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<document id="D9356B0BA585AB66B787BDD45A861136" ID-CLB-Dataset="305167" ID-DOI="10.5852/ejt.2024.959.2677" ID-GBIF-Dataset="bf132c8e-dbc7-4f3d-96e9-1a0b153e765a" ID-ISSN="2118-9773" ID-Zenodo-Dep="13945464" ID-ZooBank="30872C13-516F-42FA-AFA7-30ADC6BF1BAF" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="carolina" IM.treatments_approvedBy="carolina" checkinTime="1729166637250" checkinUser="plazi" docAuthor="Rosa, Cecilia Lozano De La, Cupello, Mario &amp; Vaz-De-Mello, Fernando Z." docDate="2024" docId="1D50766630045204ECE8FA07722F006F" docLanguage="en" docName="EJT.2024.959.1-272.pdf" docOrigin="European Journal of Taxonomy 959 (1)" docSource="https://europeanjournaloftaxonomy.eu/index.php/ejt/article/download/2677/12401" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Dichotomius costaricensis" docType="treatment" docVersion="9" lastPageNumber="97" masterDocId="E1690E1E305A5264ED17FFA7770C013F" masterDocTitle="The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution" masterLastPageNumber="272" masterPageNumber="1" pageNumber="95" updateTime="1731534417997" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="477DD14F17DAECCAACC6702B020BF5B0">The dung beetles of Venezuela (Coleoptera: Scarabaeidae: Scarabaeinae): catalogue and updated distribution</mods:title>
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<mods:name id="A243E84C9D5F3BC8AB1569BBAB3B7CD3" type="personal">
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<mods:namePart id="8F23209F77EEABA4B367EC24B8E21D4F">Rosa, Cecilia Lozano De La</mods:namePart>
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<mods:affiliation id="E7C8D6AEA7C4538984C8459C075629F9">Instituto Venezolano de Investigaciones Científicas, Centro de Estudios Botánicos y Agroforestales, Laboratorio de Ecología Espacial, Maracaibo, estado Zulia, Venezuela.</mods:affiliation>
<mods:nameIdentifier id="A4AD6FBA52EF680A36F866B4F46AED1D" type="email">lozanoceci@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name id="646CE6FE60C3FCEC0C01CCB60002CA0C" type="personal">
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<mods:namePart id="E4A8060648A738ED57D0FDA4B899A79D">Cupello, Mario</mods:namePart>
<mods:nameIdentifier id="54A66825109E6550C3A349CA1EB9FF85" type="LSID">BDB03C18-7095-4EAA-8BDD-03CB4F79676D</mods:nameIdentifier>
<mods:affiliation id="96ABF776B4B2FAC07706D6D0E74136D1">Laboratório de Sistemática e Bioecologia de Coleoptera, Departamento de Zoologia, Universidade Federal do Paraná, Curitiba, Paraná, Brazil. Currently at Department of Entomology, Texas A &amp; M University, College Station, TX, 77843, USA.</mods:affiliation>
<mods:nameIdentifier id="EFCC2BF91DD2540484C268D5317D0654" type="email">mcupello@hotmail.com</mods:nameIdentifier>
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<mods:name id="931B3398FBC4F077C9D8C9C50AD8CF9D" type="personal">
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<mods:roleTerm id="2C3C438D751CA323FBA89CBDB523FF66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="59977B24EF31AD6261D89C853F542ED9">Vaz-De-Mello, Fernando Z.</mods:namePart>
<mods:nameIdentifier id="84E99D355D9014E50EAB50605DC80A8D" type="LSID">2BC093C2-3E2B-466B-B31C-9F3D9FA844BC</mods:nameIdentifier>
<mods:affiliation id="9933AB24A2EC8F300B9658E247087F3C">Departamento de Biologia e Zoologia, Instituto de Biociências, Universidade Federal de Mato Grosso, Cuiabá, Mato Grosso, Brazil.</mods:affiliation>
<mods:nameIdentifier id="B8FBC1B648DB5EB53877F8B24E0EA3D5" type="email">vazdemello@gmail.com</mods:nameIdentifier>
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<mods:title id="546E696E06089E5B0DBA42D13E75DB64">European Journal of Taxonomy</mods:title>
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<mods:date id="70D6212AA3B6E6A83F3EAD3235D32633">2024</mods:date>
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<mods:number id="8BDDBA23C326404879F183EF5F3856F0">2024-10-11</mods:number>
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<treatment id="1D50766630045204ECE8FA07722F006F" ID-DOI="http://doi.org/10.5281/zenodo.14154554" ID-GBIF-Taxon="246297819" ID-Zenodo-Dep="14154554" LSID="urn:lsid:plazi:treatment:1D50766630045204ECE8FA07722F006F" httpUri="http://treatment.plazi.org/id/1D50766630045204ECE8FA07722F006F" lastPageId="96" lastPageNumber="97" pageId="94" pageNumber="95">
<subSubSection id="DDE394FB3004523AECE8FA0773380484" box="[511,1076,1440,1467]" pageId="94" pageNumber="95" type="nomenclature">
<paragraph id="9546C7703004523AECE8FA0773380484" blockId="94.[511,1076,1440,1502]" box="[511,1076,1440,1467]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AECE8FA0773380484" box="[511,1076,1440,1467]" centered="true" fontSize="11" level="2" pageId="94" pageNumber="95" reason="2">
<taxonomicName id="52F9BCF33004523AECE8FA0773380484" ID-CoL="7ZN3Y" authority="(Luederwaldt, 1935)" baseAuthorityName="Luederwaldt" baseAuthorityYear="1935" box="[511,1076,1440,1467]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="A78D1B623004523AECE8FA0774300485" bold="true" box="[511,828,1440,1467]" italics="true" pageId="94" pageNumber="95">Dichotomius costaricensis</emphasis>
(
<bibRefCitation id="F168BA813004523AEE5CFA0773200484" author="Luederwaldt H." box="[843,1068,1440,1467]" pageId="94" pageNumber="95" pagination="334 - 342" refId="ref128808" refString="Luederwaldt H. 1935. Zur Kenntnis der Gattung Pinotus (Col. Scarabaeidae). Revista de Entomologia 5 (3): 334 - 342." type="journal article" year="1935">Luederwaldt, 1935</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3004523AEFF1FA64744304E1" box="[742,847,1475,1502]" pageId="94" pageNumber="95" type="description">
<paragraph id="9546C7703004523AEFF1FA64744304E1" blockId="94.[511,1076,1440,1502]" box="[742,847,1475,1502]" pageId="94" pageNumber="95">
<figureCitation id="0DC2DBF53004523AEFF1FA64744304E1" box="[742,847,1475,1502]" captionStart="Fig" captionStartId="95.[189,232,1963,1989]" captionTargetBox="[274,1305,278,1891]" captionTargetId="figure-12@95.[263,1323,265,1907]" captionTargetPageId="95" captionText="Fig. 38. Species distribution. A. Dichotomius costaricensis (Luederwaldt. 1935). B. Dichotomius deyrollei (Harold, 1869). Green diamond = CEMT collection data; black circle = literature data." figureDoi="http://doi.org/10.5281/zenodo.13945569" httpUri="https://zenodo.org/record/13945569/files/figure.png" pageId="94" pageNumber="95">Fig. 38A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3004523AEDAAF9A1730B07D2" pageId="94" pageNumber="95" type="reference_group">
<paragraph id="9546C7703004523AEDAAF9A174B6077C" blockId="94.[189,1399,1541,1603]" pageId="94" pageNumber="95">
<treatmentCitationGroup id="B5E9E05E3004523AEDAAF9A174B6077C" pageId="94" pageNumber="95">
<treatmentCitation id="1458E1613004523AEDAAF9A17431071F" author="Luederwaldt H." box="[189,829,1541,1568]" page="335" pageId="94" pageNumber="95" year="1935">
<taxonomicName id="52F9BCF33004523AEDAAF9A17431071F" ID-CoL="7Y7ZD" authority="Luederwaldt, 1935: 335" authorityName="Luederwaldt" authorityPageNumber="335" authorityYear="1935" box="[189,829,1541,1568]" class="Insecta" family="Scarabaeidae" genus="Pinotus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="subSpecies" species="bitiensis" subSpecies="costaricensis">
<emphasis id="A78D1B623004523AEDAAF9A175130720" box="[189,543,1541,1567]" italics="true" pageId="94" pageNumber="95">Pinotus bitiensis costaricensis</emphasis>
<bibRefCitation id="F168BA813004523AEF32F9A27431071F" author="Luederwaldt H." box="[549,829,1541,1568]" pageId="94" pageNumber="95" pagination="334 - 342" refId="ref128808" refString="Luederwaldt H. 1935. Zur Kenntnis der Gattung Pinotus (Col. Scarabaeidae). Revista de Entomologia 5 (3): 334 - 342." type="journal article" year="1935">Luederwaldt, 1935: 335</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description). Type locality:
<collectingCountry id="EDEE87E03004523AE9E7F9A2727F0720" box="[1264,1395,1541,1567]" name="Costa Rica" pageId="94" pageNumber="95">Costa Rica</collectingCountry>
. Name-bearing type:
<typeStatus id="4A4279D23004523AECCCF98F754E077D" box="[475,578,1576,1602]" pageId="94" pageNumber="95" type="holotype">holotype</typeStatus>
(BMNH), examined by FZVM.
</treatmentCitationGroup>
</paragraph>
<paragraph id="9546C7703004523AEDAAF9CD730B07D2" blockId="94.[189,1399,1641,1774]" pageId="94" pageNumber="95">
<treatmentCitationGroup id="B5E9E05E3004523AEDAAF9CD730B07D2" pageId="94" pageNumber="95">
<taxonomicName id="52F9BCF33004523AEDAAF9CD76FD07BC" ID-CoL="7ZN3Y" box="[189,497,1642,1668]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="94" pageNumber="95" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="A78D1B623004523AEDAAF9CD76FD07BC" box="[189,497,1642,1668]" italics="true" pageId="94" pageNumber="95">Dichotomius costaricensis</emphasis>
</taxonomicName>
<treatmentCitation id="1458E1613004523AEF1DF9CD745007BB" author="Kohlmann B. &amp; Solis A." box="[522,860,1641,1668]" page="345" pageId="94" pageNumber="95" year="1997">
<bibRefCitation id="F168BA813004523AEF1DF9CD745007BB" author="Kohlmann B. &amp; Solis A." box="[522,860,1641,1668]" pageId="94" pageNumber="95" pagination="343 - 382" refId="ref126101" refString="Kohlmann B. &amp; Solis A. 1997. El genero Dichotomius en Costa Rica. Giornale Italiano di Entomologia 8: 343 - 382." type="journal article" year="1997">Kohlmann &amp; Solís 1997: 345</bibRefCitation>
</treatmentCitation>
, 360 (key, redescription, new combination). —
<treatmentCitation id="1458E1613004523AEDFFF92A76C60798" author="Escobar F." box="[232,458,1677,1703]" page="208" pageId="94" pageNumber="95" year="2000">
<bibRefCitation id="F168BA813004523AEDFFF92A76C60798" author="Escobar F." box="[232,458,1677,1703]" pageId="94" pageNumber="95" pagination="197 - 210" refId="ref118039" refString="Escobar F. 2000. Diversidad y distribution de los escarabajos del estiercol (Coleoptera: Scarabaeidae: Scarabaeinae) de Colombia. In: Martin-Piera F., Morrone J. J. &amp; Melic A. (eds.) Hacia un proyecto CYTED para el Inventario y Estimacion de la Diversidad Entomologica en Iberoamerica: PRIBES- 2000: 197 - 210. m 3 m: Monografias Tercer Milenio, vol. 1. Sociedad Entomologica Aragonesa (SEA), Zaragoza." type="book chapter" year="2000">Escobar 2000: 208</bibRefCitation>
</treatmentCitation>
(list for
<collectingCountry id="EDEE87E03004523AEF2EF92A75BE0798" box="[569,690,1677,1703]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AEFE5F92A731B0798" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[754,1047,1677,1703]" page="138" pageId="94" pageNumber="95" year="2001">
<bibRefCitation id="F168BA813004523AEFE5F92A731B0798" author="Medina C. A. &amp; Lopera-Toro A. &amp; Vitolo A. &amp; Gill B." box="[754,1047,1677,1703]" pageId="94" pageNumber="95" pagination="131 - 144" refId="ref130799" refString="Medina C. A., Lopera-Toro A., Vitolo A. &amp; Gill B. 2001. Escarabajos coprofagos (Coleoptera: Scarabaeinae) de Colombia. Biota Colombiana 2: 131 - 144." type="journal article" year="2001">
Medina
<emphasis id="A78D1B623004523AEE41F929749F0798" box="[854,915,1677,1703]" italics="true" pageId="94" pageNumber="95">et al.</emphasis>
2001: 138
</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03004523AE9DDF92A724F0798" box="[1226,1347,1677,1703]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AEDFDF91776CF07F4" author="Ratcliffe B. C." box="[234,451,1712,1739]" page="15" pageId="94" pageNumber="95" year="2002">
<bibRefCitation id="F168BA813004523AEDFDF91776CF07F4" author="Ratcliffe B. C." box="[234,451,1712,1739]" firstAuthor="Ratcliffe" pageId="94" pageNumber="95" pagination="1 - 48" refId="ref136799" refString="Ratcliffe B. C. 2002. A checklist of the Scarabaeoidea of Panama. Zootaxa 32: 1 - 48. https: // doi. org / 10.11646 / zootaxa. 32.1.1" type="journal article" year="2002">Ratcliffe 2002: 15</bibRefCitation>
</treatmentCitation>
(checklist for
<collectingCountry id="EDEE87E03004523AEF64F91775D907F5" box="[627,725,1712,1738]" name="Panama" pageId="94" pageNumber="95">Panama</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AEE06F917735407F5" author="Solis A. &amp; Kohlmann B." box="[785,1112,1712,1738]" page="6" pageId="94" pageNumber="95" year="2012">
<bibRefCitation id="F168BA813004523AEE06F917735407F5" author="Solis A. &amp; Kohlmann B." box="[785,1112,1712,1738]" pageId="94" pageNumber="95" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 6</bibRefCitation>
</treatmentCitation>
, 12 (checklist for
<collectingCountry id="EDEE87E03004523AE823F917762B07D2" name="Costa Rica" pageId="94" pageNumber="95">Costa Rica</collectingCountry>
). —
<treatmentCitation id="1458E1613004523AEC48F974745207D2" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[351,862,1747,1773]" page="84" pageId="94" pageNumber="95" year="2014">
<bibRefCitation id="F168BA813004523AEC48F974745207D2" author="Sarmiento-Garces R. &amp; Amat-Garcia G." box="[351,862,1747,1773]" pageId="94" pageNumber="95" refId="ref138148" refString="Sarmiento-Garces R. &amp; Amat-Garcia G. 2014. Escarabajos del Genero Dichotomius Hope 1838 (Scarabaeidae: Scarabaeinae) en Colombia. Fauna de Colombia, Monografia n ° 4, Universidad Nacional de Colombia, Bogota. https: // doi. org / 10.18257 / raccefyn. 33 (127). 2009.2364" type="book" year="2014">Sarmiento-Garcés &amp; Amat-García 2014: 84</bibRefCitation>
</treatmentCitation>
(monograph).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="DDE394FB3004523AEDAAF8B3759806F3" pageId="94" pageNumber="95" type="distribution">
<paragraph id="9546C7703004523AEDAAF8B3765A0611" blockId="94.[189,342,1812,1838]" box="[189,342,1812,1838]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AEDAAF8B3765A0611" bold="true" box="[189,342,1812,1838]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A78D1B623004523AEDAAF8B3765A0611" bold="true" box="[189,342,1812,1838]" pageId="94" pageNumber="95">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703004523AEDAAF8E475E70662" blockId="94.[189,747,1859,1886]" box="[189,747,1859,1886]" pageId="94" pageNumber="95">
<collectingCountry id="EDEE87E03004523AEDAAF8E4764C0662" box="[189,320,1859,1885]" name="Costa Rica" pageId="94" pageNumber="95">Costa Rica</collectingCountry>
,
<collectingCountry id="EDEE87E03004523AEC5BF8E376A00661" box="[332,428,1860,1886]" name="Panama" pageId="94" pageNumber="95">Panama</collectingCountry>
,
<collectingCountry id="EDEE87E03004523AECA0F8E475220662" box="[439,558,1859,1885]" name="Colombia" pageId="94" pageNumber="95">Colombia</collectingCountry>
, and
<collectingCountry id="EDEE87E03004523AEF7CF8E475E40662" box="[619,744,1859,1885]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
.
</paragraph>
<paragraph id="9546C7703004523AEDAAF82376EE06A1" blockId="94.[189,482,1924,1950]" box="[189,482,1924,1950]" pageId="94" pageNumber="95">
<heading id="CE0E701C3004523AEDAAF82376EE06A1" bold="true" box="[189,482,1924,1950]" fontSize="11" level="3" pageId="94" pageNumber="95" reason="3">
<emphasis id="A78D1B623004523AEDAAF82376EE06A1" bold="true" box="[189,482,1924,1950]" pageId="94" pageNumber="95">
Subregion of
<collectingCountry id="EDEE87E03004523AEC73F82376EE06A1" box="[356,482,1924,1950]" name="Venezuela" pageId="94" pageNumber="95">Venezuela</collectingCountry>
</emphasis>
</heading>
</paragraph>
<paragraph id="9546C7703004523AEDAAF815759806F3" blockId="94.[189,660,1970,1996]" box="[189,660,1970,1996]" pageId="94" pageNumber="95">
Sierra of
<collectingRegion id="573D09923004523AEC3DF815769D06F3" box="[298,401,1970,1996]" country="Argentina" name="San Luis" pageId="94" pageNumber="95">San Luis</collectingRegion>
and Cerro
<collectingRegion id="573D09923004523AEF02F815759D06F3" box="[533,657,1970,1996]" country="El Salvador" name="Santa Ana" pageId="94" pageNumber="95">Santa Ana</collectingRegion>
.
</paragraph>
</subSubSection>
<caption id="C18697F83005523BEDAAF80C722E06D7" ID-DOI="http://doi.org/10.5281/zenodo.13945569" ID-Zenodo-Dep="13945569" httpUri="https://zenodo.org/record/13945569/files/figure.png" pageId="95" pageNumber="96" startId="95.[189,232,1963,1989]" targetBox="[274,1305,278,1891]" targetPageId="95">
<paragraph id="9546C7703005523BEDAAF80C722E06D7" blockId="95.[189,1399,1963,2025]" pageId="95" pageNumber="96">
<emphasis id="A78D1B623005523BEDAAF80C762D06FA" bold="true" box="[189,289,1963,1989]" pageId="95" pageNumber="96">Fig. 38.</emphasis>
Species distribution.
<emphasis id="A78D1B623005523BEF23F80C754506FA" bold="true" box="[564,585,1963,1989]" pageId="95" pageNumber="96">A</emphasis>
.
<taxonomicName id="52F9BCF33005523BEF48F80C73AA06F9" authority="(Luederwaldt. 1935)" baseAuthorityName="Luederwaldt." baseAuthorityYear="1935" box="[607,1190,1963,1990]" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="costaricensis">
<emphasis id="A78D1B623005523BEF48F80C749006FA" box="[607,924,1963,1989]" italics="true" pageId="95" pageNumber="96">Dichotomius costaricensis</emphasis>
(Luederwaldt. 1935)
</taxonomicName>
.
<emphasis id="A78D1B623005523BE9A0F80C73C706FA" bold="true" box="[1207,1227,1963,1989]" pageId="95" pageNumber="96">B</emphasis>
.
<taxonomicName id="52F9BCF33005523BE9F6F80C76D206D7" authority="(Harold, 1869)" baseAuthorityName="Harold" baseAuthorityYear="1869" class="Insecta" family="Scarabaeidae" genus="Dichotomius" kingdom="Animalia" order="Coleoptera" pageId="95" pageNumber="96" phylum="Arthropoda" rank="species" species="deyrollei">
<emphasis id="A78D1B623005523BE9F6F80C762806D7" italics="true" pageId="95" pageNumber="96">Dichotomius deyrollei</emphasis>
(Harold, 1869)
</taxonomicName>
. Green diamond = CEMT collection data; black circle = literature data.
</paragraph>
</caption>
<subSubSection id="DDE394FB303A5204EDAAFEAF722F006F" pageId="96" pageNumber="97" type="reference_group">
<paragraph id="9546C770303A5204EDAAFEAF76A8001C" blockId="96.[189,420,264,291]" box="[189,420,264,291]" pageId="96" pageNumber="97">
<heading id="CE0E701C303A5204EDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" fontSize="11" level="3" pageId="96" pageNumber="97" reason="3">
<emphasis id="A78D1B62303A5204EDAAFEAF76A8001C" bold="true" box="[189,420,264,291]" pageId="96" pageNumber="97">Literature records</emphasis>
</heading>
</paragraph>
<paragraph id="9546C770303A5204EDAAFE91722F006F" blockId="96.[189,1315,310,337]" box="[189,1315,310,337]" pageId="96" pageNumber="97">
<bibRefCitation id="F168BA81303A5204EDAAFE91751A006F" author="Kohlmann B. &amp; Solis A." box="[189,534,310,337]" pageId="96" pageNumber="97" pagination="343 - 382" refId="ref126101" refString="Kohlmann B. &amp; Solis A. 1997. El genero Dichotomius en Costa Rica. Giornale Italiano di Entomologia 8: 343 - 382." type="journal article" year="1997">Kohlmann &amp; Solís 1997: 360</bibRefCitation>
(
<collectingCountry id="EDEE87E0303A5204EF31FE9175A9006F" box="[550,677,310,336]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
:
<collectingRegion id="573D0992303A5204EFA7FE917409006F" box="[688,773,310,336]" country="Venezuela" name="Falcon" pageId="96" pageNumber="97">Falcón</collectingRegion>
). —
<bibRefCitation id="F168BA81303A5204EE2AFE91738B006E" author="Solis A. &amp; Kohlmann B." box="[829,1159,310,337]" pageId="96" pageNumber="97" pagination="1 - 32" refId="ref139201" refString="Solis A. &amp; Kohlmann B. 2012. Checklist and distribution atlas of the Scarabaeinae of Costa Rica. Zootaxa 3482: 1 - 32. https: // doi. org / 10.11646 / zootaxa. 3482.1.1" type="journal article" year="2012">Solís &amp; Kohlmann 2012: 12</bibRefCitation>
(
<collectingCountry id="EDEE87E0303A5204E980FE91721B006F" box="[1175,1303,310,336]" name="Venezuela" pageId="96" pageNumber="97">Venezuela</collectingCountry>
).
</paragraph>
</subSubSection>
</treatment>
</document>