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<document id="7ECD90EB0CC8384D2623FD3EFA6AE347" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666306E5250ECEEFECA757C04B0" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Canthonella gomezi" docType="treatment" docVersion="2" lastPageNumber="53" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="53" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="E01D0A11FC2935A8F6A157749D6AFA74">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:name id="A7A8FA64D86ED5F1D66D8B04D6083548" type="personal">
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<mods:namePart id="5552504B18FB97B1E2E4CAF66ACC2A7A">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="5E27FB759648F72591D1E37B288544F7">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="FD9FD2DC60667DC219C3D79ED7A44025" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="FFA0877C12CE2ABF718978D2E75C71DF">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="F43277549F0438C060813005DAA4750D" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="F94E3ADD8D5564F75D76652996A0A77B">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="F17168045D734C850EDA9CA437E1C805" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="EA31343F86FE3B428D4BEBE52D29EC8A">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="773C7894415A7E51243DC920A689DCEF" type="LSID">2BC093C2-3E2B-466B-B31C-9F3D9FA844BC</mods:nameIdentifier>
<mods:affiliation id="ECFF7D1D99D7D5427E46EFC9BFD6C9C0">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="348217631C20C774D185294A55B72012" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="CC7210987BE4BFC18EA9E00334B1866A">European Journal of Taxonomy</mods:title>
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<mods:date id="DBEA675568DF5BAF649D1477BF607DE9">2024</mods:date>
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<treatment id="1D507666306E5250ECEEFECA757C04B0" ID-DOI="http://doi.org/10.5281/zenodo.13945696" ID-Zenodo-Dep="13945696" LSID="urn:lsid:plazi:treatment:1D507666306E5250ECEEFECA757C04B0" httpUri="http://treatment.plazi.org/id/1D507666306E5250ECEEFECA757C04B0" lastPageNumber="53" pageId="52" pageNumber="53">
<subSubSection id="DDE394FB306E5250ECEEFECA733600B8" box="[505,1082,365,392]" pageId="52" pageNumber="53" type="nomenclature">
<paragraph id="9546C770306E5250ECEEFECA733600B8" blockId="52.[505,1082,365,427]" box="[505,1082,365,392]" pageId="52" pageNumber="53">
<heading id="CE0E701C306E5250ECEEFECA733600B8" box="[505,1082,365,392]" centered="true" fontSize="11" level="2" pageId="52" pageNumber="53" reason="2">
<taxonomicName id="52F9BCF3306E5250ECEEFECA733600B8" ID-CoL="QPFK" authority="(Halffter &amp; Martinez, 1968)" baseAuthorityName="Halffter &amp; Martinez" baseAuthorityYear="1968" box="[505,1082,365,392]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gomezi">
<emphasis id="A78D1B62306E5250ECEEFECA75E700B8" bold="true" box="[505,747,365,392]" italics="true" pageId="52" pageNumber="53">Canthonella gomezi</emphasis>
(
<bibRefCitation id="F168BA81306E5250EFEDFEC9733E00B8" author="Halffter G. &amp; Martinez A." box="[762,1074,365,392]" pageId="52" pageNumber="53" pagination="209 - 290" refId="ref122569" refString="Halffter G. &amp; Martinez A. 1968. Revision monografica de los Canthonina Americanos III. Revista de la Sociedad Mexicana de Historia Natural 29: 209 - 290." type="journal article" year="1968">Halffter &amp; Martínez, 1968</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB306E5250EFC3FE3674530094" box="[724,863,401,427]" pageId="52" pageNumber="53" type="description">
<paragraph id="9546C770306E5250EFC3FE3674530094" blockId="52.[505,1082,365,427]" box="[724,863,401,427]" pageId="52" pageNumber="53">
<figureCitation id="0DC2DBF5306E5250EFC3FE3674530094" box="[724,863,401,427]" captionStart="Fig" captionStartId="53.[189,232,1928,1954]" captionTargetBox="[202,1357,314,1832]" captionTargetId="figure-11@53.[189,1398,281,1865]" captionTargetPageId="53" captionText="Fig. 21. Species distribution. A. Canthonella gomezi (Halffter &amp; Martínez, 1968). B. Dorsal view of C. gomezi. C. Coprophanaeus abas (MacLeay, 1819). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945515" httpUri="https://zenodo.org/record/13945515/files/figure.png" pageId="52" pageNumber="53">Fig. 21AB</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB306E5250EDAAFE75726B034B" pageId="52" pageNumber="53" type="reference_group">
<paragraph id="9546C770306E5250EDAAFE7573B9032F" blockId="52.[189,1399,466,528]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="B5E9E05E306E5250EDAAFE7573B9032F" pageId="52" pageNumber="53">
<treatmentCitation id="1458E161306E5250EDAAFE75742F00D3" author="Halffter G. &amp; Martinez A." box="[189,803,466,492]" page="221" pageId="52" pageNumber="53" year="1968">
<taxonomicName id="52F9BCF3306E5250EDAAFE75742F00D3" ID-CoL="3PXN4" authority="Halffter &amp; Martinez, 1968: 221" authorityName="Halffter &amp; Martinez" authorityPageNumber="221" authorityYear="1968" box="[189,803,466,492]" class="Insecta" family="Scarabaeidae" genus="Ipselissus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gomezi">
<emphasis id="A78D1B62306E5250EDAAFE75768300D3" box="[189,399,466,492]" italics="true" pageId="52" pageNumber="53">Ipselissus gomezi</emphasis>
<bibRefCitation id="F168BA81306E5250EC8BFE75742F00D3" author="Halffter G. &amp; Martinez A." box="[412,803,466,492]" pageId="52" pageNumber="53" pagination="209 - 290" refId="ref122569" refString="Halffter G. &amp; Martinez A. 1968. Revision monografica de los Canthonina Americanos III. Revista de la Sociedad Mexicana de Historia Natural 29: 209 - 290." type="journal article" year="1968">Halffter &amp; Martínez, 1968: 221</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0306E5250E9E1FE75727F00D3" box="[1270,1395,466,492]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992306E5250EDFDFE52765E0330" box="[234,338,501,527]" country="Venezuela" name="Miranda" pageId="52" pageNumber="53">Miranda</collectingRegion>
: Altos de Pipe. Name-bearing type:
<typeStatus id="4A4279D2306E5250EFEEFE52746C0330" box="[761,864,501,527]" pageId="52" pageNumber="53" type="holotype">holotype</typeStatus>
(MACN), examined by MC.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770306E5250EDAAFD90726B034B" blockId="52.[189,1399,567,629]" pageId="52" pageNumber="53">
<treatmentCitationGroup id="B5E9E05E306E5250EDAAFD90726B034B" pageId="52" pageNumber="53">
<taxonomicName id="52F9BCF3306E5250EDAAFD9076AB036E" ID-CoL="QPFK" baseAuthorityName="Halffter &amp; Martinez" baseAuthorityYear="1968" box="[189,423,567,593]" class="Insecta" family="Scarabaeidae" genus="Canthonella" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gomezi">
<emphasis id="A78D1B62306E5250EDAAFD9076AB036E" box="[189,423,567,593]" italics="true" pageId="52" pageNumber="53">Canthonella gomezi</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161306E5250ECD8FD90742F036D" author="Halffter G. &amp; Martinez A." box="[463,803,567,594]" page="58" pageId="52" pageNumber="53" year="1977">
<bibRefCitation id="F168BA81306E5250ECD8FD90742F036D" author="Halffter G. &amp; Martinez A." box="[463,803,567,594]" pageId="52" pageNumber="53" pagination="29 - 107" refId="ref122649" refString="Halffter G. &amp; Martinez A. 1977. Revision monografica de los Canthonina Americanos IV. Clave para generos y subgeneros. Folia Entomologica Mexicana 38: 29 - 107." type="journal article" year="1977">Halffter &amp; Martínez 1977: 58</bibRefCitation>
</treatmentCitation>
(new synonym, key). —
<bibRefCitation id="F168BA81306E5250E950FD907221036E" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[1095,1325,567,593]" pageId="52" pageNumber="53" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
<collectingRegion id="573D0992306E5250E950FD9073AD036E" box="[1095,1185,567,593]" country="Saudi Arabia" name="Al Madinah" pageId="52" pageNumber="53">Medina</collectingRegion>
<emphasis id="A78D1B62306E5250E9B1FD9F73EC036E" box="[1190,1248,567,593]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
, 2001
</bibRefCitation>
: (list); 2003: 65 (phylogenetic analysis). —
<treatmentCitation id="1458E161306E5250EF8BFDFD746E034B" author="Krajcik M." box="[668,866,602,628]" page="64" pageId="52" pageNumber="53" year="2012">
<bibRefCitation id="F168BA81306E5250EF8BFDFD746E034B" author="Krajcik M." box="[668,866,602,628]" pageId="52" pageNumber="53" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 64</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E161306E5250EEC2FDFD722D034B" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[981,1313,602,628]" page="109" pageId="52" pageNumber="53" year="2013">
<bibRefCitation id="F168BA81306E5250EEC2FDFD722D034B" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[981,1313,602,628]" pageId="52" pageNumber="53" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B62306E5250E97EFDFC73A8034B" box="[1129,1188,602,628]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
2013: 109
</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB306E5250EDAAFD3C75A602C6" pageId="52" pageNumber="53" type="materials_examined">
<paragraph id="9546C770306E5250EDAAFD3C76A6038A" blockId="52.[189,426,667,693]" box="[189,426,667,693]" pageId="52" pageNumber="53">
<heading id="CE0E701C306E5250EDAAFD3C76A6038A" bold="true" box="[189,426,667,693]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="A78D1B62306E5250EDAAFD3C76A6038A" bold="true" box="[189,426,667,693]" pageId="52" pageNumber="53">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770306E5250EDFDFD6D765003DB" blockId="52.[189,1399,714,846]" box="[234,348,714,740]" pageId="52" pageNumber="53">
<emphasis id="A78D1B62306E5250EDFDFD6D765003DB" bold="true" box="[234,348,714,740]" pageId="52" pageNumber="53">
<typeStatus id="4A4279D2306E5250EDFDFD6D765003DB" box="[234,348,714,740]" pageId="52" pageNumber="53" type="paratype">Paratype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="9546C770306E5250EDAAFD4A76360271" blockId="52.[189,1399,714,846]" pageId="52" pageNumber="53">
<materialsCitation id="2591CD2D306E5250EDAAFD4A763A0271" collectorName="Martinez" country="Venezuela" location="Altos de Pipe" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Miranda" typeStatus="paratype">
<collectingCountry id="EDEE87E0306E5250EDAAFD4A76670238" box="[189,363,749,775]" name="Venezuela" pageId="52" pageNumber="53">VENEZUELA</collectingCountry>
<emphasis id="A78D1B62306E5250EC91FD4A76F90238" bold="true" box="[390,501,749,775]" pageId="52" pageNumber="53">
<collectingRegion id="573D0992306E5250EC91FD4A76F90238" box="[390,501,749,775]" country="Venezuela" name="Miranda" pageId="52" pageNumber="53">Miranda</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9306E5250EF1AFD4A756E0237" box="[525,610,749,776]" count="1" pageId="52" pageNumber="53" type="generic">1 spec.</specimenCount>
; “Venezuela / E° Miranda /
<location id="902691AB306E5250EEB1FD4A734F0238" LSID="urn:lsid:plazi:treatment:1D507666306E5250ECEEFECA757C04B0:902691AB306E5250EEB1FD4A734F0238" box="[934,1091,749,775]" country="Venezuela" name="Altos de Pipe" pageId="52" pageNumber="53" stateProvince="Miranda">Altos de Pipe</location>
/ a la luz / coll.
<collectorName id="380CA2A6306E5250E9EAFD4A726B0238" box="[1277,1383,749,775]" pageId="52" pageNumber="53">Martínez</collectorName>
/ Jun. 948”; “
<taxonomicName id="52F9BCF3306E5250EC47FCB775130215" authorityName="Halffter &amp; Martinez" authorityYear="1968" box="[336,543,784,810]" class="Insecta" family="Scarabaeidae" genus="Ipselissus" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="gomezi">
<emphasis id="A78D1B62306E5250EC47FCB775130215" box="[336,543,784,810]" italics="true" pageId="52" pageNumber="53">Ipselissus gomezi</emphasis>
</taxonomicName>
/ Halffter y / Martínez”; “HALFFTER collection [red label]”; CEMT-00078136
</materialsCitation>
.
</paragraph>
<paragraph id="9546C770306E5250EDFDFCD276ED02B0" blockId="52.[189,1399,885,1017]" box="[234,481,885,911]" pageId="52" pageNumber="53">
<emphasis id="A78D1B62306E5250EDFDFCD276ED02B0" bold="true" box="[234,481,885,911]" pageId="52" pageNumber="53">Additional material</emphasis>
</paragraph>
<paragraph id="9546C770306E5250EDAAFC3F75A602C6" blockId="52.[189,1399,885,1017]" pageId="52" pageNumber="53">
<materialsCitation id="2591CD2D306E5250EDAAFC3F738802E9" collectingDate="2008-09-12" country="Venezuela" location="Instituto Venezolano de Investigaciones Cientificas" municipality="Altos de Pipe" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Miranda">
<collectingCountry id="EDEE87E0306E5250EDAAFC3F7667028D" box="[189,363,920,946]" name="Venezuela" pageId="52" pageNumber="53">VENEZUELA</collectingCountry>
<emphasis id="A78D1B62306E5250EC9FFC3F76FB028D" bold="true" box="[392,503,920,946]" pageId="52" pageNumber="53">
<collectingRegion id="573D0992306E5250EC9FFC3F76FB028D" box="[392,503,920,946]" country="Venezuela" name="Miranda" pageId="52" pageNumber="53">Miranda</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9306E5250EF07FC3F7569028C" box="[528,613,920,947]" count="1" pageId="52" pageNumber="53" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="75225D0A306E5250EF67FC3F741E028D" box="[624,786,920,946]" pageId="52" pageNumber="53">Altos de Pipe</collectingMunicipality>
,
<location id="902691AB306E5250EE09FC3F727A028D" LSID="urn:lsid:plazi:treatment:1D507666306E5250ECEEFECA757C04B0:902691AB306E5250EE09FC3F727A028D" box="[798,1398,920,946]" country="Venezuela" municipality="Altos de Pipe" name="Instituto Venezolano de Investigaciones Cientificas" pageId="52" pageNumber="53" stateProvince="Miranda">Instituto Venezolano de Investigaciones Científicas</location>
(“IVIC”);
<date id="E147E1B0306E5250EC20FC1B76DA02E9" box="[311,470,956,982]" pageId="52" pageNumber="53" value="2008-09-12">
<collectingDate id="F1031858306E5250EC20FC1B76DA02E9" box="[311,470,956,982]" pageId="52" pageNumber="53" value="2008-09-12">12 Sep. 2008</collectingDate>
</date>
; “pasantes BDV2008” leg.; “BV-T-021, cebo:(?)”; CEMT
</materialsCitation>
<materialsCitation id="2591CD2D306E5250E988FC1B75AA02C6" collectingDate="2008-09-12" country="Venezuela" location="Instituto Venezolano de Investigaciones Cientificas" municipality="Altos de Pipe" pageId="52" pageNumber="53" specimenCount="1" stateProvince="Miranda">
<specimenCount id="83FF0CF9306E5250E988FC1B73F802E9" box="[1183,1268,956,982]" count="1" pageId="52" pageNumber="53" type="generic">1 spec.</specimenCount>
; same data as for the preceding; “BV-T-035”; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB306E5250EDAAFB87757C04B0" pageId="52" pageNumber="53" type="distribution">
<paragraph id="9546C770306E5250EDAAFB87765A0505" blockId="52.[189,342,1056,1082]" box="[189,342,1056,1082]" pageId="52" pageNumber="53">
<heading id="CE0E701C306E5250EDAAFB87765A0505" bold="true" box="[189,342,1056,1082]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="A78D1B62306E5250EDAAFB87765A0505" bold="true" box="[189,342,1056,1082]" pageId="52" pageNumber="53">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770306E5250EDAAFBE874B00556" blockId="52.[189,956,1102,1129]" box="[189,956,1102,1129]" pageId="52" pageNumber="53">
<collectingCountry id="EDEE87E0306E5250EDAAFBE8763D0556" box="[189,305,1103,1129]" name="Colombia" pageId="52" pageNumber="53">Colombia</collectingCountry>
and
<collectingCountry id="EDEE87E0306E5250EC7EFBE876EA0556" box="[361,486,1103,1129]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
. Possibly in
<collectingCountry id="EDEE87E0306E5250EF6EFBE875A30556" box="[633,687,1103,1129]" name="Peru" pageId="52" pageNumber="53">Peru</collectingCountry>
(
<bibRefCitation id="F168BA81306E5250EFA9FBE874BC0556" author="Ratcliffe B. C. &amp; Jameson M. L. &amp; Figueroa L. &amp; Cave R. D. &amp; Paulsen M. J. &amp; Cano E. B. &amp; Beza-Beza C. &amp; Ferbans L. &amp; Reyes-Castillo P." box="[702,944,1102,1129]" pageId="52" pageNumber="53" pagination="186 - 207" refId="ref137018" refString="Ratcliffe B. C., Jameson M. L., Figueroa L., Cave R. D., Paulsen M. J., Cano E. B., Beza-Beza C., Jimenez- Ferbans L., Reyes-Castillo P. 2015. Beetles (Coleoptera) of Peru: a survey of the families. Journal of the Kansas Entomological Society 88 (2): 186 - 207." type="journal article" year="2015">
Ratcliffe
<emphasis id="A78D1B62306E5250EE39FBE874640556" box="[814,872,1103,1129]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
2015
</bibRefCitation>
).
</paragraph>
<paragraph id="9546C770306E5250EDAAFB3776EE0595" blockId="52.[189,482,1168,1194]" box="[189,482,1168,1194]" pageId="52" pageNumber="53">
<heading id="CE0E701C306E5250EDAAFB3776EE0595" bold="true" box="[189,482,1168,1194]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="A78D1B62306E5250EDAAFB3776EE0595" bold="true" box="[189,482,1168,1194]" pageId="52" pageNumber="53">
Subregion of
<collectingCountry id="EDEE87E0306E5250EC73FB3776EE0595" box="[356,482,1168,1194]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770306E5250EDAAFB18766005E6" blockId="52.[189,364,1215,1241]" box="[189,364,1215,1241]" pageId="52" pageNumber="53">Guiana Shield.</paragraph>
<paragraph id="9546C770306E5250EDAAFAA776A80425" blockId="52.[189,420,1280,1306]" box="[189,420,1280,1306]" pageId="52" pageNumber="53">
<heading id="CE0E701C306E5250EDAAFAA776A80425" bold="true" box="[189,420,1280,1306]" fontSize="11" level="3" pageId="52" pageNumber="53" reason="3">
<emphasis id="A78D1B62306E5250EDAAFAA776A80425" bold="true" box="[189,420,1280,1306]" pageId="52" pageNumber="53">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770306E5250EDAAFA88757C04B0" blockId="52.[189,1398,1326,1423]" pageId="52" pageNumber="53">
<bibRefCitation id="F168BA81306E5250EDAAFA8875460476" author="Halffter G. &amp; Martinez A." box="[189,586,1326,1353]" pageId="52" pageNumber="53" pagination="209 - 290" refId="ref122569" refString="Halffter G. &amp; Martinez A. 1968. Revision monografica de los Canthonina Americanos III. Revista de la Sociedad Mexicana de Historia Natural 29: 209 - 290." type="journal article" year="1968">Halffter &amp; Martínez 1968: 221</bibRefCitation>
(
<collectingCountry id="EDEE87E0306E5250EF74FA8875EE0476" box="[611,738,1327,1353]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992306E5250EFE3FA8874460476" box="[756,842,1327,1353]" country="Venezuela" name="Aragua" pageId="52" pageNumber="53">Aragua</collectingRegion>
and
<collectingRegion id="573D0992306E5250EE82FA8874F20476" box="[917,1022,1327,1353]" country="Venezuela" name="Miranda" pageId="52" pageNumber="53">Miranda</collectingRegion>
). —
<bibRefCitation id="F168BA81306E5250E95FFA88727A0476" author="Medina C. A. &amp; Scholtz C. H. &amp; Gill B. D." box="[1096,1398,1326,1353]" pageId="52" pageNumber="53" pagination="23 - 68" refId="ref130892" refString="Medina C. A., Scholtz C. H. &amp; Gill B. D. 2003. Morphological variation and systematics of Canthon and related genera of new world Canthonini dung beetles. Deutsche Entomologische Zeitschrift 50 (1): 23 - 68. https: // doi. org / 10.1002 / mmnd. 20030500105" type="journal article" year="2003">
Medina
<emphasis id="A78D1B62306E5250E9A5FA8873E20476" box="[1202,1262,1327,1353]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. 2003: 65
</bibRefCitation>
(
<collectingCountry id="EDEE87E0306E5250EDD2FAF576490453" box="[197,325,1362,1388]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81306E5250EC69FAF5754B0453" author="Krajcik M." box="[382,583,1362,1388]" pageId="52" pageNumber="53" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 64</bibRefCitation>
(
<collectingCountry id="EDEE87E0306E5250EF40FAF575DA0453" box="[599,726,1362,1388]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992306E5250EFF5FAF574460453" box="[738,842,1362,1388]" country="Venezuela" name="Miranda" pageId="52" pageNumber="53">Miranda</collectingRegion>
: Alto de Pipe). —
<bibRefCitation id="F168BA81306E5250E93FFAF5727A0453" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[1064,1398,1362,1388]" pageId="52" pageNumber="53" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B62306E5250E9ABFAF473FC0453" box="[1212,1264,1362,1388]" italics="true" pageId="52" pageNumber="53">et al</emphasis>
. 2013: 109
</bibRefCitation>
(
<collectingCountry id="EDEE87E0306E5250EDD1FAD2764904B0" box="[198,325,1397,1423]" name="Venezuela" pageId="52" pageNumber="53">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992306E5250EC47FAD276B404B0" box="[336,440,1397,1423]" country="Venezuela" name="Miranda" pageId="52" pageNumber="53">Miranda</collectingRegion>
: Altos de Pipe).
</paragraph>
</subSubSection>
</treatment>
</document>