="journal article" year="1843">Westwood 1843: 61</bibRefCitation>
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(redescription);
<treatmentCitation id="1458E161309452AAEE1FF98D748A077B" author="Westwood J. O." box="[776,902,1578,1604]" page="229" pageId="206" pageNumber="207" year="1847">
<bibRefCitation id="F168BA81309452AAEE1FF98D748A077B" author="Westwood J. O." box="[776,902,1578,1604]" pageId="206" pageNumber="207" pagination="225 - 232" refId="ref141995" refString="Westwood J. O. 1847. Characteres of various new groups and species amongst the coprophagous lamellicorni beetles. Transactions of Entomological Society of London 4 (4): 225 - 232. https: // doi. org / 10.1111 / j. 1365 - 2311.1847. tb 01357. x" type="journal article" year="1847">1847: 229</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E161309452AAE995F98D727A077B" author="Agassiz L." box="[1154,1398,1578,1604]" page="1111" pageId="206" pageNumber="207" year="1846">
<bibRefCitation id="F168BA81309452AAE995F98D727A077B" author="Agassiz L." box="[1154,1398,1578,1604]" pageId="206" pageNumber="207" refId="ref108975" refString="Agassiz L. 1846. Nomenclatoris zoologici index universalis, continens nomina systematica classium, ordinum, familiarum et generum animalium omnium, tam viventium quam fossilum, secundum ordinem alphabeticum unicum deposita, adjectis homonymus platarum, nec non variis adnotationibus et emendationibus. Jent and Gassman, Solothurn [Soloduri]. https: // doi. org / 10.5962 / bhl. title. 1819" type="book" year="1846">Agassiz 1846: 1111</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161309452AAECB3F9EA75990758" author="Lacordaire J. T." box="[420,661,1613,1640]" page="91" pageId="206" pageNumber="207" year="1855">
<bibRefCitation id="F168BA81309452AAECB3F9EA75990758" author="Lacordaire J. T." box="[420,661,1613,1640]" pageId="206" pageNumber="207" refId="ref126653" refString="Lacordaire J. T. 1855. Histoire naturelle des Insectes. Genera des coleopteres, ou expose methodique et critique de tous les genres proposes jusqu'ici dans cet ordre d'insectes. Tome troisieme. Contenant les familles des Pectinicornes et Lamellicornes. Librarie Encyclopedique de Roret, Paris. [Publication date following Bousquet (2016).] https: // doi. org / 10.5962 / bhl. title. 8864" type="journal volume" year="1855">Lacordaire 1855: 91</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E161309452AAEE96F9EA73580758" author="Harold E." box="[897,1108,1613,1639]" page="37" pageId="206" pageNumber="207" year="1868">
<bibRefCitation id="F168BA81309452AAEE96F9EA73580758" author="Harold E." box="[897,1108,1613,1639]" pageId="206" pageNumber="207" pagination="33 - 55" refId="ref123386" refString="Harold E. 1868 c. Die Choerididen-Gattungen Uroxys und Trichillum. Coleopterologische Hefte 3: 33 - 55. Available from https: // www. biodiversitylibrary. org / page / 16069500 [accessed 16 Feb. 2024]." type="journal article" year="1868">Harold 1868c: 37</bibRefCitation>
</treatmentCitation>
, 54 (redescription, key);
<treatmentCitation id="1458E161309452AAEDFBF9D7767107B4" author="Harold E." box="[236,381,1648,1675]" page="1001" pageId="206" pageNumber="207" year="1869">
<bibRefCitation id="F168BA81309452AAEDFBF9D7767107B4" author="Harold E." box="[236,381,1648,1675]" firstAuthor="d" pageId="206" pageNumber="207" refId="ref123629" refString="Harold E. 1869 d. Catalogus coleopterorum. Hucusque descriptorum, synonymicus et systematicus. Tom. IV. Scarabaeidae. E. H. Gummi, Munich. https: // doi. org / 10.5962 / bhl. title. 9089" type="book" year="1869">1869d: 1001</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161309452AAEF2EF9D675EB07B4" author="Bates H. W." box="[569,743,1649,1675]" page="43" pageId="206" pageNumber="207" year="1887">
<bibRefCitation id="F168BA81309452AAEF2EF9D675EB07B4" author="Bates H. W." box="[569,743,1649,1675]" pageId="206" pageNumber="207" refId="ref110759" refString="Bates H. W. 1887. Tribe Lamellicornia. In: Bates H. W. (1886 - 1889) Biologia Centrali-Americana. Insecta. Coleoptera. Vol. II. Part 2. Pectinicornia and Lamellicornia: 25 - 64. Taylor and Francis, London. https: // doi. org / 10.5962 / bhl. title. 730 [Publication date according to Lyal (2011) and Bousquet (2016).]" type="book" year="1887">Bates 1887: 43</bibRefCitation>
</treatmentCitation>
(distribution).
<emphasis id="A78D1B62309452AAEE82F9D674A307B5" box="[917,943,1649,1674]" italics="true" pageId="206" pageNumber="207"></emphasis>
<treatmentCitation id="1458E161309452AAEEA0F9D7737807B4" author="Gillet J. J. E." box="[951,1140,1648,1675]" page="49" pageId="206" pageNumber="207" year="1911">
<bibRefCitation id="F168BA81309452AAEEA0F9D7737807B4" author="Gillet J. J. E." box="[951,1140,1648,1675]" pageId="206" pageNumber="207" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 49</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161309452AAE827F9D676520792" author="Lucas R." page="666" pageId="206" pageNumber="207" year="1920">
<bibRefCitation id="F168BA81309452AAE827F9D676520792" author="Lucas R." pageId="206" pageNumber="207" refId="ref128561" refString="Lucas R. 1920. Catalogus alphabeticus generum et subgenerum Coleopterorum orbis terrarum totius (famil., trib., subtr., sect. incl.). Pars I. Nicolaische, Berlin." type="book" year="1920">Lucas 1920: 666</bibRefCitation>
</treatmentCitation>
(catalog, distribution). —
<treatmentCitation id="1458E161309452AAEF98F933745A0791" author="Arrow G. J." box="[655,854,1684,1710]" page="387" pageId="206" pageNumber="207" year="1933">
<bibRefCitation id="F168BA81309452AAEF98F933745A0791" author="Arrow G. J." box="[655,854,1684,1710]" pageId="206" pageNumber="207" pagination="385 - 399" refId="ref109993" refString="Arrow G. J. 1933. The genus Uroxys (Coleoptera, Copridӕ), with descriptions of some new species. Annals and Magazine of Natural History, Series 10 11 (63): 385 - 399. https: // doi. org / 10.1080 / 00222933308673669" type="journal article" year="1933">Arrow 1933: 387</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161309452AAEED4F93373990791" author="Paulian R." box="[963,1173,1684,1710]" page="233" pageId="206" pageNumber="207" year="1938">
<bibRefCitation id="F168BA81309452AAEED4F93373990791" author="Paulian R." box="[963,1173,1684,1710]" pageId="206" pageNumber="207" pagination="213 - 296" refId="ref134801" refString="Paulian R. 1938. Contribution a l'etude des Canthonides americans [Coleopt. Lamellic.]. Annales de la Societe entomologique de France 107: 213 - 296." type="journal article" year="1938">Paulian 1938: 233</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309452AAE81EF933769307EE" author="Pessoa S. B. &amp; Lane F." page="441" pageId="206" pageNumber="207" year="1941">
<bibRefCitation id="F168BA81309452AAE81EF933769307EE" author="Pessoa S. B. &amp; Lane F." pageId="206" pageNumber="207" pagination="389 - 504" refId="ref135859" refString="Pessoa S. B. &amp; Lane F. 1941. Coleopteros necrofagos de interesse medico-legal: ensaio monografico sobre a familia Scarabaeidae de S. Paulo e regioes vizinhas. Arquivos de Zoologia do Estado de Sao Paulo 2: 389 - 504. https: // doi. org / 10.11606 / issn. 2176 - 7793.19412389 - 504" type="journal article" year="1941">Pessôa &amp; Lane 1941: 441</bibRefCitation>
</treatmentCitation>
(diagnosis, misspelled as
<emphasis id="A78D1B62309452AAEFD9F91F741107EE" box="[718,797,1720,1745]" italics="true" pageId="206" pageNumber="207">Uroxis</emphasis>
). —
<treatmentCitation id="1458E161309452AAEE4EF910736407EE" author="Blackwelder R. E." box="[857,1128,1719,1745]" page="203" pageId="206" pageNumber="207" year="1944">
<bibRefCitation id="F168BA81309452AAEE4EF910736407EE" author="Blackwelder R. E." box="[857,1128,1719,1745]" pageId="206" pageNumber="207" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 203</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161309452AAE834F910766C07CA" author="Pereira F. S." page="56" pageId="206" pageNumber="207" year="1954">
<bibRefCitation id="F168BA81309452AAE834F910766C07CA" author="Pereira F. S." pageId="206" pageNumber="207" pagination="53 - 78" refId="ref135304" refString="Pereira F. S. 1954 a. O genero Scatonomus Er. (Coleoptera, Scarabaeidae). Revista Brasileira de Entomologia 1: 53 - 78." type="journal article" year="1954">Pereira 1954a: 56</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309452AAECF7F97C759C07CB" author="Roze J. A." box="[480,656,1754,1781]" page="43" pageId="206" pageNumber="207" year="1955">
<bibRefCitation id="F168BA81309452AAECF7F97C759C07CB" author="Roze J. A." box="[480,656,1754,1781]" pageId="206" pageNumber="207" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 43</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309452AAEE57F97D74B207CB" box="[832,958,1754,1780]" name="Venezuela" pageId="206" pageNumber="207">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E161309452AAEEEDF97D727A07CA" author="Halffter G. &amp; Matthews E. G." box="[1018,1398,1754,1781]" page="256" pageId="206" pageNumber="207" year="1966">
<bibRefCitation id="F168BA81309452AAEEEDF97D727A07CA" author="Halffter G. &amp; Matthews E. G." box="[1018,1398,1754,1781]" pageId="206" pageNumber="207" pagination="1 - 312" refId="ref122681" refString="Halffter G. &amp; Matthews E. G. 1966. The natural history of dung beetles of the subfamily Scarabaeinae (Coleoptera: Scarabaeidae). Folia Entomologica Mexicana 12 - 14: 1 - 312." type="book chapter" year="1966">Halffter &amp; Matthews 1966: 256</bibRefCitation>
</treatmentCitation>
(catalog, distribution). —
<treatmentCitation id="1458E161309452AAEF0DF95A74630627" author="Vulcano M. A. &amp; Pereira F. S." box="[538,879,1789,1816]" page="576" pageId="206" pageNumber="207" year="1967">
<bibRefCitation id="F168BA81309452AAEF0DF95A74630627" author="Vulcano M. A. &amp; Pereira F. S." box="[538,879,1789,1816]" pageId="206" pageNumber="207" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 576</bibRefCitation>
</treatmentCitation>
(key for the Amazon). —
<treatmentCitation id="1458E161309452AAE9B4F95A765B0604" author="Howden H. F. &amp; Young O. P." page="13" pageId="206" pageNumber="207" year="1981">
<bibRefCitation id="F168BA81309452AAE9B4F95A765B0604" author="Howden H. F. &amp; Young O. P." pageId="206" pageNumber="207" pagination="1 - 204" refId="ref124824" refString="Howden H. F. &amp; Young O. P. 1981. Panamanian Scarabaeinae: taxonomy, distribution and habits (Coleoptera, Scarabaeidae). Contributions of the American Entomological Institute 18 (1): 1 - 204." type="journal article" year="1981">Howden &amp; Young 1981: 13</bibRefCitation>
</treatmentCitation>
, 50 (key, redescription). —
<treatmentCitation id="1458E161309452AAEFA0F887733B0604" author="Halffter G. &amp; Edmonds W. D." box="[695,1079,1824,1851]" page="137" pageId="206" pageNumber="207" year="1982">
<bibRefCitation id="F168BA81309452AAEFA0F887733B0604" author="Halffter G. &amp; Edmonds W. D." box="[695,1079,1824,1851]" pageId="206" pageNumber="207" refId="ref122441" refString="Halffter G. &amp; Edmonds W. D. 1982. The Nesting Behavior of Dung Beetles (Scarabaeinae). An Ecological and Evolutive Approach. Instituto de Ecologia, Mexico, D. F." type="book" year="1982">Halffter &amp; Edmonds 1982: 137</bibRefCitation>
</treatmentCitation>
(catalog, distribution). —
<treatmentCitation id="1458E161309452AAEDFDF8E375820662" author="Medina C. A. &amp; Lopera-Toro A." box="[234,654,1859,1886]" page="306" pageId="206" pageNumber="207" year="2000">
<bibRefCitation id="F168BA81309452AAEDFDF8E375820662" author="Medina C. A. &amp; Lopera-Toro A." box="[234,654,1859,1886]" pageId="206" pageNumber="207" pagination="299 - 315" refId="ref130725" refString="Medina C. A. &amp; Lopera-Toro A. 2000. Clave ilustrada para la identificacion de generos de escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Caldasia 22: 299 - 315." type="journal article" year="2000">Medina &amp; Lopera-Toro 2000: 306</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309452AAEE03F8E373300661" author="Vaz-de-Mello F. Z." box="[788,1084,1859,1886]" page="195" pageId="206" pageNumber="207" year="2000">
<bibRefCitation id="F168BA81309452AAEE03F8E373300661" author="Vaz-de-Mello F. Z." box="[788,1084,1859,1886]" pageId="206" pageNumber="207" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 195</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309452AAE9E3F8E3724D0661" box="[1268,1345,1860,1886]" name="Brazil" pageId="206" pageNumber="207">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E161309452AAEDFDF8C0750906BE" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[234,517,1895,1921]" page="139" pageId="206" pageNumber="207" year="2001">
<bibRefCitation id="F168BA81309452AAEDFDF8C0750906BE" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[234,517,1895,1921]" pageId="206" pageNumber="207" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B62309452AAEC5BF8CF768B06BE" box="[332,391,1895,1921]" italics="true" pageId="206" pageNumber="207">et al.</emphasis>
2001: 139
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309452AAEFA6F8C0742606BE" box="[689,810,1895,1921]" name="Colombia" pageId="206" pageNumber="207">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161309452AAEE75F8C0733406BD" author="Ratcliffe B. C." box="[866,1080,1895,1922]" page="15" pageId="206" pageNumber="207" year="2002">
<bibRefCitation id="F168BA81309452AAEE75F8C0733406BD" author="Ratcliffe B. C." box="[866,1080,1895,1922]" firstAuthor="Ratcliffe" pageId="206" pageNumber="207" pagination="1 - 48" refId="ref136799" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea of Panama. Zootaxa 32: 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 15</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309452AAE9F4F8C0724906BE" box="[1251,1349,1895,1921]" name="Panama" pageId="206" pageNumber="207">Panama</collectingCountry>
). —
<treatmentCitation id="1458E161309452AAEDFCF82D76DF069A" author="Kohlmann B." box="[235,467,1930,1957]" page="56" pageId="206" pageNumber="207" year="2003">
<bibRefCitation id="F168BA81309452AAEDFCF82D76DF069A" author="Kohlmann B." box="[235,467,1930,1957]" firstAuthor="Kohlmann" pageId="206" pageNumber="207" pagination="45 - 58" refId="ref126021" refString="Kohlmann B. 2003. Tribu Coprini. In: Moron M. A. (ed.) Atlas de los Escarabajos de Mexico. Coleoptera: Lamellicornia. Vol. II. Familias Scarabaeidae, Trogidae, Passalidae y Lucanidae: 45 - 58. Argania Editio, Barcelona." type="book chapter" year="2003">Kohlmann 2003: 56</bibRefCitation>
</treatmentCitation>
(diagnosis). —
<treatmentCitation id="1458E161309452AAEF9BF82D74EE069B" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[652,994,1930,1956]" page="12" pageId="206" pageNumber="207" year="2006">
<bibRefCitation id="F168BA81309452AAEF9BF82D74EE069B" author="Hamel-Leigue A. &amp; Mann D. J. &amp; Vaz-de-Mello F. Z. &amp; Herzog S. K." box="[652,994,1930,1956]" pageId="206" pageNumber="207" pagination="1 - 18" refId="ref122830" refString="Hamel-Leigue A., Mann D. J., Vaz-de-Mello F. Z. &amp; Herzog S. K. 2006. Hacia un inventario de los escarabajos peloteros (Coleoptera: Scarabaeinae) de Bolivia: primera compilacion de los generos y especies registrados para el pais. Revista Boliviana de Ecologia y Conservacion Ambiental 20: 1 - 18." type="journal article" year="2006">
Hamel-Leigue
<emphasis id="A78D1B62309452AAEE2CF82C747A069B" box="[827,886,1930,1956]" italics="true" pageId="206" pageNumber="207">et al.</emphasis>
2006: 12
</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E0309452AAE953F82D73AC069B" box="[1092,1184,1930,1956]" name="Bolivia" pageId="206" pageNumber="207">Bolívia</collectingCountry>
). —
<treatmentCitation id="1458E161309452AAE9C2F82D769606F8" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." page="22" pageId="206" pageNumber="207" year="2011">
<bibRefCitation id="F168BA81309452AAE9C2F82D769606F8" author="Vaz-de-Mello F. Z. &amp; Edmonds W. D. &amp; Ocampo F. C. &amp; Schoolmeesters P." pageId="206" pageNumber="207" pagination="1 - 73" refId="ref140802" refString="Vaz-de-Mello F. Z., Edmonds W. D., Ocampo F. C., Schoolmeesters P. 2011 a. A multilingual key to the genera and subgenera of the subfamily Scarabaeinae of the New World (Coleoptera: Scarabaeidae). Zootaxa 2854 (1): 1 - 73. https: // doi. org / 10.11646 / zootaxa. 2854.1.1" type="journal article" year="2011">
Vaz-de-Mello
<emphasis id="A78D1B62309452AAEDFEF809762F06F8" box="[233,291,1965,1991]" italics="true" pageId="206" pageNumber="207">et al.</emphasis>
2011a: 22
</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309452AAEF1AF80A742606F8" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[525,810,1965,1991]" page="132" pageId="206" pageNumber="207" year="2011">
<bibRefCitation id="F168BA81309452AAEF1AF80A742606F8" author="Carvajal V. &amp; Villamarin S. &amp; Ortega A. M." box="[525,810,1965,1991]" pageId="206" pageNumber="207" refId="ref113831" refString="Carvajal V., Villamarin S. &amp; Ortega A. M. 2011. Escarabajos del Ecuador. Principales Generos. Instituto de Ciencias Biologicas, Escuela Politecnica Nacional, Quito." type="book" year="2011">
Carvajal
<emphasis id="A78D1B62309452AAEF63F80975A206F8" box="[628,686,1965,1991]" italics="true" pageId="206" pageNumber="207">et al.</emphasis>
2011: 132
</bibRefCitation>
</treatmentCitation>
, 318 (diagnosis, list for
<collectingCountry id="EDEE87E0309452AAE92CF80A73AE06F8" box="[1083,1186,1965,1991]" name="Ecuador" pageId="206" pageNumber="207">Ecuador</collectingCountry>
). —
<treatmentCitation id="1458E161309452ABE9C1F80A761A001C" author="Krajcik M." lastPageId="207" lastPageNumber="208" page="262" pageId="206" pageNumber="207" year="2012">
<bibRefCitation id="F168BA81309452ABE9C1F80A761A001C" author="Krajcik M." lastPageId="207" lastPageNumber="208" pageId="206" pageNumber="207" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 262</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161309552ABEC9AFEAE75DC001B" author="Solis A. &amp; Kohlmann B." box="[397,720,265,292]" page="5" pageId="207" pageNumber="208" year="2012">
<bibRefCitation id="F168BA81309552ABEC9AFEAE75DC001B" author="Solis A. &amp; Kohlmann B." box="[397,720,265,292]" pageId="207" pageNumber="208" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 5</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0309552ABEE68FEAE730A001C" box="[895,1030,265,291]" name="Costa Rica" pageId="207" pageNumber="208">Costa Rica</collectingCountry>
);
<treatmentCitation id="1458E161309552ABE90CFEAE739F001C" author="Solis A. &amp; Kohlmann B." box="[1051,1171,265,291]" page="290" pageId="207" pageNumber="208" year="2013">
<bibRefCitation id="F168BA81309552ABE90CFEAE739F001C" author="Solis A. &amp; Kohlmann B." box="[1051,1171,265,291]" pageId="207" pageNumber="208" pagination="289 - 340" refId="ref139240" refString="Solis A. &amp; Kohlmann B. 2013. El genero Uroxys en Costa Rica. Giornale Italiano di Entomologia 13 (58): 289 - 340." type="journal article" year="2013">2013: 290</bibRefCitation>
</treatmentCitation>
(redescription). —
<treatmentCitation id="1458E161309552ABEDFBFE8B75710079" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[236,637,300,326]" page="106" pageId="207" pageNumber="208" year="2013">
<bibRefCitation id="F168BA81309552ABEDFBFE8B75710079" author="Boilly O. &amp; Vaz-de-Mello F. Z." box="[236,637,300,326]" pageId="207" pageNumber="208" pagination="103 - 112" refId="ref111991" refString="Boilly O. &amp; Vaz-de-Mello F. Z. 2013. Les Scarabaeinae de Guyane: cle illustree des genres. ACOREP- France 7: 103 - 112." type="journal article" year="2013">Boilly &amp; Vaz-de-Mello 2013: 106</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161309552ABEFEFFE8B73290079" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[760,1061,300,326]" page="74" pageId="207" pageNumber="208" year="2018">
<bibRefCitation id="F168BA81309552ABEFEFFE8B73290079" author="Chamorro W. &amp; Marin-Armijos D. &amp; Granda V. &amp; Vaz-de-Mello F. Z." box="[760,1061,300,326]" pageId="207" pageNumber="208" pagination="72 - 100" refId="ref114035" refString="Chamorro W., Marin-Armijos D., Granda V. &amp; Vaz-de-Mello F. Z. 2018. Listado de especies y clave de generos y subgeneros de escarabajos estercoleros (Coleoptera: Scarabaeidae: Scarabaeinae) presentes y presuntos para Ecuador. Revista Colombiana de Entomologia 44 (1): 72 - 100. https: // doi. org / 10.25100 / socolen. v 44 i 1.6545" type="journal article" year="2018">
Chamorro
<emphasis id="A78D1B62309552ABEE60FE8A74BE0079" box="[887,946,300,326]" italics="true" pageId="207" pageNumber="208">et al.</emphasis>
2018: 74
</bibRefCitation>
</treatmentCitation>
, 81 (list for
<collectingCountry id="EDEE87E0309552ABE9AFFE8B72130079" box="[1208,1311,300,326]" name="Ecuador" pageId="207" pageNumber="208">Ecuador</collectingCountry>
);
<treatmentCitation id="1458E161309552ABE823FE8B761B0055" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." page="244" pageId="207" pageNumber="208" year="2019">
<bibRefCitation id="F168BA81309552ABE823FE8B761B0055" author="Chamorro W. &amp; Marin-Armijos D. &amp; Asenjo A. &amp; Vaz-De-Mello F. Z." pageId="207" pageNumber="208" pagination="1 - 343" refId="ref114107" refString="Chamorro W., Marin-Armijos D., Asenjo A. &amp; Vaz-De-Mello F. Z. 2019. Scarabaeinae dung beetles from Ecuador: a catalog, nomenclatural acts, and distribution records. ZooKeys 826: 1 - 343. https: // doi. org / 10.3897 / zookeys. 826.26488" type="journal article" year="2019">2019: 244</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161309552ABECC1FEE874420055" ID-DOI="http://doi.org/10.5281/zenodo.3684531" ID-Zenodo-Dep="3684531" author="Hielkema A. J. &amp; Hielkema M. A." box="[470,846,335,362]" httpUri="http://treatment.plazi.org/id/03D7FF1CFFE7FFC1FF48FD923373F8C9" page="43" pageId="207" pageNumber="208" year="2019">
<bibRefCitation id="F168BA81309552ABECC1FEE874420055" author="Hielkema A. J. &amp; Hielkema M. A." box="[470,846,335,362]" pageId="207" pageNumber="208" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 43</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770309552ABEDAAFED5726B00C9" blockId="207.[189,1399,265,503]" pageId="207" pageNumber="208">
<treatmentCitationGroup id="B5E9E05E309552ABEDAAFED5726B00C9" pageId="207" pageNumber="208">
<taxonomicName id="52F9BCF3309552ABEDAAFED5765E00B3" ID-CoL="745R" authorityName="Balthasar" authorityYear="1938" box="[189,338,370,396]" class="Insecta" family="Scarabaeidae" genus="Pseuduroxys" kingdom="Animalia" order="Coleoptera" pageId="207" pageNumber="208" phylum="Arthropoda" rank="genus">
<emphasis id="A78D1B62309552ABEDAAFED5765E00B3" box="[189,338,370,396]" italics="true" pageId="207" pageNumber="208">Pseuduroxys</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161309552ABEC6CFED5759500B2" author="Blackwelder R. E." box="[379,665,370,397]" page="204" pageId="207" pageNumber="208" year="1944">
<bibRefCitation id="F168BA81309552ABEC6CFED5759500B2" author="Blackwelder R. E." box="[379,665,370,397]" pageId="207" pageNumber="208" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 204</bibRefCitation>
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(catalog, distribution). —
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