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<document id="39BC2731C5A35B3C00E4D0AA20E328B6" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663043527EEF33F8BA73C803D8" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Bdelyropsis venezuelensis Howden 1976" docType="treatment" docVersion="2" lastPageNumber="27" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="26" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="56BA023569DB70AFF8A89B651B86D47E">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="57CDCDD9AB2DC265D413EE71E1BD2CA2">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:nameIdentifier id="374194C03998DCAFA17ADE8467EA67AD" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
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<mods:namePart id="A87C48377182147B15B722DF8A9B1FC0">Cupello, Mario</mods:namePart>
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<mods:affiliation id="3CD8D26B7C536D126DCAE7B6CAD958BA">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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<mods:namePart id="59460419A9C901EA01379779E27497B1">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:affiliation id="3B61F08D8B58DB97A58444AAB8A84CCA">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
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<paragraph id="9546C7703043527DEF33F8BA731C0607" blockId="25.[548,1040,1821,1883]" box="[548,1040,1821,1848]" pageId="25" pageNumber="26">
<heading id="CE0E701C3043527DEF33F8BA731C0607" box="[548,1040,1821,1848]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
<taxonomicName id="52F9BCF33043527DEF33F8BA731C0607" ID-CoL="L59J" authority="Howden, 1976" authorityName="Howden" authorityYear="1976" box="[548,1040,1821,1848]" class="Insecta" family="Scarabaeidae" genus="Bdelyropsis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="A78D1B623043527DEF33F8BA74500607" bold="true" box="[548,860,1821,1848]" italics="true" pageId="25" pageNumber="26">Bdelyropsis venezuelensis</emphasis>
<bibRefCitation id="F168BA813043527DEE74F8B9731C0607" author="Howden H. F." box="[867,1040,1822,1848]" pageId="25" pageNumber="26" pagination="95 - 103" refId="ref124712" refString="Howden H. F. 1976. New species in the genera Bdelyropsis, Cryptocanthon and Drepanocerus (Coleoptera, Scarabaeidae). Proceedings of the Entomological Society of Washington 78: 95 - 103. Available from https: // www. biodiversitylibrary. org / page / 16252570 [accessed 16 Feb. 2024]." type="journal article" year="1976">Howden, 1976</bibRefCitation>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="DDE394FB3043527DEFF1F8E674420664" box="[742,846,1857,1883]" pageId="25" pageNumber="26" type="description">
<paragraph id="9546C7703043527DEFF1F8E674420664" blockId="25.[548,1040,1821,1883]" box="[742,846,1857,1883]" pageId="25" pageNumber="26">
<figureCitation id="0DC2DBF53043527DEFF1F8E674420664" box="[742,846,1857,1883]" captionStart="Fig" captionStartId="27.[189,232,1961,1987]" captionTargetBox="[281,1298,278,1874]" captionTargetId="figure-11@27.[270,1318,265,1911]" captionTargetPageId="27" captionText="Fig. 12. Species distribution. A. Bdelyropsis venezuelensis Howden, 1976. B. Bdelyrus leptomerus Cook, 1998. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945490" httpUri="https://zenodo.org/record/13945490/files/figure.png" pageId="25" pageNumber="26">Fig. 12A</figureCitation>
</paragraph>
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<subSubSection id="DDE394FB3043527DEDAAF825766806DC" pageId="25" pageNumber="26" type="reference_group">
<paragraph id="9546C7703043527DEDAAF825766806DC" blockId="25.[189,1399,1922,2019]" pageId="25" pageNumber="26">
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<treatmentCitation id="1458E1613043527DEDAAF82575CB06A2" author="Howden H. F." box="[189,711,1922,1949]" page="95" pageId="25" pageNumber="26" year="1976">
<taxonomicName id="52F9BCF33043527DEDAAF82575CB06A2" ID-CoL="L59J" authority="Howden, 1976: 95" authorityName="Howden" authorityPageNumber="95" authorityYear="1976" box="[189,711,1922,1949]" class="Insecta" family="Scarabaeidae" genus="Bdelyropsis" kingdom="Animalia" order="Coleoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="species" species="venezuelensis">
<emphasis id="A78D1B623043527DEDAAF82576E006A3" box="[189,492,1922,1948]" italics="true" pageId="25" pageNumber="26">Bdelyropsis venezuelensis</emphasis>
<bibRefCitation id="F168BA813043527DECE6F82575CB06A2" author="Howden H. F." box="[497,711,1922,1949]" pageId="25" pageNumber="26" pagination="95 - 103" refId="ref124712" refString="Howden H. F. 1976. New species in the genera Bdelyropsis, Cryptocanthon and Drepanocerus (Coleoptera, Scarabaeidae). Proceedings of the Entomological Society of Washington 78: 95 - 103. Available from https: // www. biodiversitylibrary. org / page / 16252570 [accessed 16 Feb. 2024]." type="journal article" year="1976">Howden, 1976: 95</bibRefCitation>
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(original description). Type locality:
<collectingCountry id="EDEE87E03043527DE961F82573FF06A3" box="[1142,1267,1922,1948]" name="Venezuela" pageId="25" pageNumber="26">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923043527DE9EBF825727F06A3" box="[1276,1395,1922,1948]" country="Venezuela" name="Carabobo" pageId="25" pageNumber="26">Carabobo</collectingRegion>
: Cerro el Cafe. Name-bearing type:
<typeStatus id="4A4279D23043527DEF81F80275F10680" box="[662,765,1957,1983]" pageId="25" pageNumber="26" type="holotype">holotype</typeStatus>
(CMNC) (originally, in HAHC;
<bibRefCitation id="F168BA813043527DE99AF80272350680" author="Howden H. F." box="[1165,1337,1957,1983]" pageId="25" pageNumber="26" pagination="95 - 103" refId="ref124712" refString="Howden H. F. 1976. New species in the genera Bdelyropsis, Cryptocanthon and Drepanocerus (Coleoptera, Scarabaeidae). Proceedings of the Entomological Society of Washington 78: 95 - 103. Available from https: // www. biodiversitylibrary. org / page / 16252570 [accessed 16 Feb. 2024]." type="journal article" year="1976">Howden 1976</bibRefCitation>
), not examined.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3040527EEDAAFEAF727F004B" pageId="26" pageNumber="27" type="materials_examined">
<paragraph id="9546C7703040527EEDAAFEAF76A6001D" blockId="26.[189,426,264,290]" box="[189,426,264,290]" pageId="26" pageNumber="27">
<heading id="CE0E701C3040527EEDAAFEAF76A6001D" bold="true" box="[189,426,264,290]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="A78D1B623040527EEDAAFEAF76A6001D" bold="true" box="[189,426,264,290]" pageId="26" pageNumber="27">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703040527EEDAAFE90727F004B" blockId="26.[189,1399,311,373]" pageId="26" pageNumber="27">
<materialsCitation id="2591CD2D3040527EEDAAFE907250006E" box="[189,1372,311,338]" collectingDate="1973-01-31" collectorName="Bordon" country="Venezuela" elevation="100" location="San Esteban" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Carabobo">
<collectingCountry id="EDEE87E03040527EEDAAFE907667006E" box="[189,363,311,337]" name="Venezuela" pageId="26" pageNumber="27">VENEZUELA</collectingCountry>
<emphasis id="A78D1B623040527EEC9EFE90750A006E" bold="true" box="[393,518,311,337]" pageId="26" pageNumber="27">
<collectingRegion id="573D09923040527EEC9EFE90750A006E" box="[393,518,311,337]" country="Venezuela" name="Carabobo" pageId="26" pageNumber="27">Carabobo</collectingRegion>
</emphasis>
• 2 specs;
<location id="902691AB3040527EEF9EFE907412006E" LSID="urn:lsid:plazi:treatment:1D5076663043527EEF33F8BA73C803D8:902691AB3040527EEF9EFE907412006E" box="[649,798,311,337]" country="Venezuela" name="San Esteban" pageId="26" pageNumber="27" stateProvince="Carabobo">San Esteban</location>
;
<elevation id="1ED420433040527EEE3DFE9074BB006E" box="[810,951,311,338]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="m" value="100.0">
<quantity id="52016A953040527EEE3DFE90747A006D" box="[810,886,311,338]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="26" pageNumber="27" unit="m" value="100.0">100 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03040527EEED2FE90736C006E" box="[965,1120,311,338]" pageId="26" pageNumber="27" value="1973-01-31">
<collectingDate id="F10318583040527EEED2FE90736C006E" box="[965,1120,311,338]" pageId="26" pageNumber="27" value="1973-01-31">31 Jan. 1973</collectingDate>
</date>
;
<collectorName id="380CA2A63040527EE97AFE9073C9006E" box="[1133,1221,311,337]" pageId="26" pageNumber="27">Bordón</collectorName>
leg.; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D3040527EEDAAFEFD7263004B" box="[189,1391,346,373]" collectingDate="2005-11-27" country="Venezuela" location="Hacienda Guaquira" pageId="26" pageNumber="27" specimenCount="1" stateProvince="Yaracuy">
<emphasis id="A78D1B623040527EEDAAFEFD7629004B" bold="true" box="[189,293,346,372]" pageId="26" pageNumber="27">
<collectingRegion id="573D09923040527EEDAAFEFD7629004B" box="[189,293,346,372]" country="Venezuela" name="Yaracuy" pageId="26" pageNumber="27">Yaracuy</collectingRegion>
</emphasis>
• 2 specs;
<location id="902691AB3040527EECB4FEFD7586004B" LSID="urn:lsid:plazi:treatment:1D5076663043527EEF33F8BA73C803D8:902691AB3040527EECB4FEFD7586004B" box="[419,650,346,372]" country="Venezuela" name="Hacienda Guaquira" pageId="26" pageNumber="27" stateProvince="Yaracuy">Hacienda Guáquira</location>
;
<date id="E147E1B03040527EEF81FEFD7434004A" box="[662,824,346,373]" pageId="26" pageNumber="27" value="2005-11-27">
<collectingDate id="F10318583040527EEF81FEFD7434004A" box="[662,824,346,373]" pageId="26" pageNumber="27" value="2005-11-27">27 Nov. 2005</collectingDate>
</date>
; curso NM2006 leg.; 08:15, 25h, faeces; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3040527EEDAAFE3C73C803D8" pageId="26" pageNumber="27" type="distribution">
<paragraph id="9546C7703040527EEDAAFE3C765A008A" blockId="26.[189,342,411,437]" box="[189,342,411,437]" pageId="26" pageNumber="27">
<heading id="CE0E701C3040527EEDAAFE3C765A008A" bold="true" box="[189,342,411,437]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="A78D1B623040527EEDAAFE3C765A008A" bold="true" box="[189,342,411,437]" pageId="26" pageNumber="27">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703040527EEDAAFE6D757000DB" blockId="26.[189,636,458,484]" box="[189,636,458,484]" pageId="26" pageNumber="27">
<collectingCountry id="EDEE87E03040527EEDAAFE6D763A00DB" box="[189,310,458,484]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
(endemic) (
<bibRefCitation id="F168BA813040527EECD4FE6D756300DB" author="Howden H. F." box="[451,623,458,484]" pageId="26" pageNumber="27" pagination="95 - 103" refId="ref124712" refString="Howden H. F. 1976. New species in the genera Bdelyropsis, Cryptocanthon and Drepanocerus (Coleoptera, Scarabaeidae). Proceedings of the Entomological Society of Washington 78: 95 - 103. Available from https: // www. biodiversitylibrary. org / page / 16252570 [accessed 16 Feb. 2024]." type="journal article" year="1976">Howden 1976</bibRefCitation>
).
</paragraph>
<paragraph id="9546C7703040527EEDAAFDAC76EE0319" blockId="26.[189,482,523,550]" box="[189,482,523,550]" pageId="26" pageNumber="27">
<heading id="CE0E701C3040527EEDAAFDAC76EE0319" bold="true" box="[189,482,523,550]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="A78D1B623040527EEDAAFDAC76EE0319" bold="true" box="[189,482,523,550]" pageId="26" pageNumber="27">
Subregion of
<collectingCountry id="EDEE87E03040527EEC73FDAB76EE0319" box="[356,482,524,550]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703040527EEDAAFD9D753D036A" blockId="26.[189,561,570,597]" box="[189,561,570,597]" pageId="26" pageNumber="27">
Central
<collectingRegion id="573D09923040527EEC0CFD9D7652036B" box="[283,350,570,596]" country="New Zealand" name="West Coast" pageId="26" pageNumber="27">Coast</collectingRegion>
Mountain Range.
</paragraph>
<paragraph id="9546C7703040527EEDAAFDDC76A803A9" blockId="26.[189,420,635,662]" box="[189,420,635,662]" pageId="26" pageNumber="27">
<heading id="CE0E701C3040527EEDAAFDDC76A803A9" bold="true" box="[189,420,635,662]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
<emphasis id="A78D1B623040527EEDAAFDDC76A803A9" bold="true" box="[189,420,635,662]" pageId="26" pageNumber="27">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703040527EEDAAFD0D73C803D8" blockId="26.[189,1398,682,744]" pageId="26" pageNumber="27">
<bibRefCitation id="F168BA813040527EEDAAFD0D769003FA" author="Howden H. F." box="[189,412,682,709]" pageId="26" pageNumber="27" pagination="95 - 103" refId="ref124712" refString="Howden H. F. 1976. New species in the genera Bdelyropsis, Cryptocanthon and Drepanocerus (Coleoptera, Scarabaeidae). Proceedings of the Entomological Society of Washington 78: 95 - 103. Available from https: // www. biodiversitylibrary. org / page / 16252570 [accessed 16 Feb. 2024]." type="journal article" year="1976">Howden 1976: 95</bibRefCitation>
(
<collectingCountry id="EDEE87E03040527EECA5FD0D753D03FB" box="[434,561,682,708]" name="Venezuela" pageId="26" pageNumber="27">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923040527EEF56FD0D759B03FB" box="[577,663,682,708]" country="Venezuela" name="Aragua" pageId="26" pageNumber="27">Aragua</collectingRegion>
[actually
<collectingRegion id="573D09923040527EEE0DFD0D749803FB" box="[794,916,682,708]" country="Venezuela" name="Carabobo" pageId="26" pageNumber="27">Carabobo</collectingRegion>
]: Cerro el Cafe). —
<bibRefCitation id="F168BA813040527EE9B5FD0D727A03FA" author="Krajcik M." box="[1186,1398,682,709]" pageId="26" pageNumber="27" pagination="1 - 278" refId="ref126632" refString="Krajcik M. 2012. Checklist of the World Scarabaeoidea. Animma. X Supplement 5: 1 - 278." type="journal article" year="2012">Krajcik 2012: 52</bibRefCitation>
(
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). —
<bibRefCitation id="F168BA813040527EEC6BFD6A75CB03D8" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[380,711,717,744]" pageId="26" pageNumber="27" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B623040527EEF18FD69754F03D8" box="[527,579,717,743]" italics="true" pageId="26" pageNumber="27">et al</emphasis>
. 2013: 108
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:
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: Hacienda Guáquira).
</paragraph>
</subSubSection>
</treatment>
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