treatments-xml/data/1D/14/87/1D1487A32E4AFFF6FE85FB48FB14F9A0.xml
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<document id="02630514DE1F65BF78F3E2DA6AC58910" ID-DOI="10.5281/zenodo.7709961" ID-ISSN="1536-9307" ID-Zenodo-Dep="7709961" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1678328567740" checkinUser="felipe" docAuthor="Al-Saraireh, MOhammad, Yağmur, Ersen Aydın, Afifeh, Bassam Abu &amp; Amr, Zuhair" docDate="2023" docId="1D1487A32E4AFFF6FE85FB48FB14F9A0" docLanguage="en" docName="Euscorpius.369.1-1-19.pdf" docOrigin="Euscorpius 369" docStyle="DocumentStyle:5E96F6379CE20B0E8E4F9E7DF8E1C0DF.2:Euscorpius.2020-.journal_article" docStyleId="5E96F6379CE20B0E8E4F9E7DF8E1C0DF" docStyleName="Euscorpius.2020-.journal_article" docStyleVersion="2" docTitle="Scorpio granulomanus Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr 2023, sp. n." docType="treatment" docVersion="3" lastPageNumber="14" masterDocId="E12DFFDB2E4EFFF9FF8EFF9EFFABFFC2" masterDocTitle="A new species Of Scorpio frOm JOrdan (ScOrpiOnes: ScOrpiOnidae)" masterLastPageNumber="17" masterPageNumber="1" pageNumber="3" updateTime="1699461463767" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="EE2A01B6119DD05C83B4FDD9B0D8BF39">A new species Of Scorpio frOm JOrdan (ScOrpiOnes: ScOrpiOnidae)</mods:title>
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(Figures 159,
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<specimenCount id="83BBFD3C2E4AFFFDFEA2F95CFEE5F918" box="[300,334,1730,1754]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
(
<typeStatus id="4A0688172E4AFFFDFEEDF95CFE6FF918" box="[355,452,1730,1754]" pageId="4" pageNumber="3" type="holotype">holotype</typeStatus>
) (
<collectionCode id="F3ACAE702E4AFFFDFE6EF95CFD92F918" box="[480,569,1730,1754]" pageId="4" pageNumber="3">AZMM</collectionCode>
/Sco-2022:01),
<specimenCount id="83BBFD3C2E4AFFFDFD6CF95CFCA8F918" box="[738,771,1730,1754]" pageId="4" pageNumber="3" type="female">1♀</specimenCount>
(
<typeStatus id="4A0688172E4AFFFDFFF7F97CFF73F938" box="[121,216,1762,1786]" pageId="4" pageNumber="3" type="paratype">paratype</typeStatus>
) (
<collectionCode id="F3ACAE702E4AFFFDFF78F97CFEE5F938" box="[246,334,1762,1786]" pageId="4" pageNumber="3">AZMM</collectionCode>
/Sco-2022:02),
<specimenCount id="83BBFD3C2E4AFFFDFE74F97CFDB0F938" box="[506,539,1762,1786]" pageId="4" pageNumber="3" type="male">1♂</specimenCount>
<specimenCount id="83BBFD3C2E4AFFFDFD95F97CFDE1F938" box="[539,586,1762,1786]" pageId="4" pageNumber="3" type="female">10♀</specimenCount>
(
<typeStatus id="4A0688172E4AFFFDFDEFF97CFD66F938" box="[609,717,1762,1786]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
),
<date id="E10310752E4AFFFDFD66F97CFF4CF8D8" pageId="4" pageNumber="3" value="2022-05-20">
<collectingDate id="F147E99D2E4AFFFDFD66F97CFF4CF8D8" pageId="4" pageNumber="3" value="2022-05-20">20 May 2022</collectingDate>
</date>
, leg.
<collectorName id="384853632E4AFFFDFEA2F89DFE67F8D8" box="[300,460,1794,1818]" pageId="4" pageNumber="3">B. Abu Afifeh</collectorName>
&amp;
<collectorName id="384853632E4AFFFDFE74F89DFD0BF8D8" box="[506,672,1794,1818]" pageId="4" pageNumber="3">M. Al-Saraireh</collectorName>
(
<collectionCode id="F3ACAE702E4AFFFDFD3BF89CFD5CF8D8" box="[693,759,1794,1818]" pageId="4" pageNumber="3">JUST</collectionCode>
);
<location id="9062606E2E4AFFFDFFFFF8BCFE38F8F8" LSID="urn:lsid:plazi:treatment:1D1487A32E4AFFF6FE85FB48FB14F9A0:9062606E2E4AFFFDFFFFF8BCFE38F8F8" box="[113,403,1826,1850]" country="Jordan" county="Dibbeen Forest Reserve" latitude="32.24797" longLatPrecision="1" longitude="35.823277" municipality="Dibbeen Forest Reserve" name="Dibbeen Forest Reserve" pageId="4" pageNumber="3" stateProvince="Jarash">Dibbeen Forest Reserve</location>
,
<geoCoordinate id="F08950722E4AFFFDFE22F8BCFDE0F8F8" box="[428,587,1826,1850]" degrees="32" direction="north" minutes="14" orientation="latitude" pageId="4" pageNumber="3" precision="1" seconds="52.70" value="32.24797">32°14'52.70&quot;N</geoCoordinate>
<geoCoordinate id="F08950722E4AFFFDFDEEF8BCFD54F8F8" box="[608,767,1826,1850]" degrees="35" direction="east" minutes="49" orientation="longitude" pageId="4" pageNumber="3" precision="1" seconds="23.80" value="35.823277">35°49'23.80&quot;E</geoCoordinate>
,
<collectingRegion id="5779F8572E4AFFFDFFFFF8DCFEEDF898" box="[113,326,1858,1882]" country="Jordan" name="Jarash" pageId="4" pageNumber="3">Jerash Governorate</collectingRegion>
,
<elevation id="1E90D1862E4AFFFDFEDAF8DCFE40F898" box="[340,491,1858,1882]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="4" pageNumber="3" unit="m" value="741.0">
<quantity id="52459B502E4AFFFDFEDAF8DCFE30F898" box="[340,411,1858,1882]" metricMagnitude="2" metricUnit="m" metricValue="7.409999999999999" pageId="4" pageNumber="3" unit="m" value="741.0">741 m</quantity>
a. s. l.
</elevation>
,
<specimenCount id="83BBFD3C2E4AFFFDFE74F8DCFDB0F898" box="[506,539,1858,1882]" pageId="4" pageNumber="3" type="female">2♀</specimenCount>
(
<typeStatus id="4A0688172E4AFFFDFDA3F8DCFD32F898" box="[557,665,1858,1882]" pageId="4" pageNumber="3" type="paratype">paratypes</typeStatus>
),
<date id="E10310752E4AFFFDFD20F8DCFF02F8B8" pageId="4" pageNumber="3" value="2022-06-26">
<collectingDate id="F147E99D2E4AFFFDFD20F8DCFF02F8B8" pageId="4" pageNumber="3" value="2022-06-26">26 June 2022</collectingDate>
</date>
, leg.
<collectorName id="384853632E4AFFFDFF6CF8FDFEDCF8B8" box="[226,375,1890,1914]" pageId="4" pageNumber="3">B. Abu Afifeh</collectorName>
&amp;
<collectorName id="384853632E4AFFFDFE17F8FDFD93F8B8" box="[409,568,1890,1914]" pageId="4" pageNumber="3">M. Al-Saraireh</collectorName>
(
<collectionCode id="F3ACAE702E4AFFFDFDC9F8FCFD22F8B8" box="[583,649,1890,1914]" pageId="4" pageNumber="3">JUST</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DDA7653E2E4AFFFDFFFFF83CFF52F838" pageId="4" pageNumber="3" type="etymology">
<paragraph id="950236B52E4AFFFDFFFFF83CFF52F838" blockId="4.[113,772,1954,2042]" pageId="4" pageNumber="3">ETYMOLOGY. The specific epithet refers to a heavily granulated chela manus of this species, the most distinct character for this new species.</paragraph>
</subSubSection>
<subSubSection id="DDA7653E2E4AFFFCFCBEFBA7FDBCF839" lastPageId="5" lastPageNumber="4" pageId="4" pageNumber="3" type="diagnosis">
<paragraph id="950236B52E4AFFFDFCBEFBA7FA69F833" blockId="4.[816,1475,1081,2033]" pageId="4" pageNumber="3">
DIAGNOSIS. Medium-sized species compared with its congeners. Adults are about
<quantity id="52459B502E4AFFFDFBF8FBC7FB6DFBB3" box="[1142,1222,1113,1137]" metricMagnitude="-2" metricUnit="m" metricValue="4.8" pageId="4" pageNumber="3" unit="mm" value="48.0">48 mm</quantity>
in length. Coloration: carapace is light reddish brown in males and dark reddish brown in females with reticulations in both seXes. Mesosoma: basically, reddish brown in males and reddish black in females. Venter dark yellow in males and light brown in females. Pedipalp: femur and patella dark yellow, chela reddish yellow in both seXes. Metasoma and legs: Dark yellow in males and light brownish yellow in females. Prosoma: anterior margin of carapace bilobed, carapace finely granulated, the triangular region between the median eyes and the anterior margin is smooth with scattered moderate granules in males, with fine and dense granules laterally and scattered moderate granules anteriorly and posteriorly in females. DorsoeXternal carinae of pedipalp femur distinct with separate granules distally, sometimes with several rows of spaced granules, intercarinal surface smooth with numerous setae, dorsoeXternal surface with numerous rounded and pointed granules of different size, internal surface with some large, pointed granules. Patella with two carinae. Dorsomedian carina distinct with swollen and some fused granules and the ventrointernal carinae are obsolete with few swollen granules, intercarinal surface is smooth with numerous setae. Chela manus densely covered with large rounded and somewhat pointed granules, the posterior margin of the eXternal surface of chela manus bears very large spinoid protrusions. Internal surface of chela manus with small to moderate pointed granules along the dorsal margin. Chela manus flattened and elongated; manus length to chela depth ratio almost the same in both seXes. Chela fingers are relatively long with small pointed gradually
</paragraph>
<paragraph id="950236B52E4BFFFCFFFDF832FDBCF839" blockId="5.[115,1474,1964,2043]" pageId="5" pageNumber="4">
<emphasis id="A7C9EAA72E4BFFFCFFFDF832FF5DF803" bold="true" box="[115,246,1964,1986]" pageId="5" pageNumber="4">FiGures 512</emphasis>
:
<taxonomicName id="52BD4D362E4BFFFCFE8DF832FE74F803" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[259,479,1964,1985]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E4BFFFCFE8DF832FE74F803" box="[259,479,1964,1985]" italics="true" pageId="5" pageNumber="4">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E4BFFFCFE6BF832FDB1F803" box="[485,538,1964,1985]" pageId="5" pageNumber="4" rank="species">
<emphasis id="A7C9EAA72E4BFFFCFE6BF832FE57F803" bold="true" box="[485,508,1964,1985]" pageId="5" pageNumber="4">sp</emphasis>
.
<emphasis id="A7C9EAA72E4BFFFCFD86F832FDBEF803" bold="true" box="[520,533,1964,1985]" pageId="5" pageNumber="4">n</emphasis>
.
</taxonomicNameLabel>
, female
<typeStatus id="4A0688172E4BFFFCFDE1F832FD69F803" box="[623,706,1964,1985]" pageId="5" pageNumber="4" type="paratype">paratype</typeStatus>
(58) and male
<typeStatus id="4A0688172E4BFFFCFCE9F832FC10F800" box="[871,955,1964,1986]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
(912).
<emphasis id="A7C9EAA72E4BFFFCFB80F832FBC6F803" bold="true" box="[1038,1133,1964,1986]" pageId="5" pageNumber="4">FiGures 5</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFBF7F832FB2EF803" bold="true" box="[1145,1157,1964,1985]" pageId="5" pageNumber="4">7</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFB1FF832FB36F803" bold="true" box="[1169,1181,1964,1985]" pageId="5" pageNumber="4">9</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFB24F832FB6AF803" bold="true" box="[1194,1217,1964,1985]" pageId="5" pageNumber="4">11</emphasis>
. Carapace and mesosoma.
<emphasis id="A7C9EAA72E4BFFFCFFFDF856FF7AF81F" bold="true" box="[115,209,1992,2014]" pageId="5" pageNumber="4">FiGures 6</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFF52F856FF43F81F" bold="true" box="[220,232,1992,2013]" pageId="5" pageNumber="4">8</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFF7AF856FEA7F81F" bold="true" box="[244,268,1992,2013]" pageId="5" pageNumber="4">10</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFE99F856FE84F81F" bold="true" box="[279,303,1992,2013]" pageId="5" pageNumber="4">12</emphasis>
. CoXal and sternopectinal regions and sternites.
<emphasis id="A7C9EAA72E4BFFFCFD73F856FCDBF81F" bold="true" box="[765,880,1992,2014]" pageId="5" pageNumber="4">FiGures 5-6</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFCF5F856FC0CF81F" bold="true" box="[891,935,1992,2013]" pageId="5" pageNumber="4">9-10</emphasis>
. Under white light.
<emphasis id="A7C9EAA72E4BFFFCFBE8F856FB73F81F" bold="true" box="[1126,1240,1992,2014]" pageId="5" pageNumber="4">FiGures 7-8</emphasis>
,
<emphasis id="A7C9EAA72E4BFFFCFB6AF856FAB0F81F" bold="true" box="[1252,1307,1992,2013]" pageId="5" pageNumber="4">11-12</emphasis>
. Under UV light. Scale bars:
<quantity id="52459B502E4BFFFCFF6FF878FE8FF839" box="[225,292,2022,2043]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="4" unit="mm" value="10.0">10 mm</quantity>
(58) and
<quantity id="52459B502E4BFFFCFE03F878FE7BF839" box="[397,464,2022,2043]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="4" unit="mm" value="10.0">10 mm</quantity>
(912)
</paragraph>
</subSubSection>
<subSubSection id="DDA7653E2E48FFF6FFF8F79DFB14F9A0" lastPageId="15" lastPageNumber="14" pageId="6" pageNumber="5" type="description">
<paragraph id="950236B52E48FFFFFFF8F79DFC7EF7F4" blockId="6.[118,1477,2051,2102]" pageId="6" pageNumber="5">
<emphasis id="A7C9EAA72E48FFFFFFF8F79DFEA2F7DA" bold="true" box="[118,265,2051,2073]" pageId="6" pageNumber="5">FiGures 1316</emphasis>
:
<taxonomicName id="52BD4D362E48FFFFFE97F79AFE52F7DA" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[281,505,2051,2073]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E48FFFFFE97F79AFE52F7DA" box="[281,505,2051,2073]" italics="true" pageId="6" pageNumber="5">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E48FFFFFD8CF79DFD90F7DB" box="[514,571,2051,2073]" pageId="6" pageNumber="5" rank="species">
<emphasis id="A7C9EAA72E48FFFFFD8CF79DFDB2F7DA" bold="true" box="[514,537,2051,2072]" pageId="6" pageNumber="5">sp</emphasis>
.
<emphasis id="A7C9EAA72E48FFFFFDA6F79DFD9EF7DA" bold="true" box="[552,565,2051,2072]" pageId="6" pageNumber="5">n</emphasis>
.
</taxonomicNameLabel>
, female
<typeStatus id="4A0688172E48FFFFFD18F79AFD47F7DB" box="[662,748,2052,2073]" pageId="6" pageNumber="5" type="paratype">paratype</typeStatus>
.
<emphasis id="A7C9EAA72E48FFFFFD77F79DFCCCF7DA" bold="true" box="[761,871,2051,2073]" pageId="6" pageNumber="5">FiGures 13</emphasis>
,
<emphasis id="A7C9EAA72E48FFFFFCF9F79DFC24F7DA" bold="true" box="[887,911,2051,2072]" pageId="6" pageNumber="5">15</emphasis>
. Carapace and tergites IIII.
<emphasis id="A7C9EAA72E48FFFFFB37F79DFA83F7DA" bold="true" box="[1209,1320,2051,2073]" pageId="6" pageNumber="5">FiGures 14</emphasis>
,
<emphasis id="A7C9EAA72E48FFFFFAB9F79DFAE4F7DA" bold="true" box="[1335,1359,2051,2072]" pageId="6" pageNumber="5">16</emphasis>
. CoXal and sternopectinal region.
<emphasis id="A7C9EAA72E48FFFFFEC2F7BEFE77F7F7" bold="true" box="[332,476,2080,2102]" pageId="6" pageNumber="5">FiGures 1314</emphasis>
. Under white light.
<emphasis id="A7C9EAA72E48FFFFFD13F7BEFC87F7F7" bold="true" box="[669,812,2080,2102]" pageId="6" pageNumber="5">FiGures 1516</emphasis>
. Under UV light.
</paragraph>
<paragraph id="950236B52E49FFFEFFFAF870FD5FF7E3" blockId="7.[116,1475,2030,2081]" pageId="7" pageNumber="6">
<emphasis id="A7C9EAA72E49FFFEFFFAF870FEAEF7C1" bold="true" box="[116,261,2030,2052]" pageId="7" pageNumber="6">FiGures 1720</emphasis>
:
<taxonomicName id="52BD4D362E49FFFEFE9AF870FE59F7C1" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[276,498,2030,2051]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="7" pageNumber="6" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E49FFFEFE9AF870FE59F7C1" box="[276,498,2030,2051]" italics="true" pageId="7" pageNumber="6">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E49FFFEFE74F870FD9AF7C1" box="[506,561,2030,2051]" pageId="7" pageNumber="6" rank="species">
<emphasis id="A7C9EAA72E49FFFEFE74F870FDBAF7C1" bold="true" box="[506,529,2030,2051]" pageId="7" pageNumber="6">sp</emphasis>
.
<emphasis id="A7C9EAA72E49FFFEFD91F870FD87F7C1" bold="true" box="[543,556,2030,2051]" pageId="7" pageNumber="6">n</emphasis>
.
</taxonomicNameLabel>
, male
<typeStatus id="4A0688172E49FFFEFDF9F870FD65F7C6" box="[631,718,2030,2052]" pageId="7" pageNumber="6" type="holotype">holotype</typeStatus>
.
<emphasis id="A7C9EAA72E49FFFEFD57F870FCEDF7C1" bold="true" box="[729,838,2030,2052]" pageId="7" pageNumber="6">FiGures 17</emphasis>
,
<emphasis id="A7C9EAA72E49FFFEFCDAF870FCC7F7C1" bold="true" box="[852,876,2030,2051]" pageId="7" pageNumber="6">19</emphasis>
. Carapace.
<emphasis id="A7C9EAA72E49FFFEFC6CF870FBE4F7C1" bold="true" box="[994,1103,2030,2052]" pageId="7" pageNumber="6">FiGures 18</emphasis>
,
<emphasis id="A7C9EAA72E49FFFEFBD3F870FBDEF7C1" bold="true" box="[1117,1141,2030,2051]" pageId="7" pageNumber="6">20</emphasis>
. CoXal and sternopectinal region.
<emphasis id="A7C9EAA72E49FFFEFFFAF794FF54F7DD" bold="true" box="[116,255,2058,2080]" pageId="7" pageNumber="6">FiGures 17-18</emphasis>
. Under white light.
<emphasis id="A7C9EAA72E49FFFEFE4EF794FDE0F7DD" bold="true" box="[448,587,2058,2080]" pageId="7" pageNumber="6">FiGures 19-20</emphasis>
. Under UV light.
</paragraph>
<paragraph id="950236B52E46FFF1FFFFF877FC59F7D9" blockId="8.[113,1471,2025,2076]" pageId="8" pageNumber="7">
<emphasis id="A7C9EAA72E46FFF1FFFFF877FEAAF83C" bold="true" box="[113,257,2025,2047]" pageId="8" pageNumber="7">FiGures 2126</emphasis>
:
<taxonomicName id="52BD4D362E46FFF1FE81F877FE40F83C" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[271,491,2025,2046]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="8" pageNumber="7" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E46FFF1FE81F877FE40F83C" box="[271,491,2025,2046]" italics="true" pageId="8" pageNumber="7">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E46FFF1FE7CF877FD83F83C" box="[498,552,2025,2046]" pageId="8" pageNumber="7" rank="species">
<emphasis id="A7C9EAA72E46FFF1FE7CF877FDA2F83C" bold="true" box="[498,521,2025,2046]" pageId="8" pageNumber="7">sp</emphasis>
.
<emphasis id="A7C9EAA72E46FFF1FD98F877FD88F83C" bold="true" box="[534,547,2025,2046]" pageId="8" pageNumber="7">n</emphasis>
.
</taxonomicNameLabel>
, female
<typeStatus id="4A0688172E46FFF1FDF1F877FD7FF83C" box="[639,724,2025,2046]" pageId="8" pageNumber="7" type="paratype">paratype</typeStatus>
, metasoma and telson in lateral (21, 24), ventral (22, 25) and dorsal (23, 26) views.
<emphasis id="A7C9EAA72E46FFF1FF3BF79BFEEFF7D9" bold="true" box="[181,324,2053,2075]" pageId="8" pageNumber="7">FiGures 2123</emphasis>
. Under white light.
<emphasis id="A7C9EAA72E46FFF1FD8BF79BFD3FF7D9" bold="true" box="[517,660,2053,2075]" pageId="8" pageNumber="7">FiGures 2426</emphasis>
. Under UV light. Scale bar:
<quantity id="52459B502E46FFF1FC29F798FC46F7D9" box="[935,1005,2054,2075]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="7" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<paragraph id="950236B52E47FFF0FFE1F851FC5BF7C3" blockId="9.[111,1469,1999,2050]" pageId="9" pageNumber="8">
<emphasis id="A7C9EAA72E47FFF0FFE1F851FEABF826" bold="true" box="[111,256,1999,2021]" pageId="9" pageNumber="8">FiGures 2732</emphasis>
:
<taxonomicName id="52BD4D362E47FFF0FE80F851FE47F826" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[270,492,1999,2020]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="9" pageNumber="8" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E47FFF0FE80F851FE47F826" box="[270,492,1999,2020]" italics="true" pageId="9" pageNumber="8">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E47FFF0FE7AF851FD81F827" box="[500,554,1999,2021]" pageId="9" pageNumber="8" rank="species">
<emphasis id="A7C9EAA72E47FFF0FE7AF851FDA0F826" bold="true" box="[500,523,1999,2020]" pageId="9" pageNumber="8">sp</emphasis>
.
<emphasis id="A7C9EAA72E47FFF0FD96F851FD8EF826" bold="true" box="[536,549,1999,2020]" pageId="9" pageNumber="8">n</emphasis>
.
</taxonomicNameLabel>
, male
<typeStatus id="4A0688172E47FFF0FDFEF851FD6DF827" box="[624,710,1999,2021]" pageId="9" pageNumber="8" type="holotype">holotype</typeStatus>
, metasoma and telson in lateral (27, 30), ventral (28, 31) and dorsal (29, 32) views.
<emphasis id="A7C9EAA72E47FFF0FF3DF872FEE9F7C3" bold="true" box="[179,322,2028,2050]" pageId="9" pageNumber="8">FiGures 2729</emphasis>
. Under white light.
<emphasis id="A7C9EAA72E47FFF0FD8DF872FD39F7C3" bold="true" box="[515,658,2028,2050]" pageId="9" pageNumber="8">FiGures 3032</emphasis>
. Under UV light. Scale bar:
<quantity id="52459B502E47FFF0FC2BF872FC40F7C3" box="[933,1003,2028,2049]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="9" pageNumber="8" unit="mm" value="10.0">10 mm</quantity>
.
</paragraph>
<paragraph id="950236B52E44FFF3FFFAF924FDA2F8CB" blockId="10.[116,1475,1722,1801]" pageId="10" pageNumber="9">
<emphasis id="A7C9EAA72E44FFF3FFFAF924FEADF90D" bold="true" box="[116,262,1722,1744]" pageId="10" pageNumber="9">FiGures 3338</emphasis>
:
<taxonomicName id="52BD4D362E44FFF3FE9BF924FE5FF90D" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[277,500,1722,1743]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="10" pageNumber="9" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E44FFF3FE9BF924FE5FF90D" box="[277,500,1722,1743]" italics="true" pageId="10" pageNumber="9">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E44FFF3FE73F924FD9EF90D" box="[509,565,1722,1743]" pageId="10" pageNumber="9" rank="species">
<emphasis id="A7C9EAA72E44FFF3FE73F924FDBFF90D" bold="true" box="[509,532,1722,1743]" pageId="10" pageNumber="9">sp</emphasis>
.
<emphasis id="A7C9EAA72E44FFF3FDACF924FD84F90D" bold="true" box="[546,559,1722,1743]" pageId="10" pageNumber="9">n</emphasis>
.
</taxonomicNameLabel>
, female
<typeStatus id="4A0688172E44FFF3FD01F924FD4FF90D" box="[655,740,1722,1743]" pageId="10" pageNumber="9" type="paratype">paratype</typeStatus>
, pedipalp segments under white light. Chela ventral (33) and dorsal (33). Pedipalp ventral (35) and dorsal (36). Movable (37) and fiXed (38) fingers dentition. Trichobothrial pattern indicated by green circles. Scale bars:
<quantity id="52459B502E44FFF3FF27F96AFF47F8CB" box="[169,236,1780,1801]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="mm" value="10.0">10 mm</quantity>
(3334) and
<quantity id="52459B502E44FFF3FEE3F96AFE1BF8CB" box="[365,432,1780,1801]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="10" pageNumber="9" unit="mm" value="10.0">10 mm</quantity>
(3536).
</paragraph>
<paragraph id="950236B52E44FFF2FFFAF8AFFBECF83A" blockId="10.[116,774,1841,2025]" lastBlockId="11.[115,1473,1990,2041]" lastPageId="11" lastPageNumber="10" pageId="10" pageNumber="9">
decreased granules anteriorly. FiXed and movable fingers with 4 or 5 strong accessory denticles. Mesosoma: Tergites IVI matte, completely covered with fine granules in males; glossy and smooth, at most slightly shagreened laterally in females. Tergite VII finely granular with moderate, pointed granules posteriorly and laterally in males, but smooth anteriorly, granulate posteriorly and laterally in females. Sternites IIIVI wrinkled medially, almost smooth laterally in males, smooth and shiny in females with numerous moderate setae on the posterior margins. Sternite VII granular and feebly wrinkled medially; shagreened laterally with four tuberculated carinae with lobated granules; the posterior
<emphasis id="A7C9EAA72E45FFF2FFFDF858FEAAF819" bold="true" box="[115,257,1990,2012]" pageId="11" pageNumber="10">FiGures 3944</emphasis>
.
<taxonomicName id="52BD4D362E45FFF2FE82F859FE4CF81E" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[268,487,1991,2012]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="11" pageNumber="10" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E45FFF2FE82F859FE4CF81E" box="[268,487,1991,2012]" italics="true" pageId="11" pageNumber="10">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E45FFF2FE65F859FD8BF81E" box="[491,544,1991,2012]" pageId="11" pageNumber="10" rank="species">
<emphasis id="A7C9EAA72E45FFF2FE65F859FDA9F81E" bold="true" box="[491,514,1991,2012]" pageId="11" pageNumber="10">sp</emphasis>
.
<emphasis id="A7C9EAA72E45FFF2FD83F859FDB1F81E" bold="true" box="[525,538,1991,2012]" pageId="11" pageNumber="10">n</emphasis>
.
</taxonomicNameLabel>
, female
<typeStatus id="4A0688172E45FFF2FDFCF859FD6CF81E" box="[626,711,1991,2012]" pageId="11" pageNumber="10" type="paratype">paratype</typeStatus>
, pedipalp segments under UV light. Chela ventral (39) and dorsal (40). Pedipalp ventral (41) and dorsal (42). Movable (43) and fiXed (44) fingers dentition. Scale bar:
<quantity id="52459B502E45FFF2FC25F87DFC45F83B" box="[939,1006,2019,2041]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="11" pageNumber="10" unit="mm" value="10.0">10 mm</quantity>
(3940).
</paragraph>
<paragraph id="950236B52E42FFF5FFFDF9CBFE21F966" blockId="12.[115,1473,1621,1701]" pageId="12" pageNumber="11">
<emphasis id="A7C9EAA72E42FFF5FFFDF9CBFEAAF9A9" bold="true" box="[115,257,1621,1643]" pageId="12" pageNumber="11">FiGures 4550</emphasis>
:
<taxonomicName id="52BD4D362E42FFF5FE82F9C8FE4CF9A9" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[268,487,1622,1643]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="12" pageNumber="11" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E42FFF5FE82F9C8FE4CF9A9" box="[268,487,1622,1643]" italics="true" pageId="12" pageNumber="11">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E42FFF5FE65F9C8FD8BF9A9" box="[491,544,1622,1643]" pageId="12" pageNumber="11" rank="species">
<emphasis id="A7C9EAA72E42FFF5FE65F9C8FDA9F9A9" bold="true" box="[491,514,1622,1643]" pageId="12" pageNumber="11">sp</emphasis>
.
<emphasis id="A7C9EAA72E42FFF5FD83F9C8FDB1F9A9" bold="true" box="[525,538,1622,1643]" pageId="12" pageNumber="11">n</emphasis>
.
</taxonomicNameLabel>
, male
<typeStatus id="4A0688172E42FFF5FDD1F9C8FD1EF9AE" box="[607,693,1622,1644]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
, pedipalp segments under white light. Chela ventral (45) and dorsal (46). Pedipalp ventral (47) and dorsal (48). Movable (49) and fiXed (50) fingers dentition. Trichobothrial pattern indicated by green circles. Scale bars:
<quantity id="52459B502E42FFF5FAF0F9EDFA6BF94A" box="[1406,1472,1651,1672]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="11" unit="mm" value="10.0">10 mm</quantity>
(4546) and
<quantity id="52459B502E42FFF5FF63F911FE9BF967" box="[237,304,1679,1701]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="12" pageNumber="11" unit="mm" value="10.0">10 mm</quantity>
(4748).
</paragraph>
<paragraph id="950236B52E42FFF5FFFEF95DFB36F879" blockId="12.[112,771,1731,2044]" lastBlockId="12.[815,1473,1731,2043]" pageId="12" pageNumber="11">
margin bears numerous moderate setae. Metasoma: Segment I with 10 carinae, segment II-IV with 8, and segment V with 5 carinae. Lateral inframedian carinae on segment I indistinct. Ventral submedian and ventrolateral carinae moderate to strong, with moderate rounded granules, size and numbers of granules decrease from segment I toward segment IV, nearly obsolete on segment IV. Ventromedian and ventrolateral carinae strong with spaced large and pointed granules on segment V, pointed granules gradually increase posteriorly on ventrolateral carina of segment V. Telson globular in females and relatively slender in males. Pectinal teeth number
<quantity id="52459B502E42FFF5FC04F97DFC75F939" box="[906,990,1763,1787]" metricMagnitude="-1" metricUnit="m" metricValue="2.667" metricValueMax="3.048" metricValueMin="2.286" pageId="12" pageNumber="11" unit="in" value="10.5" valueMax="12.0" valueMin="9.0">912 in</quantity>
males,
<quantity id="52459B502E42FFF5FBB8F97DFB22F939" box="[1078,1161,1763,1787]" metricMagnitude="-1" metricUnit="m" metricValue="2.413" metricValueMax="2.794" metricValueMin="2.032" pageId="12" pageNumber="11" unit="in" value="9.5" valueMax="11.0" valueMin="8.0">811 in</quantity>
females. Genital operculum is heart-shaped in females, ovoid in males. TrichobothriotaXy of
<typeStatus id="4A0688172E42FFF5FCC0F8BAFCD7F8FE" box="[846,892,1828,1852]" pageId="12" pageNumber="11">type</typeStatus>
C, orthobothriotaXic (
<bibRefCitation id="F12C4B442E42FFF5FBFCF8BDFAA6F8F9" author="VACHON" box="[1138,1293,1827,1851]" pageId="12" pageNumber="11" pagination="857 - 958" refId="ref5949" refString="VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaXie en arachnologie. Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3 e ser., n ° 140, Zool., 104: 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
), pedipalp chela trichobothrium
<emphasis id="A7C9EAA72E42FFF5FC55F8DAFC41F899" box="[987,1002,1860,1883]" italics="true" pageId="12" pageNumber="11">it</emphasis>
placed basally along fiXed finger near
<emphasis id="A7C9EAA72E42FFF5FA2BF8DDFA11F899" box="[1445,1466,1859,1883]" italics="true" pageId="12" pageNumber="11">ib</emphasis>
. Tibial spurs absent. Tarsi armed only with prolateral pedal spurs, tarsi of legs I to IV with 4/7, 6/8, 7/8, 7/8 internal and eXternal spines arranged in series.
</paragraph>
<paragraph id="950236B52E42FFF5FCA1F85DFB4BF839" blockId="12.[815,1473,1731,2043]" pageId="12" pageNumber="11">
<emphasis id="A7C9EAA72E42FFF5FCA1F85DFC18F819" bold="true" box="[815,947,1987,2011]" pageId="12" pageNumber="11">DescriptiOn</emphasis>
: Based on male
<typeStatus id="4A0688172E42FFF5FB00F85DFB40F819" box="[1166,1259,1987,2011]" pageId="12" pageNumber="11" type="holotype">holotype</typeStatus>
(unless otherwise specified). Measurements are in
<tableCitation id="D83F030E2E42FFF5FB04F87DFB72F839" box="[1162,1241,2019,2043]" captionStart="Table 1" captionStartId="4.[119,175,983,1004]" captionText="Table 1. Comparative measurements of Scorpio granulomanus sp. n. male holotype and female paratype. Abbreviations: length (L), width (W), width anterior (Wa), width posterior (Wp), depth (D)." pageId="12" pageNumber="11">Table 1</tableCitation>
.
</paragraph>
<paragraph id="950236B52E43FFF4FFFFF97DFBEEF8D4" blockId="13.[113,1471,1763,1814]" pageId="13" pageNumber="12">
<emphasis id="A7C9EAA72E43FFF4FFFFF97DFF54F93A" bold="true" box="[113,255,1763,1785]" pageId="13" pageNumber="12">FiGures 5156</emphasis>
:
<taxonomicName id="52BD4D362E43FFF4FE82F97AFE43F93A" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[268,488,1763,1785]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E43FFF4FE82F97AFE43F93A" box="[268,488,1763,1785]" italics="true" pageId="13" pageNumber="12">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E43FFF4FE60F97AFD88F93B" box="[494,547,1764,1785]" pageId="13" pageNumber="12" rank="species">
<emphasis id="A7C9EAA72E43FFF4FE60F97AFDAEF93B" bold="true" box="[494,517,1764,1785]" pageId="13" pageNumber="12">sp</emphasis>
.
<emphasis id="A7C9EAA72E43FFF4FD9EF97AFDB6F93B" bold="true" box="[528,541,1764,1785]" pageId="13" pageNumber="12">n</emphasis>
.
</taxonomicNameLabel>
, male
<typeStatus id="4A0688172E43FFF4FDEAF97DFD11F93B" box="[612,698,1763,1785]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
, pedipalp segments under UV light. Chela ventral (51) and dorsal (52). Pedipalp ventral (53) and dorsal (54). Movable (55) and fiXed (56) fingers dentition. Scale bar:
<quantity id="52459B502E43FFF4FC27F89EFC47F8D4" box="[937,1004,1792,1814]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="12" unit="mm" value="10.0">10 mm</quantity>
(5152).
</paragraph>
<paragraph id="950236B52E43FFF4FFFFF8DEFA69F83A" blockId="13.[113,772,1856,2040]" lastBlockId="13.[816,1474,1856,2040]" pageId="13" pageNumber="12">
<emphasis id="A7C9EAA72E43FFF4FFFFF8DEFF41F89A" bold="true" box="[113,234,1856,1880]" pageId="13" pageNumber="12">COlOratiOn</emphasis>
(Figures 12). Carapace is light reddish brown with darker reticulations medially. Mesosoma: Tergites I-VI reddish brown with some reticulations medially but dark yellow laterally. Tergite VII reddish brown medially, dark yellow laterally and posteriorly. Sternites pale dark yellow. CoXae and sternum dark yellow. Genital operculum and pectines lustrous yellow. Pedipalp: Femur and patella dark yellow, dorsal carina of patella and granules reddish brown. Chela manus reddish yellow. Metasoma and legs dark yellow. The upper half of eXternal surface of chela manus and basal part of fiXed finger dark yellow, the lower half of eXternal surface of chela manus, the distal part of fiXed finger, and movable finger reddish
</paragraph>
<paragraph id="950236B52E40FFF7FFFBF845FA31F833" blockId="14.[117,1434,2011,2033]" box="[117,1434,2011,2033]" pageId="14" pageNumber="13">
<emphasis id="A7C9EAA72E40FFF7FFFBF845FEA8F832" bold="true" box="[117,259,2011,2033]" pageId="14" pageNumber="13">FiGures 5758</emphasis>
:
<taxonomicName id="52BD4D362E40FFF7FE9EF845FE47F832" authority="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr, 2023" authorityName="Al-Saraireh &amp; Yağmur &amp; Afifeh &amp; Amr" authorityYear="2023" box="[272,492,2011,2032]" class="Arachnida" family="Scorpionidae" genus="Scorpio" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="granulomanus" status="sp. nov.">
<emphasis id="A7C9EAA72E40FFF7FE9EF845FE47F832" box="[272,492,2011,2032]" italics="true" pageId="14" pageNumber="13">Scorpio granulomanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BCFA57DC2E40FFF7FE7FF845FD8DF832" box="[497,550,2011,2032]" pageId="14" pageNumber="13" rank="species">
<emphasis id="A7C9EAA72E40FFF7FE7FF845FDA3F832" bold="true" box="[497,520,2011,2032]" pageId="14" pageNumber="13">sp</emphasis>
.
<emphasis id="A7C9EAA72E40FFF7FD9AF845FD8AF832" bold="true" box="[532,545,2011,2032]" pageId="14" pageNumber="13">n</emphasis>
.
</taxonomicNameLabel>
, right legs IIV, retrolateral aspect.
<emphasis id="A7C9EAA72E40FFF7FC0FF845FC49F832" bold="true" box="[897,994,2011,2033]" pageId="14" pageNumber="13">FiGure 57</emphasis>
. Female
<typeStatus id="4A0688172E40FFF7FBB5F845FB3AF832" box="[1083,1169,2011,2032]" pageId="14" pageNumber="13" type="paratype">paratype</typeStatus>
.
<emphasis id="A7C9EAA72E40FFF7FB14F845FB50F832" bold="true" box="[1178,1275,2011,2033]" pageId="14" pageNumber="13">FiGure 58</emphasis>
. Male
<typeStatus id="4A0688172E40FFF7FAB1F845FA3DF833" box="[1343,1430,2011,2033]" pageId="14" pageNumber="13" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="950236B52E41FFF6FFFFFF75FE26FDE1" blockId="15.[113,773,235,2051]" pageId="15" pageNumber="14">brown. Ventrointernal surface of chela manus dark yellow to reddish brown. Granules reddish brown or black on apeX. Finger denticles and movable finger condyles black. Metasoma dark yellow, darker in ventral surface. Granules on dorsolateral, ventrolateral, ventral and ventral submedian carinae light reddish brown. Vesicle of telson dark yellow, aculeus reddish yellow at the base and dark reddish brown at the tip. Legs: dark yellow, spinules and condyles red or reddish black. Chelicerae: Chelicera manus dark yellow with brown reticulations, fingers brown, teeth reddish black.</paragraph>
<paragraph id="950236B52E41FFF6FFFFFDB4FE55FBC0" blockId="15.[113,773,235,2051]" pageId="15" pageNumber="14">
<emphasis id="A7C9EAA72E41FFF6FFFFFDB4FF41FD80" bold="true" box="[113,234,554,578]" pageId="15" pageNumber="14">Chelicerae</emphasis>
(Figures 17, 19). Cheliceral dentition characteristic for the family of
<taxonomicName id="52BD4D362E41FFF6FEAAFDD5FDF6FDA1" authority="(Vachon, 1963)" baseAuthorityName="Vachon" baseAuthorityYear="1963" box="[292,605,587,611]" class="Arachnida" family="Scorpionidae" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="family">
Scorpionidae (
<bibRefCitation id="F12C4B442E41FFF6FE30FDD5FDFEFDA1" author="VACHON" box="[446,597,587,611]" pageId="15" pageNumber="14" pagination="161 - 166" refId="ref5904" refString="VACHON, M. 1963. De l'utilite, en systematique, d'une nomenclature des dents des cheliceres chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 2 e ser., 35 (2): 161 - 166." type="journal article" year="1963">Vachon, 1963</bibRefCitation>
)
</taxonomicName>
; subdistal tooth and basal teeth conspicuous rather than on the movable finger.
<emphasis id="A7C9EAA72E41FFF6FFFFFD14FF7EFD60" bold="true" box="[113,213,650,674]" pageId="15" pageNumber="14">PrOsOma</emphasis>
(Figures 1720). Carapace anterior margin distinctly bilobed. Carapace acarinate, only posteriomedian carina visible, finely granular but triangular region between median eyes and anterior margin smooth with scattered moderate granules. Anterior margin with 13 medium to large setae and a distinct depression. There is a protrusion in the middle of the depression. Median ocular tubercle distinct and located in the center of the carapace. A pair of median eyes and three pairs of lateral eyes eXist. The first two lateral eyes in equal size, and the third slightly reduced. Anteriomedian furrow moderate but posterior median and posterior lateral furrows distinct. 6 moderate setae eXist on the carapace.
</paragraph>
<paragraph id="950236B52E41FFF6FFFFFB94FE81FA41" blockId="15.[113,773,235,2051]" pageId="15" pageNumber="14">
<emphasis id="A7C9EAA72E41FFF6FFFFFB94FF42FBE0" bold="true" box="[113,233,1034,1058]" pageId="15" pageNumber="14">MesOsOma</emphasis>
(Figures 912). Tergites IVI acarinate, matte, covered with fine granules completely in males. Tergite VII finely granular with moderate, pointed granules posteriorly and laterally; carinae granulate and present only in posterior half of segment, posterior margin of tergites with microsetae and a long seta on seventh tergite at end of each carina. Sternites IIIVI wrinkled medially, almost smooth laterally. Sternite VII granular and feebly wrinkled medially; shagreened laterally with four tuberculate carinae with lobated granules; posterior margin bears numerous moderate setae. Genital operculum ovoid. Pectines short. Stigmas linear, angled 45° and conspicuous.
</paragraph>
<paragraph id="950236B52E41FFF6FFFFFA14FD15F7C1" blockId="15.[113,773,235,2051]" pageId="15" pageNumber="14">
<emphasis id="A7C9EAA72E41FFF6FFFFFA14FECEFA60" bold="true" box="[113,357,1418,1443]" pageId="15" pageNumber="14">MetasOma and telsOn</emphasis>
(Figures 2732). Segment I with 10, segment II-IV with 8, and segment V with 5 carinae. Lateral inframedian carinae on segment I indistinct. Ventral submedian and ventrolateral carinae moderate to strong, with moderate rounded granules, size and numbers of granules decrease from segment I toward segment IV, nearly obsolete on segment IV. Ventromedian and ventrolateral carinae strong with spaced large and pointed granules on segment V, pointed granules gradually increase posteriorly on ventrolateral carina of segment V. Dorsolateral carinae strong on IIV with pointed and spaced, small granules; moderate on segment V with spinoid granules. Lateral supramedian carinae moderate with spaced granules on IIV. Segment V with a row of granules laterally. Intercarinal surfaces on segments I-V covered with fine granules on IIV, and with moderate granules on segment V. Segments IV covered moderately with red setae of variable size, denser on ventral surface. Vesicle globular, smooth dorsally, granulated ventrally and laterally; ventral aspect with numerous small, spinoid granules, vesicle with many red macrosetae even on the base of aculeus. Aculeus strongly curved and shorter than vesicle.
</paragraph>
<paragraph id="950236B52E41FFF6FCBEFF74FB37FC41" blockId="15.[816,1475,234,963]" pageId="15" pageNumber="14">
<emphasis id="A7C9EAA72E41FFF6FCBEFF74FC3EFEC0" bold="true" box="[816,917,234,258]" pageId="15" pageNumber="14">Pedipalp</emphasis>
(Figures 4556). DorsoeXternal carinae of pedipalp femur distinct with separate granules distally, sometimes with several rows of spaced granules, intercarinal surface smooth with numerous setae, dorsoeXternal surface with numerous rounded and pointed granules of different sizes, internal surface with some large, pointed granules. Patella with two carinae. Dorsomedian carina distinct with swollen and some fused granules and ventrointernal carinae obsolete with a few swollen granules, intercarinal surface smooth with numerous setae. Chela manus densely covered with large rounded and somewhat pointed granules, the posterior margin of the eXternal surface of chela manus bears very large spinoid protrusions. Internal surface of chela manus with small to moderate pointed granules along dorsal margin. VentroeXternal carina strong with dark rather fused granules, ventrointernal carina smooth. Chela manus flattened and elongated. Intercarinal surface smooth with numerous distinct setae. Chela fingers are relatively long with small pointed gradually decreased granules anteriorly. FiXed and movable fingers with 4 or 5 strong accessory denticles. TrichobothriotaXy of
<typeStatus id="4A0688172E41FFF6FAE1FCD5FA36FCA1" box="[1391,1437,843,867]" pageId="15" pageNumber="14">type</typeStatus>
C; orthobothriotaXic (
<bibRefCitation id="F12C4B442E41FFF6FC79FCF5FB3BFC41" author="VACHON" box="[1015,1168,875,899]" pageId="15" pageNumber="14" pagination="857 - 958" refId="ref5949" refString="VACHON, M. 1974. Etude des caracteres utilises pour classer les familles et les genres de Scorpions (Arachnides). 1. La trichobothriotaXie en arachnologie. Sigles trichobothriauX et types de trichobothriotaXie chez les Scorpions. Bulletin du Museum national d'Histoire naturelle, Paris, 3 e ser., n ° 140, Zool., 104: 857 - 958." type="journal article" year="1974">Vachon, 1974</bibRefCitation>
).
</paragraph>
<paragraph id="950236B52E41FFF6FCBEFC14FAE3FC01" blockId="15.[816,1475,234,963]" pageId="15" pageNumber="14">
<emphasis id="A7C9EAA72E41FFF6FCBEFC14FCCEFC60" bold="true" box="[816,869,906,930]" pageId="15" pageNumber="14">LeGs</emphasis>
(Figures 58). tarsi of right legs I to IV with 4/7, 6/8, 7/8, 7/8 internal and eXternal spines arranged in series.
</paragraph>
<paragraph id="950236B52E41FFF6FCBEFC74FB14F9A0" blockId="15.[816,1476,1002,1635]" pageId="15" pageNumber="14">
<emphasis id="A7C9EAA72E41FFF6FCBEFC74FBA3FBC0" bold="true" box="[816,1032,1002,1026]" pageId="15" pageNumber="14">Sexual dimOrphism</emphasis>
: Males differed from females as follows: (
<emphasis id="A7C9EAA72E41FFF6FCB7FB95FCEDFBE1" bold="true" box="[825,838,1035,1059]" pageId="15" pageNumber="14">a</emphasis>
) tergites IVI matte, completely covered with fine granules in males, but glossy and smooth in females; sternites IIIVI wrinkled medially in males, but smooth and shining in females; (
<emphasis id="A7C9EAA72E41FFF6FAFEFBD5FAD5FBA1" bold="true" box="[1392,1406,1099,1123]" pageId="15" pageNumber="14">b</emphasis>
) fiXed finger of pedipalp chela in adult females longer than in males; fiXed finger length to chela length ratio 0.370.40 with average
<quantity id="52459B502E41FFF6FCBEFB35FCD7FB01" box="[816,892,1195,1219]" metricMagnitude="-3" metricUnit="m" metricValue="9.906" pageId="15" pageNumber="14" unit="in" value="0.39">0.39 in</quantity>
males (n=7); 0.420.43 with average
<quantity id="52459B502E41FFF6FA92FB35FAC3FB01" box="[1308,1384,1195,1219]" metricMagnitude="-2" metricUnit="m" metricValue="1.0668" pageId="15" pageNumber="14" unit="in" value="0.42">0.42 in</quantity>
females (n=7); (
<emphasis id="A7C9EAA72E41FFF6FC0FFB55FC27FB21" bold="true" box="[897,908,1227,1251]" pageId="15" pageNumber="14">c</emphasis>
) vesicle of telson is more elongated in males than in females; vesicle length to width ratio 1.271.40 with average
<quantity id="52459B502E41FFF6FA1BFB75FCEFFAE1" metricMagnitude="-2" metricUnit="m" metricValue="3.3782" pageId="15" pageNumber="14" unit="in" value="1.33">1.33 in</quantity>
males (n=7), 1.231.26 with average
<quantity id="52459B502E41FFF6FB31FA95FAAEFAE1" box="[1215,1285,1291,1315]" metricMagnitude="-2" metricUnit="m" metricValue="3.1496" pageId="15" pageNumber="14" unit="in" value="1.24">1.24 in</quantity>
females (n=7); (
<emphasis id="A7C9EAA72E41FFF6FA25FA95FA12FAE1" bold="true" box="[1451,1465,1291,1315]" pageId="15" pageNumber="14">d</emphasis>
) genital operculum ovoid in males, but heart-shaped in females. (Figs. 14, 18); (
<emphasis id="A7C9EAA72E41FFF6FC44FAD5FC7EFAA1" bold="true" box="[970,981,1355,1379]" pageId="15" pageNumber="14">e</emphasis>
) pectinal teeth number in males 912 (18 combs) (1 comb with 9 teeth, 4 with 10, 12 with 11, 1 with 12), in females 8-11 (44 combs) (1 comb with 8 teeth, 12 with 9, 21 with 10, 10 with 11); average pectinal tooth count per pecten in males is 10.7 (18 combs, n=9),
<quantity id="52459B502E41FFF6FB98FA55FBE4FA21" box="[1046,1103,1483,1507]" metricMagnitude="-1" metricUnit="m" metricValue="2.5146" pageId="15" pageNumber="14" unit="in" value="9.9">9.9 in</quantity>
females (44 combs, n=22) (
<emphasis id="A7C9EAA72E41FFF6FAE2FA54FADFFA20" bold="true" box="[1388,1396,1482,1506]" pageId="15" pageNumber="14">f</emphasis>
) tergite VII finely granulate with moderate, pointed granules posteriorly and laterally in males (Fig. 11) but smooth anteriorly, granulate posteriorly and laterally in females (Fig. 7); (
<emphasis id="A7C9EAA72E41FFF6FB62F9B4FB52F980" bold="true" box="[1260,1273,1578,1602]" pageId="15" pageNumber="14">G</emphasis>
) in general, females are darker in color than males (Fig. 59).
</paragraph>
</subSubSection>
</treatment>
</document>