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<document id="7CD9EC6973E9A8721CC395984BFE1296" ID-DOI="10.5852/ejt.2024.959.2677" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D5076663004523AEF3CFEAF74FB0446" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius coenosus" docType="treatment" docVersion="2" lastPageNumber="95" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="95" updateTime="1729169574746" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="4FE65A9EB69BDD55BDAB7C097B11BE29">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:namePart id="54C3D312649F004C64AAB0D4CC90A3DE">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:namePart id="12E7233E6D4EFB0EB727884F08627FE4">Cupello, Mario</mods:namePart>
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<mods:affiliation id="0B2555B1F55CCB899293663E45322D0C">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
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<emphasis id="A78D1B623004523AEF3CFEAF7430001C" bold="true" box="[555,828,264,291]" italics="true" pageId="94" pageNumber="95">Dichotomius coenosus</emphasis>
(
<bibRefCitation id="F168BA813004523AEE5CFEAE730C001C" author="Erichson W. F." box="[843,1024,265,291]" pageId="94" pageNumber="95" pagination="553 - 1260" refId="ref117915" refString="Erichson W. F. 1848. Insecten. In: Schomburgk R. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil: 553 - 1260. J. J. Weber, Leipzig. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson, 1848</bibRefCitation>
)
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</paragraph>
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<figureCitation id="0DC2DBF53004523AEFF1FE8A74410078" box="[742,845,301,327]" captionStart="Fig" captionStartId="93.[189,232,1963,1989]" captionTargetBox="[279,1309,278,1889]" captionTargetId="figure-12@93.[268,1319,265,1907]" captionTargetPageId="93" captionText="Fig. 37. Species distribution.A. Dichotomius boreus (Olivier, 1789).B. Dichotomius coenosus (Erichson, 1848). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945567" httpUri="https://zenodo.org/record/13945567/files/figure.png" pageId="94" pageNumber="95">Fig. 37B</figureCitation>
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<taxonomicName id="52F9BCF33004523AEDAAFED776EB00B5" authorityName="Erichson" authorityYear="1849" box="[189,487,368,394]" class="Insecta" family="Scarabaeidae" genus="Copris" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="coenosa" subGenus="Pinotus">
<emphasis id="A78D1B623004523AEDAAFED7760100B5" box="[189,269,368,394]" italics="true" pageId="94" pageNumber="95">Copris</emphasis>
(
<emphasis id="A78D1B623004523AEC0AFED6767B00B5" box="[285,375,369,394]" italics="true" pageId="94" pageNumber="95">Pinotus</emphasis>
)
<emphasis id="A78D1B623004523AEC90FED676EB00B5" box="[391,487,369,394]" italics="true" pageId="94" pageNumber="95">coenosa</emphasis>
</taxonomicName>
[sic]
<treatmentCitation id="1458E1613004523AEF3FFED7740000B4" author="Erichson W. F." box="[552,780,368,395]" page="564" pageId="94" pageNumber="95" year="1848">
<bibRefCitation id="F168BA813004523AEF3FFED7740000B4" author="Erichson W. F." box="[552,780,368,395]" pageId="94" pageNumber="95" pagination="553 - 1260" refId="ref117915" refString="Erichson W. F. 1848. Insecten. In: Schomburgk R. Reisen in Britisch-Guiana in den Jahren 1840 - 1844. Dritter Theil: 553 - 1260. J. J. Weber, Leipzig. https: // doi. org / 10.5962 / bhl. title. 109982" type="book chapter" year="1848">Erichson 1848: 564</bibRefCitation>
</treatmentCitation>
(original description). Type locality:
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. Name-bearing type:
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(MFNB), examined by FZVM.
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<paragraph id="9546C7703004523AEDAAFE717650032C" blockId="94.[189,1399,469,707]" pageId="94" pageNumber="95">
<treatmentCitationGroup id="B5E9E05E3004523AEDAAFE717650032C" pageId="94" pageNumber="95">
<emphasis id="A78D1B623004523AEDAAFE71769300D0" box="[189,415,470,495]" italics="true" pageId="94" pageNumber="95">
<taxonomicName id="52F9BCF33004523AEDAAFE71768500D0" ID-CoL="9JZYK" authorityName="Erichson" authorityYear="1848" box="[189,393,470,495]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="coenosus">Pinotus coenosus</taxonomicName>
</emphasis>
<treatmentCitation id="1458E1613004523AECB3FE72756E00D0" author="Gillet J. J. E." box="[420,610,469,495]" page="60" pageId="94" pageNumber="95" year="1911">
<bibRefCitation id="F168BA813004523AECB3FE72756E00D0" author="Gillet J. J. E." box="[420,610,469,495]" pageId="94" pageNumber="95" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 60</bibRefCitation>
</treatmentCitation>
(catalogue). —
<treatmentCitation id="1458E1613004523AEE37FE72732D00CF" author="Luederwaldt H." box="[800,1057,469,496]" page="17" pageId="94" pageNumber="95" year="1929">
<bibRefCitation id="F168BA813004523AEE37FE72732D00CF" author="Luederwaldt H." box="[800,1057,469,496]" pageId="94" pageNumber="95" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 17</bibRefCitation>
</treatmentCitation>
(key). —
<treatmentCitation id="1458E1613004523AE98EFE72761A032C" author="Blackwelder R. E." page="207" pageId="94" pageNumber="95" year="1944">
<bibRefCitation id="F168BA813004523AE98EFE72761A032C" author="Blackwelder R. E." pageId="94" pageNumber="95" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
</treatmentCitation>
(list).
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</paragraph>
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<treatmentCitationGroup id="B5E9E05E3004523AEDAAFDBB75AD03FC" pageId="94" pageNumber="95">
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<emphasis id="A78D1B623004523AEDAAFDBB765F0309" box="[189,339,540,566]" italics="true" pageId="94" pageNumber="95">Dichotomius</emphasis>
(
<emphasis id="A78D1B623004523AEC75FDBB76F40309" box="[354,504,540,566]" italics="true" pageId="94" pageNumber="95">Dichotomius</emphasis>
)
<emphasis id="A78D1B623004523AEF1FFDBA75780309" box="[520,628,541,566]" italics="true" pageId="94" pageNumber="95">coenosus</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613004523AEF99FDBB74390309" author="Roze J. A." box="[654,821,540,567]" page="44" pageId="94" pageNumber="95" year="1955">
<bibRefCitation id="F168BA813004523AEF99FDBB74390309" author="Roze J. A." box="[654,821,540,567]" pageId="94" pageNumber="95" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03004523AEECCFDBB73550309" box="[987,1113,540,566]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AE99AFDBB72430309" author="Blanco J." box="[1165,1359,540,566]" page="40" pageId="94" pageNumber="95" year="1988">
<bibRefCitation id="F168BA813004523AE99AFDBB72430309" author="Blanco J." box="[1165,1359,540,566]" pageId="94" pageNumber="95" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 40</bibRefCitation>
</treatmentCitation>
, 41 (catalogue). —
<treatmentCitation id="1458E1613004523AECB6FD9875CC0366" author="Vaz-de-Mello F. Z." box="[417,704,575,601]" page="193" pageId="94" pageNumber="95" year="2000">
<bibRefCitation id="F168BA813004523AECB6FD9875CC0366" author="Vaz-de-Mello F. Z." box="[417,704,575,601]" pageId="94" pageNumber="95" pagination="183 - 195" refId="ref140384" refString="Vaz-de-Mello F. Z. 2000. Estado atual de conhecimentos dos Scarabaeidae s. str. (Coleoptera: Scarabaeoidea) do Brasil. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds) Hacia un Proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 183 - 195. m 3 m: Monografias Tercer Milenio vol. 1, Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Vaz-de-Mello 2000: 193</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03004523AEE7DFD9874BB0366" box="[874,951,575,601]" name="Brazil" pageId="94" pageNumber="95">Brazil</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AEEF9FDE772000366" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[1006,1292,575,602]" page="1294" pageId="94" pageNumber="95" year="2008">
<bibRefCitation id="F168BA813004523AEEF9FDE772000366" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[1006,1292,575,602]" pageId="94" pageNumber="95" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B623004523AE952FDE7738C0366" box="[1093,1152,575,601]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
2008: 1294
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613004523AEDFDFDC575410343" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[234,589,610,636]" page="109" pageId="94" pageNumber="95" year="2013">
<bibRefCitation id="F168BA813004523AEDFDFDC575410343" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[234,589,610,636]" pageId="94" pageNumber="95" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B623004523AEC94FDC476C80343" box="[387,452,610,636]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
2013: 109
</bibRefCitation>
</treatmentCitation>
(list). —
<treatmentCitation id="1458E1613004523AEFC6FDC573E50343" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[721,1257,610,636]" page="29" pageId="94" pageNumber="95" year="2014">
<bibRefCitation id="F168BA813004523AEFC6FDC573E50343" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[721,1257,610,636]" pageId="94" pageNumber="95" refId="ref138148" refString="Sarmiento-Garces R. &amp; Amat-Garcia G. 2014. Escarabajos del Genero Dichotomius Hope 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia, Monografia n ° 4, Universidad Nacional de Colombia, Bogota. https: // doi. org / 10.18257 / raccefyn. 33 (127). 2009.2364" type="book" year="2014">Sarmiento-Garcés &amp; Amat-García 2014: 29</bibRefCitation>
</treatmentCitation>
(diagnosis, distribution for
<collectingCountry id="EDEE87E03004523AECBCFD22752803A0" box="[427,548,645,671]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AEF73FD22749D039F" author="Pardo-Diaz C. &amp; Lopera T. A. &amp; Pena S. A. &amp; Sarmiento-Garces R. &amp; Sanchez-Herrera M. &amp; Salazar C." box="[612,913,645,672]" page="8" pageId="94" pageNumber="95" year="2019">
<bibRefCitation id="F168BA813004523AEF73FD22749D039F" author="Pardo-Diaz C. &amp; Lopera T. A. &amp; Pena S. A. &amp; Sarmiento-Garces R. &amp; Sanchez-Herrera M. &amp; Salazar C." box="[612,913,645,672]" pageId="94" pageNumber="95" refId="ref134659" refString="Pardo-Diaz C., Lopera T. A., Pena S. A., Sarmiento-Garces R., Sanchez-Herrera M. &amp; Salazar C. 2019. Taxonomic reassessment of the genus Dichotomius (Coleoptera: Scarabaeinae) through integrative taxonomy. PeerJ 7: e 7332. https: // doi. org / 10.7717 / peerj. 7332" type="journal volume" year="2019">
Pardo-Díaz
<emphasis id="A78D1B623004523AEFE2FD21743803A0" box="[757,820,645,671]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
2019:8
</bibRefCitation>
</treatmentCitation>
(comments). —
<treatmentCitation id="1458E1613004523AE973FD22765D03FC" ID-DOI="http://doi.org/10.5281/zenodo.3684823" ID-Zenodo-Dep="3684823" author="Hielkema A. J. &amp; Hielkema M. A." httpUri="http://treatment.plazi.org/id/03D7FF1CFFFEFFD8FF48F8813593FAED" page="52" pageId="94" pageNumber="95" year="2019">
<bibRefCitation id="F168BA813004523AE973FD22765D03FC" author="Hielkema A. J. &amp; Hielkema M. A." pageId="94" pageNumber="95" pagination="1 - 306" refId="ref124225" refString="Hielkema A. J. &amp; Hielkema M. A. 2019. An annotated checklist of the Scarabaeoidea (Insecta: Coleoptera) of the Guianas. Insecta Mundi 732 (732): 1 - 306." type="journal article" year="2019">Hielkema &amp; Hielkema 2019: 52</bibRefCitation>
</treatmentCitation>
(catalogue for the Guianas).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3004523AEDAAFD4D73AF02A3" pageId="94" pageNumber="95" type="materials_examined">
<paragraph id="9546C7703004523AEDAAFD4D76A6023B" blockId="94.[189,426,746,772]" box="[189,426,746,772]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AEDAAFD4D76A6023B" bold="true" box="[189,426,746,772]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A78D1B623004523AEDAAFD4D76A6023B" bold="true" box="[189,426,746,772]" pageId="94" pageNumber="95">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703004523AEDAAFCBE73AF02A3" blockId="94.[189,1399,792,925]" pageId="94" pageNumber="95">
<materialsCitation id="2591CD2D3004523AEDAAFCBE737F020C" box="[189,1139,792,819]" collectingDate="1984-05" collectorName="El Vigia &amp; Joffre B." country="Venezuela" location="Merida" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Merida">
<collectingCountry id="EDEE87E03004523AEDAAFCBE7667020C" box="[189,363,793,819]" name="Venezuela" pageId="94" pageNumber="95">VENEZUELA</collectingCountry>
<emphasis id="A78D1B623004523AEC9BFCBF76E5020C" bold="true" box="[396,489,792,819]" pageId="94" pageNumber="95">
<collectingRegion id="573D09923004523AEC9BFCBF76E5020C" box="[396,489,792,819]" country="Venezuela" name="Merida" pageId="94" pageNumber="95">Mérida</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93004523AEF1FFCBE7553020C" box="[520,607,793,819]" count="1" pageId="94" pageNumber="95" type="generic">1 spec.</specimenCount>
;
<collectorName id="380CA2A63004523AEF79FCBE75D8020C" box="[622,724,793,819]" pageId="94" pageNumber="95">El Vigia</collectorName>
;
<date id="E147E1B03004523AEFF4FCBE746D020C" box="[739,865,793,819]" pageId="94" pageNumber="95" value="1984-05">
<collectingDate id="F10318583004523AEFF4FCBE746D020C" box="[739,865,793,819]" pageId="94" pageNumber="95" value="1984-05">May 1984</collectingDate>
</date>
;
<collectorName id="380CA2A63004523AEE67FCBE74D4020C" box="[880,984,793,819]" pageId="94" pageNumber="95">Joffre B.</collectorName>
leg.; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D3004523AE98FFCBF74380269" collectingDate="1986-05" collectorName="Joffre B." country="Venezuela" location="San Pedro del Rio" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Tachira">
<emphasis id="A78D1B623004523AE98FFCBF73F7020C" bold="true" box="[1176,1275,792,819]" pageId="94" pageNumber="95">
<collectingRegion id="573D09923004523AE98FFCBF73F7020C" box="[1176,1275,792,819]" country="Venezuela" name="Tachira" pageId="94" pageNumber="95">Táchira</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93004523AE80DFCBE727E020C" box="[1306,1394,793,819]" count="1" pageId="94" pageNumber="95" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3004523AEDAAFC9B76970269" LSID="urn:lsid:plazi:treatment:1D5076663004523AEF3CFEAF74FB0446:902691AB3004523AEDAAFC9B76970269" box="[189,411,828,854]" country="Venezuela" name="San Pedro del Rio" pageId="94" pageNumber="95" stateProvince="Tachira">San Pedro del Río</location>
;
<date id="E147E1B03004523AECBEFC9B752A0269" box="[425,550,828,854]" pageId="94" pageNumber="95" value="1986-05">
<collectingDate id="F10318583004523AECBEFC9B752A0269" box="[425,550,828,854]" pageId="94" pageNumber="95" value="1986-05">May 1986</collectingDate>
</date>
;
<collectorName id="380CA2A63004523AEF23FC9B75970269" box="[564,667,828,854]" pageId="94" pageNumber="95">Joffre B.</collectorName>
leg.; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D3004523AEE4FFC9B761C02A3" collectingDate="2009-07-19" collectorName="M. Asmussen &amp; P. Colmenares &amp; H. Martinez" country="Venezuela" elevation="464" latitude="10.0" location="Aroa" longLatPrecision="21" longitude="-68.0" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Yaracuy">
<emphasis id="A78D1B623004523AEE4FFC9B74CC0269" bold="true" box="[856,960,828,854]" pageId="94" pageNumber="95">
<collectingRegion id="573D09923004523AEE4FFC9B74CC0269" box="[856,960,828,854]" country="Venezuela" name="Yaracuy" pageId="94" pageNumber="95">Yaracuy</collectingRegion>
</emphasis>
• 2 specs;
<collectingRegion id="573D09923004523AE951FC9B73AC0269" box="[1094,1184,828,854]" country="Venezuela" name="Bolivar" pageId="94" pageNumber="95">Bolívar</collectingRegion>
,
<location id="902691AB3004523AE9BBFC9B73E60269" LSID="urn:lsid:plazi:treatment:1D5076663004523AEF3CFEAF74FB0446:902691AB3004523AE9BBFC9B73E60269" box="[1196,1258,828,854]" country="Venezuela" latitude="10.0" longLatPrecision="21" longitude="-68.0" name="Aroa" pageId="94" pageNumber="95" stateProvince="Yaracuy">Aroa</location>
;
<geoCoordinate id="F0CDA1B73004523AE9EFFC9D727E0269" box="[1272,1394,826,854]" degrees="10" direction="north" minutes="0" orientation="latitude" pageId="94" pageNumber="95" precision="15" seconds="0" value="10.0">10°00″ N</geoCoordinate>
,
<geoCoordinate id="F0CDA1B73004523AEDAAFCFA76360245" box="[189,314,861,890]" degrees="68" direction="west" minutes="0" orientation="longitude" pageId="94" pageNumber="95" precision="15" seconds="0" value="-68.0">68°00″ W</geoCoordinate>
;
<elevation id="1ED420433004523AEC52FCF876C10246" box="[325,461,863,890]" metricMagnitude="2" metricUnit="m" metricValue="4.64" pageId="94" pageNumber="95" unit="m" value="464.0">
<quantity id="52016A953004523AEC52FCF876830245" box="[325,399,863,890]" metricMagnitude="2" metricUnit="m" metricValue="4.64" pageId="94" pageNumber="95" unit="m" value="464.0">464 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03004523AECCDFCF875600246" box="[474,620,863,889]" pageId="94" pageNumber="95" value="2009-07-19">
<collectingDate id="F10318583004523AECCDFCF875600246" box="[474,620,863,889]" pageId="94" pageNumber="95" value="2009-07-19">19 Jul. 2009</collectingDate>
</date>
;
<collectorName id="380CA2A63004523AEF60FCC774150246" box="[631,793,863,890]" pageId="94" pageNumber="95">M. Asmüssen</collectorName>
,
<collectorName id="380CA2A63004523AEE33FCC774C60246" box="[804,970,863,890]" pageId="94" pageNumber="95">P. Colmenares</collectorName>
and
<collectorName id="380CA2A63004523AE915FCC773830246" box="[1026,1167,863,890]" pageId="94" pageNumber="95">H. Martínez</collectorName>
leg.; human faeces; CEMT
</materialsCitation>
.
<materialsCitation id="2591CD2D3004523AEC38FC25739302A3" box="[303,1183,898,925]" collectingDate="2004-07-06" collectorName="D. Garcia" country="Venezuela" elevation="50" location="Cuyagua" pageId="94" pageNumber="95" specimenCount="1" stateProvince="Aragua">
<emphasis id="A78D1B623004523AEC38FC25768202A3" bold="true" box="[303,398,898,924]" pageId="94" pageNumber="95">
<collectingRegion id="573D09923004523AEC38FC25768202A3" box="[303,398,898,924]" country="Venezuela" name="Aragua" pageId="94" pageNumber="95">Aragua</collectingRegion>
</emphasis>
<specimenCount id="83FF0CF93004523AECB2FC2576F602A2" box="[421,506,898,925]" count="1" pageId="94" pageNumber="95" type="generic">1 spec.</specimenCount>
;
<location id="902691AB3004523AEF11FC25757E02A3" LSID="urn:lsid:plazi:treatment:1D5076663004523AEF3CFEAF74FB0446:902691AB3004523AEF11FC25757E02A3" box="[518,626,898,924]" country="Venezuela" name="Cuyagua" pageId="94" pageNumber="95" stateProvince="Aragua">Cuyagua</location>
;
<elevation id="1ED420433004523AEF69FC2575F502A3" box="[638,761,898,925]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="94" pageNumber="95" unit="m" value="50.0">
<quantity id="52016A953004523AEF69FC2575B602A2" box="[638,698,898,925]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="94" pageNumber="95" unit="m" value="50.0">50 m</quantity>
a.s.l.
</elevation>
;
<date id="E147E1B03004523AEE10FC25748002A3" box="[775,908,898,924]" pageId="94" pageNumber="95" value="2004-07-06">
<collectingDate id="F10318583004523AEE10FC25748002A3" box="[775,908,898,924]" pageId="94" pageNumber="95" value="2004-07-06">6 Jul. 2004</collectingDate>
</date>
;
<collectorName id="380CA2A63004523AEE8FFC24730602A3" box="[920,1034,898,925]" pageId="94" pageNumber="95">D. García</collectorName>
leg.; CEMT
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3004523AEDAAFC6474FB0446" pageId="94" pageNumber="95" type="distribution">
<paragraph id="9546C7703004523AEDAAFC64765A02E2" blockId="94.[189,342,963,989]" box="[189,342,963,989]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AEDAAFC64765A02E2" bold="true" box="[189,342,963,989]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A78D1B623004523AEDAAFC64765A02E2" bold="true" box="[189,342,963,989]" pageId="94" pageNumber="95">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703004523AEDAAFC5474900533" blockId="94.[189,924,1010,1037]" box="[189,924,1010,1037]" pageId="94" pageNumber="95">
<collectingCountry id="EDEE87E03004523AEDAAFC5476110532" box="[189,285,1011,1037]" name="Panama" pageId="94" pageNumber="95">Panama</collectingCountry>
,
<collectingCountry id="EDEE87E03004523AEC3EFC5576AC0533" box="[297,416,1010,1036]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
,
<collectingCountry id="EDEE87E03004523AECBDFC55752A0533" box="[426,550,1010,1036]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
,
<collectingCountry id="EDEE87E03004523AEF25FC55759C0533" box="[562,656,1010,1036]" name="Guyana" pageId="94" pageNumber="95">Guyana</collectingCountry>
,
<collectingCountry id="EDEE87E03004523AEF8CFC5574020533" box="[667,782,1010,1036]" name="Suriname" pageId="94" pageNumber="95">Suriname</collectingCountry>
, and
<collectingCountry id="EDEE87E03004523AEE5BFC55749B0533" box="[844,919,1010,1036]" name="Brazil" pageId="94" pageNumber="95">Brazil</collectingCountry>
.
</paragraph>
<paragraph id="9546C7703004523AEDAAFB9476E10571" blockId="94.[189,493,1075,1102]" box="[189,493,1075,1102]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AEDAAFB9476E10571" bold="true" box="[189,493,1075,1102]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A78D1B623004523AEDAAFB9476E10571" bold="true" box="[189,493,1075,1102]" pageId="94" pageNumber="95">
Subregions of
<collectingCountry id="EDEE87E03004523AEC78FB9376E10571" box="[367,493,1076,1102]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703004523AEDAAFBC5756E059F" blockId="94.[188,1398,1122,1184]" pageId="94" pageNumber="95">
Coastal mainland, System of hills and low piedmont mountains of the Guiana Shield, Andes mountains, and Central
<collectingRegion id="573D09923004523AEC5BFB22768305A0" box="[332,399,1157,1183]" country="New Zealand" name="West Coast" pageId="94" pageNumber="95">Coast</collectingRegion>
Mountain Range.
</paragraph>
<paragraph id="9546C7703004523AEDAAFB6176A805DE" blockId="94.[189,420,1222,1249]" box="[189,420,1222,1249]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AEDAAFB6176A805DE" bold="true" box="[189,420,1222,1249]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A78D1B623004523AEDAAFB6176A805DE" bold="true" box="[189,420,1222,1249]" pageId="94" pageNumber="95">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703004523AEDAAFB5274FB0446" blockId="94.[189,1399,1269,1401]" pageId="94" pageNumber="95">
<bibRefCitation id="F168BA813004523AEDAAFB5276870430" author="Gillet J. J. E." box="[189,395,1269,1295]" pageId="94" pageNumber="95" refId="ref121235" refString="Gillet J. J. E. 1911 b. Coleopterorum Catalogus. Pars 38: Scarabaeidae: Coprinae I. W. Junk, Berlin. Available from https: // www. biodiversitylibrary. org / page / 48405679 [accesseed 16 Feb. 2024]." type="url" year="1911">Gillet 1911b: 60</bibRefCitation>
(
<collectingCountry id="EDEE87E03004523AECB6FB52752D0430" box="[417,545,1269,1295]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813004523AEF7FFB5274750430" author="Luederwaldt H." box="[616,889,1269,1295]" pageId="94" pageNumber="95" pagination="603 - 775" refId="ref128687" refString="Luederwaldt H. 1929. As especies brasileiras do genero Pinotus, com algumas consideracoes tambem sobre outras especies. Revista do Museu Paulista 16: 603 - 775." type="journal article" year="1929">Luederwaldt 1929: 17</bibRefCitation>
(
<collectingCountry id="EDEE87E03004523AEE87FB52731C0430" box="[912,1040,1269,1295]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813004523AE941FB52727A0430" author="Blackwelder R. E." box="[1110,1398,1269,1295]" pageId="94" pageNumber="95" pagination="189 - 341" refId="ref111083" refString="Blackwelder R. E. 1944. Checklist of the coleopterous insects of Mexico, Central America, the West Indies, and South America. Part 2. United States National Museum Bulletin 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 207</bibRefCitation>
(
<collectingCountry id="EDEE87E03004523AEDD2FABF7649040D" box="[197,325,1304,1330]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
). —
<bibRefCitation id="F168BA813004523AEC92FABE7535040C" author="Roze J. A." box="[389,569,1304,1331]" pageId="94" pageNumber="95" pagination="39 - 63" refId="ref137914" refString="Roze J. A. 1955. Lista preliminar de la familia Scarabaeidae sensu lato de Venezuela. Boletin del Museo de Ciencias Naturales 1 (1): 39 - 63." type="journal article" year="1955">Roze 1955: 44</bibRefCitation>
(
<collectingCountry id="EDEE87E03004523AEF5AFABF75C0040D" box="[589,716,1304,1330]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923004523AEFCCFABF7499040D" box="[731,917,1304,1330]" country="Venezuela" name="Distrito Capital" pageId="94" pageNumber="95">Distrito Capital</collectingRegion>
and
<collectingRegion id="573D09923004523AEEC0FABF734C040D" box="[983,1088,1304,1330]" country="Venezuela" name="Miranda" pageId="94" pageNumber="95">Miranda</collectingRegion>
). —
<bibRefCitation id="F168BA813004523AE997FABF727B040C" author="Blanco J." box="[1152,1399,1304,1331]" pageId="94" pageNumber="95" pagination="39 - 48" refId="ref111584" refString="Blanco J. 1988. Catalogo de los Scarabaeinae (Coleoptera) coprofagos y necrofagos, del Estado Tachira. Venezuela. Revista Cientifica Unet 2 (1): 39 - 48." type="journal article" year="1988">Blanco 1988: 4041</bibRefCitation>
(
<collectingCountry id="EDEE87E03004523AEDD2FA9B76480469" box="[197,324,1340,1366]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923004523AEC59FA9B76A10469" box="[334,429,1340,1366]" country="Venezuela" name="Tachira" pageId="94" pageNumber="95">Táchira</collectingRegion>
). —
<bibRefCitation id="F168BA813004523AECF3FA9B740D0469" author="Larsen T. H. &amp; Lopera A. &amp; Forsyth A." box="[484,769,1340,1366]" pageId="94" pageNumber="95" pagination="1288 - 1298" refId="ref127038" refString="Larsen T. H., Lopera A. &amp; Forsyth A. 2008. Understanding trait-dependent community disassembly: dung beetles, density functions, and forest fragmentation. Conservation Biology 22 (5): 1288 - 1298. https: // doi. org / 10.1111 / j. 1523 - 1739.2008.00969. x" type="journal article" year="2008">
Larsen
<emphasis id="A78D1B623004523AEF2DFA9A75610469" box="[570,621,1340,1366]" italics="true" pageId="94" pageNumber="95">et al</emphasis>
. 2008: 1294
</bibRefCitation>
(
<collectingCountry id="EDEE87E03004523AEE18FA9B74820469" box="[783,910,1340,1366]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
:
<collectingRegion id="573D09923004523AEE8EFA9B74FA0469" box="[921,1014,1340,1366]" country="Venezuela" name="Bolivar" pageId="94" pageNumber="95">Bolívar</collectingRegion>
). —
<bibRefCitation id="F168BA813004523AE93AFA9B727A0469" author="Ferrer-Paris J. &amp; Sanchez-Mercado A. &amp; Rodriguez J. P." box="[1069,1398,1340,1366]" pageId="94" pageNumber="95" pagination="89 - 110" refId="ref119203" refString="Ferrer-Paris J. R, Sanchez-Mercado A. &amp; Rodriguez J. P. 2013. Optimizacion del muestreo de invertebrados tropicales: un ejemplo con escarabajos coprofagos (Coleoptera: Scarabaeinae) en Venezuela. Revista de Biologia Tropical 61 (1): 89 - 110. https: // doi. org / 10.15517 / rbt. v 61 i 1.10941" type="journal article" year="2013">
Ferrer-Paris
<emphasis id="A78D1B623004523AE9A8FA9A73FE0469" box="[1215,1266,1340,1366]" italics="true" pageId="94" pageNumber="95">et al</emphasis>
. 2013: 109
</bibRefCitation>
(
<collectingRegion id="573D09923004523AEDD2FAF8762F0446" box="[197,291,1375,1401]" country="Venezuela" name="Bolivar" pageId="94" pageNumber="95">Bolívar</collectingRegion>
: Sabanas de Guri,
<collectingRegion id="573D09923004523AECEAFAF875480446" box="[509,580,1375,1401]" country="Venezuela" name="Sucre" pageId="94" pageNumber="95">Sucre</collectingRegion>
: Araya and
<collectingRegion id="573D09923004523AEFD9FAF8741C0446" box="[718,784,1375,1401]" country="Venezuela" name="Zulia" pageId="94" pageNumber="95">Zulia</collectingRegion>
: Rosario de Perijá).
</paragraph>
</subSubSection>
</treatment>
</document>