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<document id="E342D890E16609589C3862C8F942A52E" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663076524AEF29F8BE7463057C" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Canthon subcyaneus Erichson 1848" docType="treatment" docVersion="2" lastPageNumber="47" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="45" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="6ADE76C3637706A072189C6436F0BB51">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="A8F71F45B021DB71AEEDC3DB472AC266">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="01F7544B2F70E0E4DCC2D42E1E6068BA">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
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<mods:namePart id="BD63116DFBAC78FD555D1A0B1A8D3C82">Cupello, Mario</mods:namePart>
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<mods:affiliation id="2251B59DC7C905F9EDB63136D2ED31D8">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="39E8B769B432B8F1100FB6B2B8A0AC85" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="1AA35DC4D2806CB11FD13B613D66FAA4">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="F926D3267CBA1464D50285F901E45B92">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<subSubSection id="DDE394FB30765248EF29F8BE74F9060B" box="[574,1013,1817,1844]" pageId="44" pageNumber="45" type="nomenclature">
<paragraph id="9546C77030765248EF29F8BE74F9060B" blockId="44.[574,1013,1817,1879]" box="[574,1013,1817,1844]" pageId="44" pageNumber="45">
<heading id="CE0E701C30765248EF29F8BE74F9060B" box="[574,1013,1817,1844]" centered="true" fontSize="11" level="2" pageId="44" pageNumber="45" reason="2">
<taxonomicName id="52F9BCF330765248EF29F8BE74F9060B" ID-CoL="68XHL" authority="Erichson, 1848" authorityName="Erichson" authorityYear="1848" box="[574,1013,1817,1844]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subcyaneus">
<emphasis id="A78D1B6230765248EF29F8BE7430060B" bold="true" box="[574,828,1817,1844]" italics="true" pageId="44" pageNumber="45">Canthon subcyaneus</emphasis>
<bibRefCitation id="F168BA8130765248EE54F8BD74F9060B" author="Erichson W. F." box="[835,1013,1818,1844]" pageId="44" pageNumber="45" pagination="553 - 1260" refId="ref117915" refString="Erichson W. F. 1848. Insecten. In: Schomburgk R. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil: 553 - 1260. J. J. Weber, Leipzig. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson, 1848</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="DDE394FB30765248EFF1F89A74430668" box="[742,847,1853,1879]" pageId="44" pageNumber="45" type="description">
<paragraph id="9546C77030765248EFF1F89A74430668" blockId="44.[574,1013,1817,1879]" box="[742,847,1853,1879]" pageId="44" pageNumber="45">
<figureCitation id="0DC2DBF530765248EFF1F89A74430668" box="[742,847,1853,1879]" captionStart="Fig" captionStartId="48.[189,232,1963,1989]" captionTargetBox="[274,1307,278,1893]" captionTargetId="figure-18@48.[263,1325,265,1909]" captionTargetPageId="48" captionText="Fig. 19. Species distribution. A. Canthon subcyaneus Erichson, 1848. B. Canthon subhyalinus subhyalinus Harold, 1867. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945510" httpUri="https://zenodo.org/record/13945510/files/figure.png" pageId="44" pageNumber="45">Fig. 19A</figureCitation>
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<paragraph id="9546C77030765248EDAAF8DA74B90684" blockId="44.[189,1400,1917,1979]" pageId="44" pageNumber="45">
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<treatmentCitation id="1458E16130765248EDAAF8DA75A506A7" author="Erichson W. F." box="[189,681,1917,1944]" page="563" pageId="44" pageNumber="45" year="1848">
<taxonomicName id="52F9BCF330765248EDAAF8DA75A506A7" ID-CoL="68XHL" authority="Erichson, 1848: 563" authorityName="Erichson" authorityPageNumber="563" authorityYear="1848" box="[189,681,1917,1944]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="subcyaneus">
<emphasis id="A78D1B6230765248EDAAF8DA76BD06A8" box="[189,433,1917,1943]" italics="true" pageId="44" pageNumber="45">Canthon subcyaneus</emphasis>
<bibRefCitation id="F168BA8130765248ECACF8DA75A506A7" author="Erichson W. F." box="[443,681,1917,1944]" pageId="44" pageNumber="45" pagination="553 - 1260" refId="ref117915" refString="Erichson W. F. 1848. Insecten. In: Schomburgk R. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil: 553 - 1260. J. J. Weber, Leipzig. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson, 1848: 563</bibRefCitation>
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(original description). Type locality:
<collectingCountry id="EDEE87E030765248E972F8D973C806A7" box="[1125,1220,1918,1944]" name="Guyana" pageId="44" pageNumber="45">Guyana</collectingCountry>
. Name-bearing type: a single known
<typeStatus id="4A4279D230765248ECF2F806754C0684" box="[485,576,1953,1979]" pageId="44" pageNumber="45" type="syntype">syntype</typeStatus>
(MFNB), examined by FZVM.
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</paragraph>
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<paragraph id="9546C77030775249EDAAF80C734C06D7" blockId="45.[189,1398,1963,2025]" pageId="45" pageNumber="46">
<emphasis id="A78D1B6230775249EDAAF80C761206FA" bold="true" box="[189,286,1963,1989]" pageId="45" pageNumber="46">Fig. 18.</emphasis>
Species distribution.
<emphasis id="A78D1B6230775249EF3FF80C753106FA" bold="true" box="[552,573,1963,1989]" pageId="45" pageNumber="46">A</emphasis>
.
<taxonomicName id="52F9BCF330775249EF47F80C730E06F9" authority="(Martinez, 1950)" baseAuthorityName="Martinez" baseAuthorityYear="1950" box="[592,1026,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="simulans">
<emphasis id="A78D1B6230775249EF47F80C742406FA" box="[592,808,1963,1989]" italics="true" pageId="45" pageNumber="46">Canthon simulans</emphasis>
(
<bibRefCitation id="F168BA8130775249EE2CF80C74F506F9" author="Martinez A." box="[827,1017,1963,1990]" pageId="45" pageNumber="46" pagination="159 - 172" refId="ref129703" refString="Martinez A. 1950. Notas coleopterologicas IV. Nuevo subgenero y especies de Glaphyrocanthon Martinez, 1948 (Col. Scarabaeineae [sic]). Anales de la Sociedad Cientifica Argentina 150: 159 - 172." type="journal article" year="1950">Martínez, 1950</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="A78D1B6230775249E906F80C732906FA" bold="true" box="[1041,1061,1963,1989]" pageId="45" pageNumber="46">B</emphasis>
.
<taxonomicName id="52F9BCF330775249E920F80C77F706D7" authority="Harold, 1880" authorityName="Harold" authorityYear="1880" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="45" pageNumber="46" phylum="Arthropoda" rank="species" species="steinheili">
<emphasis id="A78D1B6230775249E920F80C721F06FA" box="[1079,1299,1963,1989]" italics="true" pageId="45" pageNumber="46">Canthon steinheili</emphasis>
<bibRefCitation id="F168BA8130775249E809F80C77F706D7" author="Harold E." pageId="45" pageNumber="46" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold, 1880</bibRefCitation>
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<paragraph id="9546C7703074524AEDAAFEAE723700B3" blockId="46.[189,1399,265,537]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="B5E9E05E3074524AEDAAFEAE723700B3" pageId="46" pageNumber="47">
<taxonomicName id="52F9BCF33074524AEDAAFEAE76B8001C" ID-CoL="68XHL" authorityName="Erichson" authorityYear="1848" box="[189,436,265,291]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subcyaneus">
<emphasis id="A78D1B623074524AEDAAFEAE76B8001C" box="[189,436,265,291]" italics="true" pageId="46" pageNumber="47">Canthon subcyaneus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613074524AECCEFEAE75BC001C" author="Harold E." box="[473,688,265,291]" page="16" pageId="46" pageNumber="47" year="1868">
<bibRefCitation id="F168BA813074524AECCEFEAE75BC001C" author="Harold E." box="[473,688,265,291]" pageId="46" pageNumber="47" pagination="1 - 144" refId="ref123312" refString="Harold E. 1868 a. Monographie der gattung Canthon. Berliner entomologische Zeitschrift 12: 1 - 144. https: // doi. org / 10.1002 / mmnd. 18680120105" type="journal article" year="1868">Harold 1868a: 16</bibRefCitation>
</treatmentCitation>
, 117 (key, redescription); 1869d: 994 (catalogue, cited for Cayenne).
<emphasis id="A78D1B623074524AEC78FE8A76850079" box="[367,393,301,326]" italics="true" pageId="46" pageNumber="47"></emphasis>
<treatmentCitation id="1458E1613074524AEC82FE8B75530079" author="Gillet J. J. E." box="[405,607,300,326]" page="34" pageId="46" pageNumber="47" year="1911">
<bibRefCitation id="F168BA813074524AEC82FE8B75530079" author="Gillet J. J. E." box="[405,607,300,326]" pageId="46" pageNumber="47" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 34</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613074524AEE3AFE8B73040079" author="Schmidt A." box="[813,1032,300,326]" page="81" pageId="46" pageNumber="47" year="1922">
<bibRefCitation id="F168BA813074524AEE3AFE8B73040079" author="Schmidt A." box="[813,1032,300,326]" pageId="46" pageNumber="47" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 81</bibRefCitation>
</treatmentCitation>
(distribution). —
<treatmentCitation id="1458E1613074524AE9F9FE8A76580055" author="Boucomont A." page="2" pageId="46" pageNumber="47" year="1928">
<bibRefCitation id="F168BA813074524AE9F9FE8A76580055" author="Boucomont A." pageId="46" pageNumber="47" pagination="1 - 8" refId="ref112356" refString="Boucomont A. 1928 b. Wissenschaftliche Ergebnisse der schwedischen entomologischen Reise des Herrn. Dr. A. Roman in Amazonas 1914 - 1915 (und 1923 - 1924), 12. Scarabaeidae. Arkiv for Zoologi 19 A (22): 1 - 8. https: // doi. org / 10.5962 / bhl. part. 1064" type="journal article" year="1928">Boucomont 1928b: 2</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613074524AECDAFEE875EF0055" author="Blackwelder R. E." box="[461,739,335,362]" page="202" pageId="46" pageNumber="47" year="1944">
<bibRefCitation id="F168BA813074524AECDAFEE875EF0055" author="Blackwelder R. E." box="[461,739,335,362]" pageId="46" pageNumber="47" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 202</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613074524AEE4CFEE873B70056" author="Vulcano M. A. &amp; Pereira F. S." box="[859,1211,335,362]" page="631" pageId="46" pageNumber="47" year="1964">
<bibRefCitation id="F168BA813074524AEE4CFEE873B70056" author="Vulcano M. A. &amp; Pereira F. S." box="[859,1211,335,362]" pageId="46" pageNumber="47" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 631</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613074524AEDFAFED576C400B2" author="Escobar F." box="[237,456,370,397]" page="207" pageId="46" pageNumber="47" year="2000">
<bibRefCitation id="F168BA813074524AEDFAFED576C400B2" author="Escobar F." box="[237,456,370,397]" pageId="46" pageNumber="47" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 207</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03074524AEF64FED575E000B3" box="[627,748,370,396]" name="Colombia" pageId="46" pageNumber="47">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E1613074524AEE33FED5733600B3" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[804,1082,370,397]" page="136" pageId="46" pageNumber="47" year="2001">
<bibRefCitation id="F168BA813074524AEE33FED5733600B3" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[804,1082,370,397]" pageId="46" pageNumber="47" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B623074524AEE95FED474B100B3" box="[898,957,370,396]" italics="true" pageId="46" pageNumber="47">et al.</emphasis>
2001: 136
</bibRefCitation>
</treatmentCitation>
(cited for
<collectingCountry id="EDEE87E03074524AE9A2FED5722200B3" box="[1205,1326,370,396]" name="Colombia" pageId="46" pageNumber="47">Colombia</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703074524AEDAAFE31721B008F" blockId="46.[189,1399,265,537]" box="[189,1303,406,433]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="B5E9E05E3074524AEDAAFE31721B008F" box="[189,1303,406,433]" pageId="46" pageNumber="47">
<taxonomicName id="52F9BCF33074524AEDAAFE3176B6008F" authorityName="Erichson" authorityYear="1849" box="[189,442,406,432]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subcyaneum">
<emphasis id="A78D1B623074524AEDAAFE3176B6008F" box="[189,442,406,432]" italics="true" pageId="46" pageNumber="47">Canthon subcyaneum</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613074524AECCFFE3175CA008F" author="Gacharna C." box="[472,710,406,433]" page="221" pageId="46" pageNumber="47" year="1951">
<bibRefCitation id="F168BA813074524AECCFFE3175CA008F" author="Gacharna C." box="[472,710,406,433]" pageId="46" pageNumber="47" pagination="220 - 229" refId="ref119803" refString="Gacharna C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Fisicas y Naturales 8: 220 - 229." type="journal article" year="1951">Gacharná 1951: 221</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613074524AEE2FFE3174EE008F" author="Roze J. A." box="[824,994,406,433]" page="43" pageId="46" pageNumber="47" year="1955">
<bibRefCitation id="F168BA813074524AEE2FFE3174EE008F" author="Roze J. A." box="[824,994,406,433]" pageId="46" pageNumber="47" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 43</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03074524AE99BFE317206008F" box="[1164,1290,406,432]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703074524AEDAAFE1E76BF0326" blockId="46.[189,1399,265,537]" pageId="46" pageNumber="47">
<treatmentCitationGroup id="B5E9E05E3074524AEDAAFE1E76BF0326" pageId="46" pageNumber="47">
<taxonomicName id="52F9BCF33074524AEDAAFE1E75A900EC" ID-CoL="68XHL" authorityName="Erichson" authorityYear="1848" box="[189,677,441,467]" class="Insecta" family="Scarabaeidae" genus="Canthon" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="subcyaneus" subGenus="Glaphyrocanthon">
<emphasis id="A78D1B623074524AEDAAFE1E762E00EC" box="[189,290,441,467]" italics="true" pageId="46" pageNumber="47">Canthon</emphasis>
(
<emphasis id="A78D1B623074524AEC2EFE1E750B00EC" box="[313,519,441,467]" italics="true" pageId="46" pageNumber="47">Glaphyrocanthon</emphasis>
)
<emphasis id="A78D1B623074524AEF09FE1E75A900EC" box="[542,677,441,467]" italics="true" pageId="46" pageNumber="47">subcyaneus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613074524AEFD8FE1E734E00EC" author="Halffter G. &amp; Martinez A." box="[719,1090,441,467]" page="80" pageId="46" pageNumber="47" year="1977">
<bibRefCitation id="F168BA813074524AEFD8FE1E734E00EC" author="Halffter G. &amp; Martinez A." box="[719,1090,441,467]" pageId="46" pageNumber="47" pagination="29 - 107" refId="ref122649" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina Americanos IV. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 80</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613074524AE9D0FE1D727A00EC" author="Feer F." box="[1223,1398,441,468]" page="62" pageId="46" pageNumber="47" year="2008">
<bibRefCitation id="F168BA813074524AE9D0FE1D727A00EC" author="Feer F." box="[1223,1398,441,468]" pageId="46" pageNumber="47" pagination="49 - 62" refId="ref118745" refString="Feer F. 2008. Responses of dung beetle assemblages to characteristics of rain forest edges. Ecotropica 14: 49 - 62." type="journal article" year="2008">Feer 2008: 62</bibRefCitation>
</treatmentCitation>
(ecology, list); 2013: 676 (list for
<collectingCountry id="EDEE87E03074524AEF95FE7B743B00C9" box="[642,823,476,502]" name="French Guiana" pageId="46" pageNumber="47">French Guiana</collectingCountry>
). —
<treatmentCitation id="1458E1613074524AEE64FE7B73FF00C9" ID-DOI="http://doi.org/10.5281/zenodo.3683546" ID-Zenodo-Dep="3683546" author="Hielkema A. J. &amp; Hielkema M. A." box="[883,1267,476,502]" httpUri="http://treatment.plazi.org/id/03D7FF1CFF8FFFA9FF48FFE4356CF908" page="67" pageId="46" pageNumber="47" year="2019">
<bibRefCitation id="F168BA813074524AEE64FE7B73FF00C9" author="Hielkema A. J. &amp; Hielkema M. A." box="[883,1267,476,502]" pageId="46" pageNumber="47" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 67</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3074524AEDAAFDE676190392" pageId="46" pageNumber="47" type="materials_examined">
<paragraph id="9546C7703074524AEDAAFDE676A60364" blockId="46.[189,426,577,603]" box="[189,426,577,603]" pageId="46" pageNumber="47">
<heading id="CE0E701C3074524AEDAAFDE676A60364" bold="true" box="[189,426,577,603]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A78D1B623074524AEDAAFDE676A60364" bold="true" box="[189,426,577,603]" pageId="46" pageNumber="47">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703074524AEDAAFDD776190392" blockId="46.[189,1399,622,685]" pageId="46" pageNumber="47">
<materialsCitation id="2591CD2D3074524AEDAAFDD7761D0392" collectingDate="2006-08-09" collectorName="de Anacoco" country="Venezuela" location="Isla de Anacoco" pageId="46" pageNumber="47" specimenCount="1" stateProvince="Bolivar">
<collectingCountry id="EDEE87E03074524AEDAAFDD7766703B5" box="[189,363,624,650]" name="Venezuela" pageId="46" pageNumber="47">VENEZUELA</collectingCountry>
<emphasis id="A78D1B623074524AEC9EFDC976E903B6" bold="true" box="[393,485,622,649]" pageId="46" pageNumber="47">
<collectingRegion id="573D09923074524AEC9EFDC976E903B6" box="[393,485,622,649]" country="Venezuela" name="Bolivar" pageId="46" pageNumber="47">Bolívar</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93074524AEF16FDC8755B03B5" box="[513,599,623,650]" count="1" pageId="46" pageNumber="47" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3074524AEF73FDC8742903B5" LSID="urn:lsid:plazi:treatment:1D5076663076524AEF29F8BE7463057C:902691AB3074524AEF73FDC8742903B5" box="[612,805,623,650]" country="Venezuela" name="Isla de Anacoco" pageId="46" pageNumber="47" stateProvince="Bolivar">
Isla
<collectorName id="380CA2A63074524AEF80FDC8742903B5" box="[663,805,623,650]" pageId="46" pageNumber="47">de Anacoco</collectorName>
</location>
;
<date id="E147E1B03074524AEE25FDC874C403B6" box="[818,968,623,650]" pageId="46" pageNumber="47" value="2006-08-09">
<collectingDate id="F10318583074524AEE25FDC874C403B6" box="[818,968,623,650]" pageId="46" pageNumber="47" value="2006-08-09">9 Aug. 2006</collectingDate>
</date>
; curso de NM2006 leg.; 12:42, 48h; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3074524AEDAAFD737463057C" pageId="46" pageNumber="47" type="distribution">
<paragraph id="9546C7703074524AEDAAFD73765A03D1" blockId="46.[189,342,724,750]" box="[189,342,724,750]" pageId="46" pageNumber="47">
<heading id="CE0E701C3074524AEDAAFD73765A03D1" bold="true" box="[189,342,724,750]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A78D1B623074524AEDAAFD73765A03D1" bold="true" box="[189,342,724,750]" pageId="46" pageNumber="47">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703074524AEDAAFCA57528027F" blockId="46.[189,1398,770,832]" pageId="46" pageNumber="47">
<collectingCountry id="EDEE87E03074524AEDAAFCA576380223" box="[189,308,770,796]" name="Colombia" pageId="46" pageNumber="47">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E03074524AEC52FCA576CD0223" box="[325,449,770,796]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E03074524AECC6FCA475230222" box="[465,559,771,797]" name="Guyana" pageId="46" pageNumber="47">Guyana</collectingCountry>
,
<collectingCountry id="EDEE87E03074524AEF56FCA575B80223" box="[577,692,770,796]" name="Suriname" pageId="46" pageNumber="47">Suriname</collectingCountry>
, and
<collectingCountry id="EDEE87E03074524AEFEAFCA574440223" box="[765,840,770,796]" name="Brazil" pageId="46" pageNumber="47">Brazil</collectingCountry>
. Presence in French Guiana to be confirmed (
<bibRefCitation id="F168BA813074524AEDD2FC817514027F" author="Hielkema A. J. &amp; Hielkema M. A." box="[197,536,806,832]" pageId="46" pageNumber="47" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019</bibRefCitation>
).
</paragraph>
<paragraph id="9546C7703074524AEDAAFCC076EE02BE" blockId="46.[189,482,871,897]" box="[189,482,871,897]" pageId="46" pageNumber="47">
<heading id="CE0E701C3074524AEDAAFCC076EE02BE" bold="true" box="[189,482,871,897]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A78D1B623074524AEDAAFCC076EE02BE" bold="true" box="[189,482,871,897]" pageId="46" pageNumber="47">
Subregion of
<collectingCountry id="EDEE87E03074524AEC73FCC076EE02BE" box="[356,482,871,897]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703074524AEDAAFC317362028F" blockId="46.[189,1134,918,944]" box="[189,1134,918,944]" pageId="46" pageNumber="47">
System of low mountains and hills Imataca-Cuyuní of Northeast
<collectingCountry id="EDEE87E03074524AEEAFFC31731F028F" box="[952,1043,918,944]" name="Guyana" pageId="46" pageNumber="47">Guyana</collectingCountry>
Shield.
</paragraph>
<paragraph id="9546C7703074524AEDAAFC7076A802CE" blockId="46.[189,420,983,1009]" box="[189,420,983,1009]" pageId="46" pageNumber="47">
<heading id="CE0E701C3074524AEDAAFC7076A802CE" bold="true" box="[189,420,983,1009]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
<emphasis id="A78D1B623074524AEDAAFC7076A802CE" bold="true" box="[189,420,983,1009]" pageId="46" pageNumber="47">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703074524AEDAAFBA17463057C" blockId="46.[189,1398,1030,1091]" pageId="46" pageNumber="47">
<bibRefCitation id="F168BA813074524AEDAAFBA17697051F" author="Schmidt A." box="[189,411,1030,1056]" pageId="46" pageNumber="47" pagination="61 - 103" refId="ref138378" refString="Schmidt A. 1922. 1. Bestimmungstabelle der mir bekannten Canthon - Arten. 2. Verbreitungsgebiete der Canthon - Arten. 3. Neubeschreibungen von Canthon, Saprositis, Mendidius, Euparia und Ataenius. Archiv fur Naturgeschichte Abteilung A 88 (3): 61 - 103. Available from https: // www. biodiversitylibrary. org / page / 13259701 [accessed 14 Feb. 2024]." type="journal article" year="1922">Schmidt 1922: 81</bibRefCitation>
(
<collectingCountry id="EDEE87E03074524AECA6FBA1753D051F" box="[433,561,1030,1056]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813074524AEF61FBA17470051F" author="Boucomont A." box="[630,892,1030,1056]" pageId="46" pageNumber="47" pagination="1 - 8" refId="ref112356" refString="Boucomont A. 1928 b. Wissenschaftliche Ergebnisse der schwedischen entomologischen Reise des Herrn. Dr. A. Roman in Amazonas 1914 - 1915 (und 1923 - 1924), 12. Scarabaeidae. Arkiv for Zoologi 19 A (22): 1 - 8. https: // doi. org / 10.5962 / bhl. part. 1064" type="journal article" year="1928">Boucomont 1928b: 2</bibRefCitation>
(
<collectingCountry id="EDEE87E03074524AEE85FBA1731E051F" box="[914,1042,1030,1056]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813074524AE940FBA1727A051F" author="Blackwelder R. E." box="[1111,1398,1030,1056]" pageId="46" pageNumber="47" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 202</bibRefCitation>
(
<collectingCountry id="EDEE87E03074524AEDD2FB8E7649057C" box="[197,325,1065,1091]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813074524AEC6BFB8E75D8057C" author="Vulcano M. A. &amp; Pereira F. S." box="[380,724,1065,1091]" pageId="46" pageNumber="47" pagination="570 - 685" refId="ref141320" refString="Vulcano M. A. &amp; Pereira F. S. 1964. Catalogue of the Canthonini (Col. Scarab.). Inhabiting the western hemisphere. Entomologische Arbeiten aus dem Museum G Frey 15 (2): 570 - 685." type="journal article" year="1964">Vulcano &amp; Pereira 1964: 631</bibRefCitation>
(
<collectingCountry id="EDEE87E03074524AEFF4FB8E746F057C" box="[739,867,1065,1091]" name="Venezuela" pageId="46" pageNumber="47">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>