treatments-xml/data/1D/50/76/1D507666303A5205EF5CF90C76A305EF.xml

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<document id="F01D601DE41EEC293A59C6C7B8FFC885" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D507666303A5205EF5CF90C76A305EF" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius fallax" docType="treatment" docVersion="2" lastPageNumber="98" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="97" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:titleInfo id="4CC233003DED431353F9A999586AE1F3">
<mods:title id="3F9D3C172F1F0CD741F5822B28E9EB95">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
</mods:titleInfo>
<mods:name id="5A7B964ADA53190238C10CFE01B3B5BC" type="personal">
<mods:role id="44A519DCE14F20FEB073EC9013AA1FB4">
<mods:roleTerm id="71AA6E97294645DAB8DA4F30799FDDEB">Author</mods:roleTerm>
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<mods:namePart id="CC50C1226FB4012542B9E378AEC9D2EF">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="B97578F09B9EAFF7FC8845542D8F10BE">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="CD30B763866FC3082A0834450DB6CC17" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="B88BA35E511EA140F5772764EF46DFF4" type="personal">
<mods:role id="73A271077ADED6AB5801C4962F564580">
<mods:roleTerm id="60AE1F8B7C27CD56011F00793486375B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5B0D6E384BC5C0EBF0AD204C3C581990">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="E51F6501A542B689FAF73F540158808B" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="71158A3021E9EA758B90EDD0E4F9F1E2">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="5C2F797E62D6E99A0AF17DAD0929C203" type="email">mcupello@hotmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="F70032F01FD499140242178F6F140211" type="personal">
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<mods:roleTerm id="0E37907EE6625581057211DEEB35479B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="129AF0BC5A097C44DBB9792BF8F8EFE5">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="9EB831028654BAC8371A8FF11401C78A" type="LSID">2BC093C2-3E2B-466B-B31C-9F3D9FA844BC</mods:nameIdentifier>
<mods:affiliation id="BCB40F4B6566DBC9D1C230C28DF94E97">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="FC793AC94C0318287EEBB90614ECC2DC" type="email">vazdemello@gmail.com</mods:nameIdentifier>
</mods:name>
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<mods:title id="0C7E395A088A9205C644752CA4CA5D43">European Journal of Taxonomy</mods:title>
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<mods:date id="68735C0E1C35FA1DFC548F1AE5EF3496">2024</mods:date>
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<treatment id="1D507666303A5205EF5CF90C76A305EF" ID-DOI="http://doi.org/10.5281/zenodo.13945760" ID-Zenodo-Dep="13945760" LSID="urn:lsid:plazi:treatment:1D507666303A5205EF5CF90C76A305EF" httpUri="http://treatment.plazi.org/id/1D507666303A5205EF5CF90C76A305EF" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97">
<subSubSection id="DDE394FB303A5204EF5CF90C74E407FA" box="[587,1000,1707,1734]" pageId="96" pageNumber="97" type="nomenclature">
<paragraph id="9546C770303A5204EF5CF90C74E407FA" blockId="96.[587,1000,1707,1768]" box="[587,1000,1707,1734]" pageId="96" pageNumber="97">
<heading id="CE0E701C303A5204EF5CF90C74E407FA" box="[587,1000,1707,1734]" centered="true" fontSize="11" level="2" pageId="96" pageNumber="97" reason="2">
<taxonomicName id="52F9BCF3303A5204EF5CF90C74E407FA" ID-CoL="35NBL" authority="(Harold, 1880)" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[587,1000,1707,1734]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="A78D1B62303A5204EF5CF90C743F07F9" bold="true" box="[587,819,1707,1734]" italics="true" pageId="96" pageNumber="97">Dichotomius fallax</emphasis>
(
<bibRefCitation id="F168BA81303A5204EE55F90C74EC07FA" author="Harold E." box="[834,992,1707,1733]" pageId="96" pageNumber="97" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold, 1880</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB303A5204EFF1F969744307D7" box="[742,847,1742,1768]" pageId="96" pageNumber="97" type="description">
<paragraph id="9546C770303A5204EFF1F969744307D7" blockId="96.[587,1000,1707,1768]" box="[742,847,1742,1768]" pageId="96" pageNumber="97">
<figureCitation id="0DC2DBF5303A5204EFF1F969744307D7" box="[742,847,1742,1768]" captionStart="Fig" captionStartId="98.[189,232,1928,1954]" captionTargetBox="[202,1346,295,1830]" captionTargetId="figure-18@98.[189,1398,281,1866]" captionTargetPageId="98" captionText="Fig. 39. Species distribution. A. Dichotomius fallax (Harold, 1880). B. Dichotomius henripittieri Montoya-Molina &amp; Vaz-de-Mello, 2021. C. Male of D. henripittieri. D. Female of D. henripittieri. Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945573" httpUri="https://zenodo.org/record/13945573/files/figure.png" pageId="96" pageNumber="97">Fig. 39A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB303A5205EDAAF8A8723800B3" lastPageId="97" lastPageNumber="98" pageId="96" pageNumber="97" type="reference_group">
<paragraph id="9546C770303A5204EDAAF8A875520650" blockId="96.[189,1399,1807,1903]" pageId="96" pageNumber="97">
<treatmentCitationGroup id="B5E9E05E303A5204EDAAF8A875520650" pageId="96" pageNumber="97">
<treatmentCitation id="1458E161303A5204EDAAF8A8755F0616" author="Harold E." box="[189,595,1807,1833]" page="26" pageId="96" pageNumber="97" year="1880">
<taxonomicName id="52F9BCF3303A5204EDAAF8A8755F0616" ID-CoL="4HZTX" authority="Harold, 1880: 26" authorityName="Harold" authorityPageNumber="26" authorityYear="1880" box="[189,595,1807,1833]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="A78D1B62303A5204EDAAF8A876660616" box="[189,362,1807,1833]" italics="true" pageId="96" pageNumber="97">Pinotus fallax</emphasis>
<bibRefCitation id="F168BA81303A5204EC6DF8A8755F0616" author="Harold E." box="[378,595,1807,1833]" pageId="96" pageNumber="97" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold, 1880: 26</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E0303A5204E920F8A873A30616" box="[1079,1199,1807,1833]" name="Colombia" pageId="96" pageNumber="97">Colombia</collectingCountry>
:
<collectingRegion id="573D0992303A5204E9D4F8A8727F0616" box="[1219,1395,1807,1833]" country="Colombia" name="Cundinamarca" pageId="96" pageNumber="97">Cundinamarca</collectingRegion>
: Tequendama: La Mesa; and
<collectingCountry id="EDEE87E0303A5204EF5EF89675CA0674" box="[585,710,1841,1867]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992303A5204EFC1F89674990674" box="[726,917,1841,1867]" country="Venezuela" name="Distrito Capital" pageId="96" pageNumber="97">Distrito Capital</collectingRegion>
: Caracas. Name-bearing type:
<typeStatus id="4A4279D2303A5204E807F895727B0673" box="[1296,1399,1842,1868]" pageId="96" pageNumber="97" type="syntype">syntypes</typeStatus>
(MNHN), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770303A5204EDAAF831748C0690" blockId="96.[189,896,1941,1968]" box="[189,896,1941,1968]" pageId="96" pageNumber="97">
<treatmentCitationGroup id="B5E9E05E303A5204EDAAF831748C0690" box="[189,896,1941,1968]" pageId="96" pageNumber="97">
<taxonomicName id="52F9BCF3303A5204EDAAF831751F0690" ID-CoL="4HZTX" authorityName="Harold" authorityYear="1880" box="[189,531,1941,1968]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="96" pageNumber="97" phylum="Arthropoda" rank="species" species="fallax" subGenus="Selenocopris">
<emphasis id="A78D1B62303A5204EDAAF831761B0690" box="[189,279,1942,1967]" italics="true" pageId="96" pageNumber="97">Pinotus</emphasis>
(
<emphasis id="A78D1B62303A5204EC3FF83276CC0690" box="[296,448,1941,1967]" italics="true" pageId="96" pageNumber="97">Selenocopris</emphasis>
)
<emphasis id="A78D1B62303A5204ECC7F832751F0690" box="[464,531,1941,1967]" italics="true" pageId="96" pageNumber="97">fallax</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161303A5204EF27F832743F0690" author="Luederwaldt H." box="[560,819,1941,1968]" page="64" pageId="96" pageNumber="97" year="1929">
<bibRefCitation id="F168BA81303A5204EF27F832743F0690" author="Luederwaldt H." box="[560,819,1941,1968]" pageId="96" pageNumber="97" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 64</bibRefCitation>
</treatmentCitation>
(key).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770303B5205EDAAFEAD73E70079" blockId="97.[189,1399,265,397]" pageId="97" pageNumber="98">
<treatmentCitationGroup id="B5E9E05E303B5205EDAAFEAD73E70079" pageId="97" pageNumber="98">
<emphasis id="A78D1B62303B5205EDAAFEAD7674001C" box="[189,376,265,291]" italics="true" pageId="97" pageNumber="98">
<taxonomicName id="52F9BCF3303B5205EDAAFEAD766D001C" ID-CoL="4HZTX" authorityName="Harold" authorityYear="1880" box="[189,353,265,291]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="fallax">Pinotus fallax</taxonomicName>
</emphasis>
<treatmentCitation id="1458E161303B5205EC97FEAE754C001C" author="Gillet J. J. E." box="[384,576,265,291]" page="60" pageId="97" pageNumber="98" year="1911">
<bibRefCitation id="F168BA81303B5205EC97FEAE754C001C" author="Gillet J. J. E." box="[384,576,265,291]" pageId="97" pageNumber="98" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 60</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E161303B5205EE17FEAE74E3001C" author="Gacharna C." box="[768,1007,265,292]" page="222" pageId="97" pageNumber="98" year="1951">
<bibRefCitation id="F168BA81303B5205EE17FEAE74E3001C" author="Gacharna C." box="[768,1007,265,292]" pageId="97" pageNumber="98" pagination="220 - 229" refId="ref119803" refString="Gacharna C. 1951. Catalogo de los coleopteros colombianos. Revista de la Academia Colombiana de Ciencias Fisicas y Naturales 8: 220 - 229." type="journal article" year="1951">Gacharná 1951: 222</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E0303B5205E941FEAE73C3001C" box="[1110,1231,265,291]" name="Colombia" pageId="97" pageNumber="98">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161303B5205E81FFEAE766D0079" author="Balthasar V." page="349" pageId="97" pageNumber="98" year="1941">
<bibRefCitation id="F168BA81303B5205E81FFEAE766D0079" author="Balthasar V." pageId="97" pageNumber="98" pagination="337 - 358" refId="ref110399" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 1: 337 - 358." type="book chapter" year="1941">Balthasar 1941: 349</bibRefCitation>
</treatmentCitation>
(list, characteristics); 1951: 334 (list). —
<treatmentCitation id="1458E161303B5205EE59FE8B736C0079" author="Blackwelder R. E." box="[846,1120,300,326]" page="207" pageId="97" pageNumber="98" year="1944">
<bibRefCitation id="F168BA81303B5205EE59FE8B736C0079" author="Blackwelder R. E." box="[846,1120,300,326]" pageId="97" pageNumber="98" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
</treatmentCitation>
(checklist).
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C770303B5205EDAAFEE8723800B3" blockId="97.[189,1399,265,397]" pageId="97" pageNumber="98">
<treatmentCitationGroup id="B5E9E05E303B5205EDAAFEE8723800B3" pageId="97" pageNumber="98">
<taxonomicName id="52F9BCF3303B5205EDAAFEE876920056" ID-CoL="35NBL" baseAuthorityName="Harold" baseAuthorityYear="1880" box="[189,414,335,361]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="97" pageNumber="98" phylum="Arthropoda" rank="species" species="fallax">
<emphasis id="A78D1B62303B5205EDAAFEE876920056" box="[189,414,335,361]" italics="true" pageId="97" pageNumber="98">Dichotomius fallax</emphasis>
</taxonomicName>
<treatmentCitation id="1458E161303B5205ECA8FEF775610055" author="Roze J. A." box="[447,621,335,362]" page="44" pageId="97" pageNumber="98" year="1955">
<bibRefCitation id="F168BA81303B5205ECA8FEF775610055" author="Roze J. A." box="[447,621,335,362]" pageId="97" pageNumber="98" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
</treatmentCitation>
(cited, checklist for
<collectingCountry id="EDEE87E0303B5205EE74FEE874ED0056" box="[867,993,335,361]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E161303B5205E90DFEE8727A0055" author="Vulcano M. A. &amp; Pereira F. S." box="[1050,1398,335,362]" page="587" pageId="97" pageNumber="98" year="1967">
<bibRefCitation id="F168BA81303B5205E90DFEE8727A0055" author="Vulcano M. A. &amp; Pereira F. S." box="[1050,1398,335,362]" pageId="97" pageNumber="98" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 587</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E161303B5205EC4AFED5757900B2" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[349,629,370,397]" page="138" pageId="97" pageNumber="98" year="2001">
<bibRefCitation id="F168BA81303B5205EC4AFED5757900B2" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[349,629,370,397]" pageId="97" pageNumber="98" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B62303B5205ECABFED476FB00B3" box="[444,503,370,396]" italics="true" pageId="97" pageNumber="98">et al.</emphasis>
2001: 138
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E0303B5205EE08FED5749400B3" box="[799,920,370,396]" name="Colombia" pageId="97" pageNumber="98">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E161303B5205EEC7FED473E200B2" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[976,1262,370,397]" page="1294" pageId="97" pageNumber="98" year="2008">
<bibRefCitation id="F168BA81303B5205EEC7FED473E200B2" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[976,1262,370,397]" pageId="97" pageNumber="98" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B62303B5205E930FED4736E00B3" box="[1063,1122,370,396]" italics="true" pageId="97" pageNumber="98">et al.</emphasis>
2008: 1294
</bibRefCitation>
</treatmentCitation>
(list).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB303B5205EDAAFE1376AD0392" pageId="97" pageNumber="98" type="materials_examined">
<paragraph id="9546C770303B5205EDAAFE1376A600F1" blockId="97.[189,426,436,462]" box="[189,426,436,462]" pageId="97" pageNumber="98">
<heading id="CE0E701C303B5205EDAAFE1376A600F1" bold="true" box="[189,426,436,462]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="A78D1B62303B5205EDAAFE1376A600F1" bold="true" box="[189,426,436,462]" pageId="97" pageNumber="98">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770303B5205EDAAFE4476AD0392" blockId="97.[189,1399,482,685]" pageId="97" pageNumber="98">
<materialsCitation id="2591CD2D303B5205EDAAFE44761C031F" collectingDate="2006-08-05" collectorName="El Manteco" country="Venezuela" location="Bolivar" pageId="97" pageNumber="98" specimenCount="1" stateProvince="Bolivar">
<collectingCountry id="EDEE87E0303B5205EDAAFE44766700C2" box="[189,363,483,509]" name="Venezuela" pageId="97" pageNumber="98">VENEZUELA</collectingCountry>
<emphasis id="A78D1B62303B5205EC9DFE4576EA00C2" bold="true" box="[394,486,482,509]" pageId="97" pageNumber="98">
<collectingRegion id="573D0992303B5205EC9DFE4576EA00C2" box="[394,486,482,509]" country="Venezuela" name="Bolivar" pageId="97" pageNumber="98">Bolívar</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9303B5205EF14FE44755500C2" box="[515,601,483,509]" count="1" pageId="97" pageNumber="98" type="generic">1 spec.</specimenCount>
;
<collectorName id="380CA2A6303B5205EF70FE4575F800C2" box="[615,756,482,509]" pageId="97" pageNumber="98">El Manteco</collectorName>
;
<date id="E147E1B0303B5205EE15FE45749400C3" box="[770,920,482,509]" pageId="97" pageNumber="98" value="2006-08-05">
<collectingDate id="F1031858303B5205EE15FE45749400C3" box="[770,920,482,509]" pageId="97" pageNumber="98" value="2006-08-05">5 Aug. 2006</collectingDate>
</date>
; curso NM2006 leg.; heces, 24h, 08:00; CEMT
</materialsCitation>
<materialsCitation id="2591CD2D303B5205EC3EFDA1720D031F" box="[297,1281,518,544]" collectingDate="1987-05-23" collectingDateMax="1987-05-30" collectingDateMin="1987-05-23" collectorName="El Dorado" country="Venezuela" elevation="190" location="Rio Yuruani" municipality="Campo Minero" pageId="97" pageNumber="98" specimenCount="1" stateProvince="Bolivar">
2 specs;
<collectingMunicipality id="75225D0A303B5205EC98FDA17549031F" box="[399,581,518,544]" pageId="97" pageNumber="98">Campo Minero</collectingMunicipality>
,
<location id="902691AB303B5205EF45FDA175E8031F" LSID="urn:lsid:plazi:treatment:1D507666303A5205EF5CF90C76A305EF:902691AB303B5205EF45FDA175E8031F" box="[594,740,518,544]" country="Venezuela" municipality="Campo Minero" name="Rio Yuruani" pageId="97" pageNumber="98" stateProvince="Bolivar">Río Yuruani</location>
,
<collectorName id="380CA2A6303B5205EFE7FDA17462031F" box="[752,878,518,544]" pageId="97" pageNumber="98">El Dorado</collectorName>
;
<quantity id="52016A95303B5205EE6CFDA174C5031F" box="[891,969,518,544]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="97" pageNumber="98" unit="m" value="190.0">
<elevation id="1ED42043303B5205EE6CFDA174C5031F" box="[891,969,518,544]" metricMagnitude="2" metricUnit="m" metricValue="1.9" pageId="97" pageNumber="98" unit="m" value="190.0">190 m</elevation>
</quantity>
;
<date id="E147E1B0303B5205EEC1FDA173AF031F" box="[982,1187,518,544]" pageId="97" pageNumber="98" value="1987-05-23" valueMax="1987-05-30" valueMin="1987-05-23">
<collectingDate id="F1031858303B5205EEC1FDA173AF031F" box="[982,1187,518,544]" pageId="97" pageNumber="98" value="1987-05-23" valueMax="1987-05-30" valueMin="1987-05-23">2330 May 1987</collectingDate>
</date>
; CEMT
</materialsCitation>
<materialsCitation id="2591CD2D303B5205E80BFDA17305037C" collectingDate="1983-04-12" collectorName="C. Bordon" country="Venezuela" location="Las Pinas" pageId="97" pageNumber="98" specimenCount="1" stateProvince="Bolivar">
<specimenCount id="83FF0CF9303B5205E80BFDA1727E031F" box="[1308,1394,518,544]" count="1" pageId="97" pageNumber="98" type="generic">1 spec.</specimenCount>
;
<location id="902691AB303B5205EDAAFD8E763F037C" LSID="urn:lsid:plazi:treatment:1D507666303A5205EF5CF90C76A305EF:902691AB303B5205EDAAFD8E763F037C" box="[189,307,553,579]" country="Venezuela" name="Las Pinas" pageId="97" pageNumber="98" stateProvince="Bolivar">Las Piñas</location>
,
<locationDeviation id="84F4DE52303B5205EC56FD8E753B037C" box="[321,567,553,579]" location="Paragua" pageId="97" pageNumber="98">
<quantity id="52016A95303B5205EC56FD8E7681037C" box="[321,397,553,579]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="97" pageNumber="98" unit="km" value="30.0">30 km</quantity>
E of Paragua
</locationDeviation>
;
<date id="E147E1B0303B5205EF52FD8E75E9037C" box="[581,741,553,579]" pageId="97" pageNumber="98" value="1983-04-12">
<collectingDate id="F1031858303B5205EF52FD8E75E9037C" box="[581,741,553,579]" pageId="97" pageNumber="98" value="1983-04-12">12 Apr. 1983</collectingDate>
</date>
;
<collectorName id="380CA2A6303B5205EFE4FD8E747C037C" box="[755,880,553,579]" pageId="97" pageNumber="98">C. Bordon</collectorName>
leg.; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D303B5205E93AFD8E76CF03B6" collectingDate="2009-07-19" collectorName="M. Assmussen &amp; P Colmenares &amp; H. Martinez" country="Venezuela" elevation="483" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" municipality="Bolivar" pageId="97" pageNumber="98" specimenCount="1" stateProvince="Yaracuy">
<emphasis id="A78D1B62303B5205E93AFD8E7399037C" bold="true" box="[1069,1173,553,579]" pageId="97" pageNumber="98">
<collectingRegion id="573D0992303B5205E93AFD8E7399037C" box="[1069,1173,553,579]" country="Venezuela" name="Yaracuy" pageId="97" pageNumber="98">Yaracuy</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9303B5205E9A4FD8E7206037C" box="[1203,1290,553,579]" count="1" pageId="97" pageNumber="98" type="generic">1 spec.</specimenCount>
;
<collectingMunicipality id="75225D0A303B5205E80FFD8E727E037C" box="[1304,1394,553,579]" pageId="97" pageNumber="98">Bolívar</collectingMunicipality>
,
<location id="902691AB303B5205EDAAFDEB77F70359" LSID="urn:lsid:plazi:treatment:1D507666303A5205EF5CF90C76A305EF:902691AB303B5205EDAAFDEB77F70359" box="[189,251,588,614]" country="Venezuela" latitude="10.0" longLatPrecision="21" longitude="-68.0" municipality="Bolivar" name="Aroa" pageId="97" pageNumber="98" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B7303B5205EC1FFDEC768D0358" box="[264,385,587,615]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="97" pageNumber="98" precision="15" seconds="0" value="10.0">10°00″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B7303B5205EC9AFDEC75070358" box="[397,523,587,615]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="97" pageNumber="98" precision="15" seconds="0" value="-68.0">68°00″ W</geoCoordinate>
;
<quantity id="52016A95303B5205EF0FFDEB756A0358" box="[536,614,588,615]" metricMagnitude="2" metricUnit="m" metricValue="4.83" pageId="97" pageNumber="98" unit="m" value="483.0">
<elevation id="1ED42043303B5205EF0FFDEB756A0358" box="[536,614,588,615]" metricMagnitude="2" metricUnit="m" metricValue="4.83" pageId="97" pageNumber="98" unit="m" value="483.0">483 m</elevation>
</quantity>
;
<date id="E147E1B0303B5205EF64FDEB74040359" box="[627,776,588,614]" pageId="97" pageNumber="98" value="2009-07-19">
<collectingDate id="F1031858303B5205EF64FDEB74040359" box="[627,776,588,614]" pageId="97" pageNumber="98" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A6303B5205EE03FDEA74CF0359" box="[788,963,588,615]" pageId="97" pageNumber="98">M. Assmüssen</collectorName>
,
<collectorName id="380CA2A6303B5205EED8FDEA737F0359" box="[975,1139,588,615]" pageId="97" pageNumber="98">P Colmenares</collectorName>
and
<collectorName id="380CA2A6303B5205E9BAFDEA72370359" box="[1197,1339,588,615]" pageId="97" pageNumber="98">H. Martínez</collectorName>
leg.; human faeces; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D303B5205ECFFFDC876910392" collectingDate="2006-07-18" country="Venezuela" location="Rosario de Perija" pageId="97" pageNumber="98" specimenCount="1" stateProvince="Zulia">
<emphasis id="A78D1B62303B5205ECFFFDC8752703B6" bold="true" box="[488,555,623,649]" pageId="97" pageNumber="98">
<collectingRegion id="573D0992303B5205ECFFFDC8752703B6" box="[488,555,623,649]" country="Venezuela" name="Zulia" pageId="97" pageNumber="98">Zulia</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF9303B5205EF50FDC8759103B5" box="[583,669,623,650]" count="1" pageId="97" pageNumber="98" type="generic">1 spec.</specimenCount>
;
<location id="902691AB303B5205EFBCFDC8747003B6" LSID="urn:lsid:plazi:treatment:1D507666303A5205EF5CF90C76A305EF:902691AB303B5205EFBCFDC8747003B6" box="[683,892,623,649]" country="Venezuela" name="Rosario de Perija" pageId="97" pageNumber="98" stateProvince="Zulia">Rosario de Perijá</location>
;
<date id="E147E1B0303B5205EE9EFDC8732C03B6" box="[905,1056,623,649]" pageId="97" pageNumber="98" value="2006-07-18">
<collectingDate id="F1031858303B5205EE9EFDC8732C03B6" box="[905,1056,623,649]" pageId="97" pageNumber="98" value="2006-07-18">18 Jul. 2006</collectingDate>
</date>
; curso NM2006 leg.; faeces, 23h, 13:09; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB303B5205EDAAFD7376A305EF" pageId="97" pageNumber="98" type="distribution">
<paragraph id="9546C770303B5205EDAAFD73765A03D1" blockId="97.[189,342,724,750]" box="[189,342,724,750]" pageId="97" pageNumber="98">
<heading id="CE0E701C303B5205EDAAFD73765A03D1" bold="true" box="[189,342,724,750]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="A78D1B62303B5205EDAAFD73765A03D1" bold="true" box="[189,342,724,750]" pageId="97" pageNumber="98">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770303B5205EDAAFCA57683027F" blockId="97.[189,1398,770,832]" pageId="97" pageNumber="98">
<collectingCountry id="EDEE87E0303B5205EDAAFCA576380223" box="[189,308,770,796]" name="Colombia" pageId="97" pageNumber="98">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E0303B5205EC2AFCA576B50223" box="[317,441,770,796]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E0303B5205ECD5FCA575BE0223" box="[450,690,770,796]" name="Trinidad and Tobago" pageId="97" pageNumber="98">Trinidad and Tobago</collectingCountry>
(
<emphasis id="A78D1B62303B5205EFD6FCA474BF0222" bold="true" box="[705,947,771,797]" pageId="97" pageNumber="98">new country record</emphasis>
, CEMT),
<collectingCountry id="EDEE87E0303B5205E930FCA573630223" box="[1063,1135,770,796]" name="Brazil" pageId="97" pageNumber="98">Brazil</collectingCountry>
(
<emphasis id="A78D1B62303B5205E969FCA4727C0222" bold="true" box="[1150,1392,771,797]" pageId="97" pageNumber="98">new country record</emphasis>
, CEMT) and
<collectingCountry id="EDEE87E0303B5205EC46FC817687027F" box="[337,395,806,832]" name="Peru" pageId="97" pageNumber="98">Peru</collectingCountry>
.
</paragraph>
<paragraph id="9546C770303B5205EDAAFCC076E102BE" blockId="97.[189,493,871,897]" box="[189,493,871,897]" pageId="97" pageNumber="98">
<heading id="CE0E701C303B5205EDAAFCC076E102BE" bold="true" box="[189,493,871,897]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="A78D1B62303B5205EDAAFCC076E102BE" bold="true" box="[189,493,871,897]" pageId="97" pageNumber="98">
Subregions of
<collectingCountry id="EDEE87E0303B5205EC78FCC076E102BE" box="[367,493,871,897]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770303B5205EDAAFC31761C02C8" blockId="97.[189,1400,918,1015]" pageId="97" pageNumber="98">
Maracaibo Depression, System of hills and low piedmont mountains of the Guiana Shield, System of low mountains and hills Imataca-Cuyuní del Escudo Northeast Guiana, and Central
<collectingRegion id="573D0992303B5205E9AEFC1E73F002EC" box="[1209,1276,953,979]" country="New Zealand" name="West Coast" pageId="97" pageNumber="98">Coast</collectingRegion>
Mountain Range.
</paragraph>
<paragraph id="9546C770303B5205EDAAFBBA76A80507" blockId="97.[189,420,1053,1080]" box="[189,420,1053,1080]" pageId="97" pageNumber="98">
<heading id="CE0E701C303B5205EDAAFBBA76A80507" bold="true" box="[189,420,1053,1080]" fontSize="11" level="3" pageId="97" pageNumber="98" reason="3">
<emphasis id="A78D1B62303B5205EDAAFBBA76A80507" bold="true" box="[189,420,1053,1080]" pageId="97" pageNumber="98">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770303B5205EDAAFBEB76A305EF" blockId="97.[189,1399,1100,1232]" pageId="97" pageNumber="98">
<bibRefCitation id="F168BA81303B5205EDAAFBEB768A0559" author="Harold E." box="[189,390,1100,1126]" pageId="97" pageNumber="98" pagination="13 - 46" refId="ref123840" refString="Harold E. 1880. Verzeichniss der von Dr. Steinheil in Neu-Granada gesammelten Coprophagen Lamellicornien. Entomologische Zeitung 41: 13 - 46." type="journal article" year="1880">Harold 1880: 26</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205EC8CFBEB75160559" box="[411,538,1100,1126]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
: Caracas). —
<bibRefCitation id="F168BA81303B5205EFDAFBEB74990559" author="Gillet J. J. E." box="[717,917,1100,1126]" pageId="97" pageNumber="98" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 60</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205EEBEFBEB73250559" box="[937,1065,1100,1126]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81303B5205E97DFBEB727A0559" author="Luederwaldt H." box="[1130,1398,1100,1126]" pageId="97" pageNumber="98" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 64</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205EDD2FBC8764905B6" box="[197,325,1135,1161]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81303B5205EC94FBC8757A05B6" author="Balthasar V." box="[387,630,1135,1161]" pageId="97" pageNumber="98" pagination="337 - 358" refId="ref110399" refString="Balthasar V. 1941. Scarabaeidae Laparostictae (Col.). Beitrage Fauna Perus 1: 337 - 358." type="book chapter" year="1941">Balthasar 1941: 349</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205EF9FFBC8740405B6" box="[648,776,1135,1161]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
); 1951: 334 (
<collectingCountry id="EDEE87E0303B5205EEBCFBC8732605B6" box="[939,1066,1135,1161]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
).—
<bibRefCitation id="F168BA81303B5205E949FBC8727A05B6" author="Blackwelder R. E." box="[1118,1398,1135,1161]" pageId="97" pageNumber="98" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205EDD2FB3576490593" box="[197,325,1170,1196]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81303B5205EC95FB3575EB0592" author="Vulcano M. A. &amp; Pereira F. S." box="[386,743,1170,1197]" pageId="97" pageNumber="98" pagination="533 - 603" refId="ref141365" refString="Vulcano M. A. &amp; Pereira F. S. 1967. Sinopse dos Passalidae e Scarabaeidae s. str. da regiao amazonica (Insecta, Coleoptera). Atas do Simposio sobre a Biota Amazonica 5: 533 - 603." type="journal article" year="1967">Vulcano &amp; Pereira 1967: 587</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205EFEEFB3574750593" box="[761,889,1170,1196]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA81303B5205EEA0FB3473EE0592" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[951,1250,1170,1197]" pageId="97" pageNumber="98" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B62303B5205E906FB3473440593" box="[1041,1096,1170,1196]" italics="true" pageId="97" pageNumber="98">et al</emphasis>
. 2008: 1294
</bibRefCitation>
(
<collectingCountry id="EDEE87E0303B5205E9E3FB35727F0593" box="[1268,1395,1170,1196]" name="Venezuela" pageId="97" pageNumber="98">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992303B5205EDAAFB11761605EF" box="[189,282,1206,1232]" country="Venezuela" name="Bolivar" pageId="97" pageNumber="98">Bolívar</collectingRegion>
: Lago Guri).
</paragraph>
</subSubSection>
</treatment>
</document>