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<document id="8229139892CF3A334F60BD9FFA3B33C6" ID-DOI="10.3390/insects15050318" ID-Zenodo-Dep="11120833" IM.bibliography_approvedBy="tatiana" IM.illustrations_approvedBy="tatiana" IM.materialsCitations_approvedBy="tatiana" IM.metadata_approvedBy="tatiana" IM.taxonomicNames_approvedBy="tatiana" IM.treatments_approvedBy="tatiana" checkinTime="1714997800831" checkinUser="tatiana" docAuthor="Brazidec, Manuel, Lohrmann, Volker &amp; Perrichot, Vincent" docDate="2024" docId="195487B5937C4638FE3CFD67D583FBC0" docLanguage="en" docName="Insects.15.1-12.pdf" docOrigin="Insects (Clifton, N. J.) 15 (5)" docSource="http://dx.doi.org/10.3390/insects15050318" docTitle="Hukawngepyris setosus Brazidec &amp; Lohrmann &amp; Perrichot 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="7" masterDocId="E56DFFCD937F463EFF87FFDFD045FFA2" masterDocTitle="The First Cretaceous Epyrine Wasp (Hymenoptera: Bethylidae): A New Genus and Species from Early Cenomanian Kachin Amber" masterLastPageNumber="200" masterPageNumber="191" pageNumber="4" updateTime="1714998355944" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="83DFFC94F95A29A2D86D388C01DB06BD">The First Cretaceous Epyrine Wasp (Hymenoptera: Bethylidae): A New Genus and Species from Early Cenomanian Kachin Amber</mods:title>
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<mods:name id="DA17093072171C4BAD52BDC6352998BA" type="personal">
<mods:role id="A4CDD854743497627D69D6F9722F14AA">
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<mods:namePart id="74C530F166C4839E8F71D7ED65956B18">Brazidec, Manuel</mods:namePart>
<mods:affiliation id="520B6A2098DDF205D6BCEF03F378A216">CNRS, Géosciences Rennes, University Rennes, UMR 6118, 35000 Rennes, France &amp; Institut de Systématique, Évolution, Biodiversité (ISYEB), Muséum national dHistoire naturelle, CNRS, Sorbonne Université, EPHE, Université des Antilles, CP 50, 57 rue Cuvier, F- 75005 Paris, France &amp; State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology and Center for Excellence in Life and Paleoenvironment, Chinese Academy of Sciences, 39 East Beijing Road, Nanjing 210008, China</mods:affiliation>
<mods:nameIdentifier id="ED03ACBA5039E15A5B845748B96A14A6" type="email">manuel.brazidec@gmail.com</mods:nameIdentifier>
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<mods:namePart id="54FDC0D5F20DADF73362AA8371351DAD">Lohrmann, Volker</mods:namePart>
<mods:affiliation id="06EF0987E90AC42F59210EFA35B7D3A1">Übersee-Museum Bremen, Bahnhofsplatz 13, 28195 Bremen, Germany</mods:affiliation>
<mods:nameIdentifier id="1CA0F175713D6385B8E7EC8460F9B2A7" type="email">v.lohrmann@uebersee-museum.de</mods:nameIdentifier>
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<mods:namePart id="DF153153C3D02BAA60D81244A6DA16C6">Perrichot, Vincent</mods:namePart>
<mods:nameIdentifier id="2F01700FF4DC0DBBFB0736BEC44DA560" type="LSID">vincent.perrichot@univ-rennes.fr</mods:nameIdentifier>
<mods:nameIdentifier id="EAC1F7E50949FE174ACBBF354F8BA948" type="ORCID">0000-0002-7973-0430</mods:nameIdentifier>
<mods:affiliation id="BF7DA1CEA013AD5DEA1C190C14A35B01">CNRS, Géosciences Rennes, University Rennes, UMR 6118, 35000 Rennes, France</mods:affiliation>
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<mods:title id="04C11F44B106738FC03FF7EC45827A38">Insects</mods:title>
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<mods:part id="937D02BF2409221EDC7F5DC0D150239C">
<mods:date id="3D0210EB9D1E98108B759C51CC99FAD6">2024</mods:date>
<mods:detail id="8277EFE3CBD18D4DAE95C6680F18C231" type="series">
<mods:title id="066F6311AD7B26843D683DC19804FEB0">Clifton, N. J.</mods:title>
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<mods:detail id="DD5865E348527FAFFE1BBAAE6C0813EA" type="pubDate">
<mods:number id="930F7A153C9E849E9C63CA151BB69921">2024-04-30</mods:number>
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<mods:detail id="853B7A2EA66DD770DDDF4980A52A8386" type="volume">
<mods:number id="CB3B01012511DF1BA080ED2C32EE7FFF">15</mods:number>
</mods:detail>
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<mods:number id="D9CE685C14C13E15E1F0E40F536B3272">5</mods:number>
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<mods:start id="61A9F5B79BC0A1C471E0A91B3F61CA75">191</mods:start>
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<mods:url id="2857A279E13E354C8348CA11628AC537">http://dx.doi.org/10.3390/insects15050318</mods:url>
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<mods:identifier id="C1DC43E898A6B29B8A9EE588BC6E4B9C" type="DOI">10.3390/insects15050318</mods:identifier>
<mods:identifier id="F2DF15AD48899F62416C8CF180AA073E" type="Zenodo-Dep">11120833</mods:identifier>
</mods:mods>
<treatment id="195487B5937C4638FE3CFD67D583FBC0" ID-DOI="http://doi.org/10.5281/zenodo.11120815" ID-Zenodo-Dep="11120815" LSID="urn:lsid:plazi:treatment:195487B5937C4638FE3CFD67D583FBC0" httpUri="http://treatment.plazi.org/id/195487B5937C4638FE3CFD67D583FBC0" lastPageId="6" lastPageNumber="7" pageId="3" pageNumber="4">
<subSubSection id="D9E76528937C463DFE3CFD67D35EFD72" box="[443,795,696,720]" pageId="3" pageNumber="4" type="nomenclature">
<paragraph id="914236A3937C463DFE3CFD67D35EFD72" blockId="3.[443,1497,261,921]" box="[443,795,696,720]" pageId="3" pageNumber="4">
<taxonomicName id="56FD4D20937C463DFE3CFD67D2FFFD72" authority="Brazidec &amp; Lohrmann &amp; Perrichot, 2024" authorityName="Brazidec &amp; Lohrmann &amp; Perrichot" authorityYear="2024" box="[443,698,696,720]" class="Insecta" family="Bethylidae" genus="Hukawngepyris" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="setosus" status="sp. nov.">
<emphasis id="A389EAB1937C463DFE3CFD67D2FFFD72" box="[443,698,696,720]" italics="true" pageId="3" pageNumber="4">Hukawngepyris setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8BA57CA937C463DFD47FD66D35EFD72" box="[704,795,697,720]" pageId="3" pageNumber="4" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="D9E76528937C463DFE3BFD05D59CFD50" box="[444,1497,730,754]" pageId="3" pageNumber="4" type="description">
<paragraph id="914236A3937C463DFE3BFD05D59CFD50" blockId="3.[443,1497,261,921]" box="[444,1497,730,754]" pageId="3" pageNumber="4">https://zoobank.org/676B52A5-2D79-433C-B379-0D15A40BC31C (accessed on 25 April 2024).</paragraph>
</subSubSection>
<subSubSection id="D9E76528937C463DFE3BFD24D2EFFCF4" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="914236A3937C463DFE3BFD24D2EFFCF4" blockId="3.[443,1497,261,921]" pageId="3" pageNumber="4">
<emphasis id="A389EAB1937C463DFE3BFD24D26FFCB1" bold="true" box="[444,554,763,787]" pageId="3" pageNumber="4">Material.</emphasis>
<typeStatus id="4E468801937C463DFDB2FD24D2E3FCB1" box="[565,678,763,787]" pageId="3" pageNumber="4">Holotype</typeStatus>
number
<specimenCode id="C15B9ED8937C463DFC94FD24D3C8FCB1" box="[787,909,763,787]" pageId="3" pageNumber="4">MB.I.8638</specimenCode>
, a nearly complete
<specimenCount id="87FBFD2A937C463DFBF0FD24D48FFCB1" box="[1143,1226,763,787]" pageId="3" pageNumber="4" type="female">female</specimenCount>
, with right side of the first and second metasomal terga damaged and parts of the right midleg and right fore and hind wings missing.
</paragraph>
</subSubSection>
<subSubSection id="D9E76528937C463DFE3BFCBFD396FC3B" pageId="3" pageNumber="4" type="etymology">
<paragraph id="914236A3937C463DFE3BFCBFD396FC3B" blockId="3.[443,1497,261,921]" pageId="3" pageNumber="4">
<emphasis id="A389EAB1937C463DFE3BFCBFD206FCDA" bold="true" box="[444,579,864,888]" pageId="3" pageNumber="4">Etymology.</emphasis>
This species epithet refers to the conspicuous setation of the new species. The species epithet is to be treated as an adjective.
</paragraph>
</subSubSection>
<caption id="C582662B937C463DFE3BF8B4D2A5F801" ID-DOI="http://doi.org/10.5281/zenodo.11120837" ID-Zenodo-Dep="11120837" httpUri="https://zenodo.org/record/11120837/files/figure.png" pageId="3" pageNumber="4" startId="3.[444,514,1899,1921]" targetBox="[130,1456,950,1864]" targetPageId="3" targetType="figure">
<paragraph id="914236A3937C463DFE3BF8B4D2A5F801" blockId="3.[444,1495,1899,1955]" pageId="3" pageNumber="4">
<emphasis id="A389EAB1937C463DFE3BF8B4D25FF822" bold="true" box="[444,538,1899,1921]" pageId="3" pageNumber="4">Figure 2.</emphasis>
<taxonomicName id="56FD4D20937C463DFDA6F8B4D346F823" authority="Brazidec &amp; Lohrmann &amp; Perrichot, 2024" authorityName="Brazidec &amp; Lohrmann &amp; Perrichot" authorityYear="2024" box="[545,771,1899,1921]" class="Insecta" family="Bethylidae" genus="Hukawngepyris" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="setosus" status="gen. et sp. nov.">
<emphasis id="A389EAB1937C463DFDA6F8B4D346F823" box="[545,771,1899,1921]" italics="true" pageId="3" pageNumber="4">Hukawngepyris setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8BA57CA937C463DFC8EF8B3D3EDF823" box="[777,936,1900,1921]" pageId="3" pageNumber="4" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, female holotype, MB.I.8638, habitus in left lateroventral view. Scale bar = 1 mm.
</paragraph>
</caption>
<subSubSection id="D9E76528937C463DFE3BF818D2A4F842" box="[444,737,1991,2016]" pageId="3" pageNumber="4" type="diagnosis">
<paragraph id="914236A3937C463DFE3BF818D2A4F842" blockId="3.[443,1492,1991,2082]" box="[444,737,1991,2016]" pageId="3" pageNumber="4">
<emphasis id="A389EAB1937C463DFE3BF818D278F87D" bold="true" box="[444,573,1991,2015]" pageId="3" pageNumber="4">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection id="D9E76528937C463DFE3CF836D47EF780" pageId="3" pageNumber="4" type="materials_examined">
<paragraph id="914236A3937C463DFE3CF836D47EF780" blockId="3.[443,1492,1991,2082]" pageId="3" pageNumber="4">
<emphasis id="A389EAB1937C463DFE3CF836D284F7A3" bold="true" box="[443,705,2024,2049]" pageId="3" pageNumber="4">Type locality and age.</emphasis>
<materialsCitation id="21953CFE937C463DFD4EF836D47EF780" collectionCode="MB" country="Myanmar" location="Noije Bum" pageId="3" pageNumber="4" specimenCode="MB.I.8638" specimenCount="1" specimenCount-female="1" stateProvince="Kachin" typeStatus="holotype">
<location id="94226078937C463DFD4EF836D303F7A2" LSID="urn:lsid:plazi:treatment:195487B5937C4638FE3CFD67D583FBC0:94226078937C463DFD4EF836D303F7A2" box="[713,838,2025,2049]" country="Myanmar" name="Noije Bum" pageId="3" pageNumber="4" stateProvince="Kachin">Noije Bum</location>
,
<location id="94226078937C463DFCD6F836D457F7A3" LSID="urn:lsid:plazi:treatment:195487B5937C4638FE3CFD67D583FBC0:94226078937C463DFCD6F836D457F7A3" box="[849,1042,2025,2049]" country="Myanmar" name="Hukawng Valley" pageId="3" pageNumber="4" stateProvince="Kachin">Hukawng Valley</location>
,
<collectingRegion id="5339F841937C463DFB9AF836D42BF7A3" box="[1053,1134,2025,2049]" country="Myanmar" name="Kachin" pageId="3" pageNumber="4">Kachin</collectingRegion>
State, Northern
<collectingCountry id="E9EA7633937C463DFAACF836D5DAF7A3" box="[1323,1439,2025,2049]" name="Myanmar" pageId="3" pageNumber="4">Myanmar</collectingCountry>
; late Albianearly Cenomanian, ca. 99 Ma, mid-Cretaceous.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="D9E76528937B4638FE3BFEDAD583FBC0" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
<paragraph id="914236A3937B463AFE3BFEDAD320FE23" blockId="4.[444,1493,261,386]" pageId="4" pageNumber="5">
<emphasis id="A389EAB1937B463AFE3BFEDAD217FEBF" bold="true" box="[444,594,261,285]" pageId="4" pageNumber="5">Description.</emphasis>
As for the genus diagnosis, with the following emendation: Body stout, length 4.39 mm, with some metallic reflections. Head rugulose; anteromesoscutum minutely rugulose; metasoma smooth. Head and anterior mesosoma with setae longer than the distance between neighboring setae.
</paragraph>
<caption id="C582662B937B463AFE3BF8C3D417F83A" ID-DOI="http://doi.org/10.5281/zenodo.11120839" ID-Zenodo-Dep="11120839" httpUri="https://zenodo.org/record/11120839/files/figure.png" pageId="4" pageNumber="5" startId="4.[444,514,1820,1842]" targetBox="[108,1476,413,1780]" targetPageId="4" targetType="figure">
<paragraph id="914236A3937B463AFE3BF8C3D417F83A" blockId="4.[443,1491,1820,1945]" pageId="4" pageNumber="5">
<emphasis id="A389EAB1937B463AFE3BF8C3D25FF893" bold="true" box="[444,538,1820,1842]" pageId="4" pageNumber="5">Figure 3.</emphasis>
<taxonomicName id="56FD4D20937B463AFDA6F8C3D34CF890" authority="Brazidec &amp; Lohrmann &amp; Perrichot, 2024" authorityName="Brazidec &amp; Lohrmann &amp; Perrichot" authorityYear="2024" box="[545,777,1820,1842]" class="Insecta" family="Bethylidae" genus="Hukawngepyris" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="setosus" status="gen. et sp. nov.">
<emphasis id="A389EAB1937B463AFDA6F8C3D34CF890" box="[545,777,1820,1842]" italics="true" pageId="4" pageNumber="5">Hukawngepyris setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8BA57CA937B463AFC88F8C2D3F7F890" box="[783,946,1821,1842]" pageId="4" pageNumber="5" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, female holotype, MB.I.8638, (
<emphasis id="A389EAB1937B463AFB75F8C3D540F893" bold="true" box="[1266,1285,1820,1841]" pageId="4" pageNumber="5">A</emphasis>
) head in left lateral view; (
<emphasis id="A389EAB1937B463AFD85F8E0D257F8F6" bold="true" box="[514,530,1855,1876]" pageId="4" pageNumber="5">B</emphasis>
) head and partial mesosoma in dorsal view (black arrow: notaulus; white arrow: parapsidal signum); (
<emphasis id="A389EAB1937B463AFDADF8BED27EF8D5" bold="true" box="[554,571,1889,1911]" pageId="4" pageNumber="5">C</emphasis>
) mesosoma in left lateroventral view (black arrow: propleural epicoxal sulcus; white arrow: protibial spur). Scale bars: A = 0.25 mm, BC = 0.5 mm.
</paragraph>
</caption>
<caption id="C582662B937A463BFE3BF96CD48AF8AF" ID-DOI="http://doi.org/10.5281/zenodo.11120841" ID-Zenodo-Dep="11120841" httpUri="https://zenodo.org/record/11120841/files/figure.png" pageId="5" pageNumber="6" startId="5.[444,514,1715,1737]" targetBox="[109,1477,260,1674]" targetPageId="5" targetType="figure">
<paragraph id="914236A3937A463BFE3BF96CD48AF8AF" blockId="5.[443,1492,1715,1806]" pageId="5" pageNumber="6">
<emphasis id="A389EAB1937A463BFE3BF96CD259F96A" bold="true" box="[444,540,1715,1737]" pageId="5" pageNumber="6">Figure 4.</emphasis>
<taxonomicName id="56FD4D20937A463BFDACF96CD353F96B" authority="Brazidec &amp; Lohrmann &amp; Perrichot, 2024" authorityName="Brazidec &amp; Lohrmann &amp; Perrichot" authorityYear="2024" box="[555,790,1715,1737]" class="Insecta" family="Bethylidae" genus="Hukawngepyris" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="setosus" status="gen. et sp. nov.">
<emphasis id="A389EAB1937A463BFDACF96CD353F96B" box="[555,790,1715,1737]" italics="true" pageId="5" pageNumber="6">Hukawngepyris setosus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="B8BA57CA937A463BFC98F96BD396F96B" box="[799,979,1716,1737]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
, female holotype, MB.I.8638, (
<emphasis id="A389EAB1937A463BFAA6F96CD571F96A" bold="true" box="[1313,1332,1715,1736]" pageId="5" pageNumber="6">A</emphasis>
) mesosoma in dorsolateral view; (
<emphasis id="A389EAB1937A463BFD17F909D2E5F949" bold="true" box="[656,672,1750,1771]" pageId="5" pageNumber="6">B</emphasis>
) metasoma in left posterolateral view; (
<emphasis id="A389EAB1937A463BFBD3F90AD420F949" bold="true" box="[1108,1125,1749,1771]" pageId="5" pageNumber="6">C</emphasis>
) drawing of fore and hind wings (dashed lines indicate flexion lines; pt = pterostigma). Scale bars = 0.5 mm.
</paragraph>
</caption>
<paragraph id="914236A3937A4638FE3BF8EDD264FE01" blockId="5.[443,1495,1841,2066]" lastBlockId="6.[443,1496,261,1122]" lastPageId="6" lastPageNumber="7" pageId="5" pageNumber="6">
<emphasis id="A389EAB1937A463BFE3BF8EDD1B8F8E8" bold="true" box="[444,509,1842,1866]" pageId="5" pageNumber="6">Head</emphasis>
prognathous, ovoid in lateral view, slightly longer than wide; LH: 0.58 mm, WH: 0.46 mm; WF: 0.34 mm, HE: 0.31 mm; OOL: 0.19 mm; WOT: 0.18 mm; DAO: 0.06 mm; VOL: 0.13 mm; frons convex in lateral view; compound eye elliptical, protruding, located laterally on head, covering 0.6 × LH; median clypeal lobe slightly projecting forward, lateral lobe reduced, median clypeal carina not particularly developed, not continuing on frons; mandible 3 × longer than basal width, 4 short teeth of similar size; palpal formula 6:3; antenna inserted close to median clypeal carina; scape cylindrical, length 0.24 mm, 1.7 × longer than pedicel; pedicel pyriform, length 0.14 mm, base narrower than apex, shorter than flagellomere 1; flagellum with 11 flagellomeres, flagellomeres elongate, cylindrical, distinctly longer than wide (length 0.100.17 mm; width 0.040.05 mm), no conspicuous setation; flagellomere 11 longest, tapering at apex; ocellar triangle posteriad on head.
</paragraph>
<paragraph id="914236A393794638FE3BFE72D2A1FC5C" blockId="6.[443,1496,261,1122]" pageId="6" pageNumber="7">
<emphasis id="A389EAB193794638FE3BFE72D27BFE67" bold="true" box="[444,574,429,453]" pageId="6" pageNumber="7">Mesosoma</emphasis>
shorter than metasoma, length 1.12 mm; propleuron barely visible in dorsal view; prosternum reduced; dorsal pronotal area slender (length 0.28 mm), without carina, lateral pronotal margins slightly incurved, posterior margin almost straight; anteromesoscutum slightly shorter than dorsal pronotal area (length 0.24 mm), notauli deeply impressed, converging posterad, posteriorly separated by width of sulcus of mesoscuto-mesoscutellar suture; sulcus of mesoscuto-mesoscutellar suture reniform; metanotum almost not visible; metapectalpropodeal complex wider than long, dorsally carinate, metapostnotal-propodeal suture present, short; propodeal declivity concave, long.
<emphasis id="A389EAB193794638FB5AFD48D515FD0D" bold="true" box="[1245,1360,663,687]" pageId="6" pageNumber="7">Forewing</emphasis>
developed (LFW: 1.62 mm), hyaline, homogeneously micro-pubescent; pterostigma elongate, 3.9 × longer than wide, narrow; 2r-rs&amp;Rs originating from distal half of pterostigma and widely arched; costal [C], radial [R] and cubital [Cu] cells closed; 1Rs slightly angled at junction with 1M (both veins of similar length); 1M evenly incurved; 1Rs straight; no other tubular veins present but patterns of flexion lines visible in the distal section of the wing.
<emphasis id="A389EAB193794638FA13FCE1D1BCFCDA" bold="true" pageId="6" pageNumber="7">Hind wing</emphasis>
hyaline, homogeneously micro-pubescent, 1.01 mm long; Sc + R and A veins tubular, Sc + R reaching half the distance to distal hamuli, anal vein length about 0.25 × length of clavus of hind wing.
<emphasis id="A389EAB193794638FD5FFC7CD355FC19" bold="true" box="[728,784,931,955]" pageId="6" pageNumber="7">Legs</emphasis>
particularly pubescent, slender; tibial spurs formula 112; protibial spur with bifurcate apex; tarsal claws slightly curved, at least metatarsal claw with one subapical tooth.
</paragraph>
<paragraph id="914236A393794638FE3BFBD8D583FBC0" blockId="6.[443,1496,261,1122]" pageId="6" pageNumber="7">
<emphasis id="A389EAB193794638FE3BFBD8D27CFBBD" bold="true" box="[444,569,1031,1055]" pageId="6" pageNumber="7">Metasoma</emphasis>
smooth, fusiform, length 1.23 mm (indicative), posterior segments partly retracted; six terga visible; petiole short; length of measurable terga: T1 = 0.24 mm; T2 = 0.35 mm; T3 = 0.15); metasomal sterna with few isolated erect bristles; apex of sting exserted.
</paragraph>
</subSubSection>
</treatment>
</document>