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<document id="982C6376415B20BA10E568212BE29285" ID-CLB-Dataset="55711" ID-DOI="10.11646/zootaxa.5133.3.2" ID-GBIF-Dataset="637a139f-7734-4d00-b27a-9e01d794cb2e" ID-ISSN="1175-5326" ID-Zenodo-Dep="6522526" ID-ZooBank="7A7C9379-36FA-4BEE-9031-3ADDD7D1561F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651756113647" checkinUser="plazi" docAuthor="Makbun, Noppadon" docDate="2022" docId="0D2387CD8E359F40009B3F7E7B96F842" docLanguage="en" docName="zootaxa.5133.3.2.pdf" docOrigin="Zootaxa 5133 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Macromia siamensis Makbun 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="352" masterDocId="F11AFFB58E379F46000C3E117E0FFF97" masterDocTitle="Macromia siamensis, a new species from North Thailand (Odonata: Anisoptera Macromiidae)" masterLastPageNumber="354" masterPageNumber="346" pageNumber="348" updateTime="1699329833206" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="23622727FD9A92AF639F131FE9C310D9">Macromia siamensis, a new species from North Thailand (Odonata: Anisoptera Macromiidae)</mods:title>
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<treatment id="0D2387CD8E359F40009B3F7E7B96F842" ID-DOI="http://doi.org/10.5281/zenodo.6522562" ID-GBIF-Taxon="195192072" ID-Zenodo-Dep="6522562" LSID="urn:lsid:plazi:treatment:0D2387CD8E359F40009B3F7E7B96F842" httpUri="http://treatment.plazi.org/id/0D2387CD8E359F40009B3F7E7B96F842" lastPageId="6" lastPageNumber="352" pageId="2" pageNumber="348">
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<emphasis id="B7FEEAC98E359F44009B3F7E7FF9FE1E" bold="true" box="[151,502,367,393]" pageId="2" pageNumber="348">
<taxonomicName id="428A4D588E359F44009B3F7E7F9EFE1E" authority="Makbun, 2022" authorityName="Makbun" authorityYear="2022" box="[151,401,367,393]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="348" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="B7FEEAC98E359F44009B3F7E7F9EFE1E" bold="true" box="[151,401,367,393]" italics="true" pageId="2" pageNumber="348">Macromia siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACCD57B28E359F4401953F7E7FF9FE1E" box="[409,502,367,393]" pageId="2" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="853536DB8E359F44009B3F827F10FE3A" blockId="2.[151,502,367,429]" box="[151,287,403,429]" pageId="2" pageNumber="348">
(
<figureCitation id="1DB12A5E8E359F4400933F827F19FE3A" box="[159,278,403,429]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 27" captionStart-2="FIGURES 811" captionStartId-0="2.[151,250,1657,1681]" captionStartId-1="3.[151,264,1906,1930]" captionStartId-2="4.[151,264,1370,1394]" captionTargetBox-0="[151,1436,774,1631]" captionTargetBox-1="[151,1436,597,1883]" captionTargetBox-2="[151,1436,489,1346]" captionTargetId-0="figure-196@2.[151,1436,774,1631]" captionTargetId-1="figure-235@3.[151,1436,597,1883]" captionTargetId-2="figure-153@4.[151,1436,489,1346]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. General habitus of Macromia siamensis sp. nov. holotype. Bar = 10 mm." captionText-1="FIGURES 27. Morphological features of Macromia siamensis sp. nov. holotype: (2) head, frontal view; (3) head, dorsal view; (4) synthorax, oblique frontal view; (5) synthorax, lateral view; (6) secondary genitalia, lateral view; (7) fore- and hindwing, lateral view." captionText-2="FIGURES 811. Abdomen (8, 10) and caudal appendages (9, 11) of Macromia siamensis sp. nov. holotype: (8, 9) lateral view; (10, 11) dorsal view." figureDoi-0="http://doi.org/10.5281/zenodo.6522528" figureDoi-1="http://doi.org/10.5281/zenodo.6522530" figureDoi-2="http://doi.org/10.5281/zenodo.6522532" httpUri-0="https://zenodo.org/record/6522528/files/figure.png" httpUri-1="https://zenodo.org/record/6522530/files/figure.png" httpUri-2="https://zenodo.org/record/6522532/files/figure.png" pageId="2" pageNumber="348">Figs. 111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CD9065508E359F44009B3FCA7A7FFDAA" pageId="2" pageNumber="348" type="materials_examined">
<paragraph id="853536DB8E359F44009B3FCA7EFDFD8E" blockId="2.[151,1437,474,753]" pageId="2" pageNumber="348">
<materialsCitation id="35E23C868E359F44009B3FCA7EFDFD8E" ID-GBIF-Occurrence="3763084301" collectingDate="2015-03-13" collectorName="N. Makbun" country="Thailand" elevation="580" location="Ban Luang" municipality="Chom Thong" pageId="2" pageNumber="348" specimenCount="1" specimenCount-male="1" stateProvince="Chiang Mai" typeStatus="holotype">
<emphasis id="B7FEEAC98E359F44009B3FCA7F04FE62" bold="true" box="[151,267,475,501]" pageId="2" pageNumber="348">
<typeStatus id="5A3188798E359F44009B3FCA7F09FE62" box="[151,262,475,501]" pageId="2" pageNumber="348" type="holotype">Holotype</typeStatus>
.
</emphasis>
<specimenCount id="938CFD528E359F44011F3FCA7F32FE63" box="[275,317,475,501]" pageId="2" pageNumber="348" type="male">1 ♂</specimenCount>
,
<collectingCountry id="FD9D764B8E359F4401473FCA7FBEFE62" box="[331,433,475,501]" name="Thailand" pageId="2" pageNumber="348">Thailand</collectingCountry>
,
<collectingRegion id="474EF8398E359F4401B23FCA7CBFFE62" box="[446,688,475,501]" country="Thailand" name="Chiang Mai" pageId="2" pageNumber="348">Chiang Mai province</collectingRegion>
,
<collectingMunicipality id="6551ACA18E359F4402B73FCA7D5CFE62" box="[699,851,475,501]" pageId="2" pageNumber="348">Chom Thong</collectingMunicipality>
,
<location id="805560008E359F4403533FCA7DD4FE62" LSID="urn:lsid:plazi:treatment:0D2387CD8E359F40009B3F7E7B96F842:805560008E359F4403533FCA7DD4FE62" box="[863,987,475,501]" country="Thailand" municipality="Chom Thong" name="Ban Luang" pageId="2" pageNumber="348" stateProvince="Chiang Mai">Ban Luang</location>
(altitude
<quantity id="42729B3E8E359F44044B3FCB7A9DFE62" box="[1095,1170,474,501]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="348" unit="m" value="580.0">
<elevation id="0EA7D1E88E359F44044B3FCB7A9DFE62" box="[1095,1170,474,501]" metricMagnitude="2" metricUnit="m" metricValue="5.8" pageId="2" pageNumber="348" unit="m" value="580.0">580 m</elevation>
</quantity>
),
<date id="F134101B8E359F4404AB3FCB7B3EFE62" box="[1191,1329,474,501]" pageId="2" pageNumber="348" value="2015-03-13">
<collectingDate id="E170E9F38E359F4404AB3FCB7B3EFE62" box="[1191,1329,474,501]" pageId="2" pageNumber="348" value="2015-03-13">13iii2015</collectingDate>
</date>
,
<collectorName id="287F530D8E359F4405313FCA7ECEFD8E" pageId="2" pageNumber="348">N. Makbun</collectorName>
leg.
</materialsCitation>
</paragraph>
<paragraph id="853536DB8E359F44009B3C327A7FFDAA" blockId="2.[151,1437,474,753]" box="[151,1136,547,573]" pageId="2" pageNumber="348">
<emphasis id="B7FEEAC98E359F44009B3C327F19FDAA" bold="true" box="[151,278,547,573]" pageId="2" pageNumber="348">
<typeStatus id="5A3188798E359F44009B3C327F1EFDAA" box="[151,273,547,573]" pageId="2" pageNumber="348" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="938CFD528E359F4401113C327F53FDAB" box="[285,348,547,573]" pageId="2" pageNumber="348" type="male">2 ♂♂</specimenCount>
,
<date id="F134101B8E359F4401663C327FFBFDAA" box="[362,500,547,573]" pageId="2" pageNumber="348" value="2013-03-14">14iii2013</date>
, locality, data and collector the same as for the
<typeStatus id="5A3188798E359F44040A3C327A63FDAA" box="[1030,1132,547,573]" pageId="2" pageNumber="348" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="CD9065508E359F4400CB3C567F19FD66" pageId="2" pageNumber="348" type="etymology">
<paragraph id="853536DB8E359F4400CB3C567F19FD66" blockId="2.[151,1437,474,753]" pageId="2" pageNumber="348">
<emphasis id="B7FEEAC98E359F4400CB3C567F41FDF6" bold="true" box="[199,334,583,609]" pageId="2" pageNumber="348">Etymology.</emphasis>
The new species is named after Siamensis, a Thai non-profit organization, in recognition of their continued support for my work, and in honor of
<collectingCountry id="FD9D764B8E359F4402B53C7A7CFAFD12" box="[697,757,619,645]" name="Thailand" pageId="2" pageNumber="348">Siam</collectingCountry>
, the former name of
<collectingCountry id="FD9D764B8E359F4403EE3C7A7A47FD12" box="[994,1096,619,645]" name="Thailand" pageId="2" pageNumber="348">Thailand</collectingCountry>
, the country in which the new taxon was found. Siamensis is a group of taxonomists in various fields and keen enthusiasts who contribute and give public awareness on biodiversity and the environment in
<collectingCountry id="FD9D764B8E359F4403243CA27D80FD5A" box="[808,911,691,717]" name="Thailand" pageId="2" pageNumber="348">Thailand</collectingCountry>
. Also, siamensis is a Latin adjective derived from
<collectingCountry id="FD9D764B8E359F4400D93CC67F1DFD66" box="[213,274,727,753]" name="Thailand" pageId="2" pageNumber="348">Siam</collectingCountry>
.
</paragraph>
</subSubSection>
<caption id="D1F566538E359F44009B38687DF2F907" ID-DOI="http://doi.org/10.5281/zenodo.6522528" ID-Zenodo-Dep="6522528" box="[151,1021,1657,1681]" httpUri="https://zenodo.org/record/6522528/files/figure.png" pageId="2" pageNumber="348" startId="2.[151,250,1657,1681]" targetBox="[151,1436,774,1631]" targetPageId="2">
<paragraph id="853536DB8E359F44009B38687DF2F907" blockId="2.[151,1021,1657,1681]" box="[151,1021,1657,1681]" pageId="2" pageNumber="348">
<emphasis id="B7FEEAC98E359F44009B38687F1CF906" bold="true" box="[151,275,1657,1681]" pageId="2" pageNumber="348">FIGURE 1.</emphasis>
General habitus of
<taxonomicName id="428A4D588E359F4401D7386B7CA2F906" authority="Makbun, 2022" authorityName="Makbun" authorityYear="2022" box="[475,685,1658,1681]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="348" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="B7FEEAC98E359F4401D7386B7CA2F906" box="[475,685,1658,1681]" italics="true" pageId="2" pageNumber="348">Macromia siamensis</emphasis>
</taxonomicName>
<emphasis id="B7FEEAC98E359F4402BF38687D0CF906" bold="true" box="[691,771,1657,1681]" pageId="2" pageNumber="348">
<taxonomicNameLabel id="ACCD57B28E359F4402BF38687D0CF906" box="[691,771,1657,1681]" pageId="2" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype. Bar = 10 mm.
</paragraph>
</caption>
<subSubSection id="CD9065508E359F4200CB38AE7F98F921" lastPageId="4" lastPageNumber="350" pageId="2" pageNumber="348" type="description">
<paragraph id="853536DB8E359F4400CB38AE7FD1F94E" blockId="2.[151,1437,1727,2042]" box="[199,478,1727,1753]" pageId="2" pageNumber="348">
<emphasis id="B7FEEAC98E359F4400CB38AE7FD1F94E" bold="true" box="[199,478,1727,1753]" pageId="2" pageNumber="348">
Description of
<typeStatus id="5A3188798E359F44017B38AE7FD1F94E" box="[375,478,1727,1753]" pageId="2" pageNumber="348" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="853536DB8E359F4400CB38F27B27F8FE" blockId="2.[151,1437,1727,2042]" pageId="2" pageNumber="348">
<emphasis id="B7FEEAC98E359F4400CB38F27F0BF96A" box="[199,260,1763,1789]" italics="true" pageId="2" pageNumber="348">Head</emphasis>
(
<figureCitation id="1DB12A5E8E359F44011F38F27F71F96A" box="[275,382,1763,1789]" captionStart-0="FIGURE 1" captionStart-1="FIGURES 27" captionStartId-0="2.[151,250,1657,1681]" captionStartId-1="3.[151,264,1906,1930]" captionTargetBox-0="[151,1436,774,1631]" captionTargetBox-1="[151,1436,597,1883]" captionTargetId-0="figure-196@2.[151,1436,774,1631]" captionTargetId-1="figure-235@3.[151,1436,597,1883]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. General habitus of Macromia siamensis sp. nov. holotype. Bar = 10 mm." captionText-1="FIGURES 27. Morphological features of Macromia siamensis sp. nov. holotype: (2) head, frontal view; (3) head, dorsal view; (4) synthorax, oblique frontal view; (5) synthorax, lateral view; (6) secondary genitalia, lateral view; (7) fore- and hindwing, lateral view." figureDoi-0="http://doi.org/10.5281/zenodo.6522528" figureDoi-1="http://doi.org/10.5281/zenodo.6522530" httpUri-0="https://zenodo.org/record/6522528/files/figure.png" httpUri-1="https://zenodo.org/record/6522530/files/figure.png" pageId="2" pageNumber="348">Figs. 13</figureCitation>
). Face black with yellow pattern. Compound eyes dark brown. Vertex with two pointed metallic black protuberances. Antenna black. Postfrons yellow with black upper margin and with a groove in the middle. Antefrons metallic black with yellow lower margin. Postclypeus wholly yellow. Anteclypeus black with both sides pale. Labrum and base of mandible black. Labium yellow with darkened upper part on the inner margin.
</paragraph>
<paragraph id="853536DB8E359F4500CB39627FE2FF6E" blockId="2.[151,1437,1727,2042]" lastBlockId="3.[151,1437,151,573]" lastPageId="3" lastPageNumber="349" pageId="2" pageNumber="348">
<emphasis id="B7FEEAC98E359F4400CB39627F17F81A" box="[199,280,1907,1933]" italics="true" pageId="2" pageNumber="348">Thorax</emphasis>
(
<figureCitation id="1DB12A5E8E359F44012439627F99F819" box="[296,406,1907,1934]" captionStart="FIGURES 27" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure-235@3.[151,1436,597,1883]" captionTargetPageId="3" captionText="FIGURES 27. Morphological features of Macromia siamensis sp. nov. holotype: (2) head, frontal view; (3) head, dorsal view; (4) synthorax, oblique frontal view; (5) synthorax, lateral view; (6) secondary genitalia, lateral view; (7) fore- and hindwing, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6522530" httpUri="https://zenodo.org/record/6522530/files/figure.png" pageId="2" pageNumber="348">Figs. 45</figureCitation>
). Prothorax black with yellow markings, as follows: middle lobe with lateral spot, posterior lobe entirely black. Synthorax black with iridescent blue reflection and covered with white setae. Antealar sinus yellow. Mesepisternum with broad, dorsally tapering antehumeral stripe and extended throughout its length, upper margin almost touching antealar sinus, lower margin running down across mesokatepisternum. Broad yellow lateral stripe cover metepisternum and metastigma except for upper and posterior margins. Posterior margin of metepimeron yellow, occupying about one fourth of its area. Poststernum is entire yellow. Leg black except for coxae largely yellow. Tibial keels yellowish.
</paragraph>
<paragraph id="853536DB8E349F4500CB3F127B89FE1E" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="349">
<emphasis id="B7FEEAC98E349F4500CB3F127F04FE8B" box="[199,267,259,284]" italics="true" pageId="3" pageNumber="349">Wings</emphasis>
hyaline with dark veins (
<figureCitation id="1DB12A5E8E349F4502253F127C62FE8A" box="[553,621,259,285]" captionStart="FIGURES 27" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure-235@3.[151,1436,597,1883]" captionTargetPageId="3" captionText="FIGURES 27. Morphological features of Macromia siamensis sp. nov. holotype: (2) head, frontal view; (3) head, dorsal view; (4) synthorax, oblique frontal view; (5) synthorax, lateral view; (6) secondary genitalia, lateral view; (7) fore- and hindwing, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6522530" httpUri="https://zenodo.org/record/6522530/files/figure.png" pageId="3" pageNumber="349">Fig. 7</figureCitation>
). Ax 13 on FW, 8 on HW. Px 6 (right), 7 (left) on FW, 8 (right), 10 (left) on HW. The anal angle of the wing rather rounded. Anal triangle with one crossvein. Anal loop 5-celled, without central cell. Two crossveins in cubital space. Discoidal field widening from one to ten cells in both wings. Triangles in both wings without crossvein. Supratriangles with 2 (right) 3 (left) cells in FW, and one cell in HW. Pt black.
</paragraph>
<paragraph id="853536DB8E349F4500CB3F827B28FDAA" blockId="3.[151,1437,151,573]" pageId="3" pageNumber="349">
<emphasis id="B7FEEAC98E349F4500CB3F827F3EFE3A" box="[199,305,403,429]" italics="true" pageId="3" pageNumber="349">Abdomen</emphasis>
black with yellow markings (
<figureCitation id="1DB12A5E8E349F4502883F827CF0FE3A" box="[644,767,403,429]" captionStart="FIGURES 811" captionStartId="4.[151,264,1370,1394]" captionTargetBox="[151,1436,489,1346]" captionTargetId="figure-153@4.[151,1436,489,1346]" captionTargetPageId="4" captionText="FIGURES 811. Abdomen (8, 10) and caudal appendages (9, 11) of Macromia siamensis sp. nov. holotype: (8, 9) lateral view; (10, 11) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6522532" httpUri="https://zenodo.org/record/6522532/files/figure.png" pageId="3" pageNumber="349">Figs. 8, 10</figureCitation>
) as follows: S1 with large anterolateral spot, S2 with basal ring covering auricle, S3 with large middorsal spot and anterolateral spot separated from each other, S46 each with a large basal ring, S7 with basal ring connected to irregular markings at some points. S8 with dorsal irregular triangle, and a ventrolateral spot extending to basal half. S9 with anterolateral spot. The posterior margin of S9 pointed when seen laterally. S10 entirely black, its dorsal outline slightly arched but do not form a protuberance.
</paragraph>
<caption id="D1F566538E349F45009B39637F1BF845" ID-DOI="http://doi.org/10.5281/zenodo.6522530" ID-Zenodo-Dep="6522530" httpUri="https://zenodo.org/record/6522530/files/figure.png" pageId="3" pageNumber="349" startId="3.[151,264,1906,1930]" targetBox="[151,1436,597,1883]" targetPageId="3">
<paragraph id="853536DB8E349F45009B39637F1BF845" blockId="3.[151,1437,1906,2002]" pageId="3" pageNumber="349">
<emphasis id="B7FEEAC98E349F45009B39637F35F81D" bold="true" box="[151,314,1906,1930]" pageId="3" pageNumber="349">FIGURES 27.</emphasis>
Morphological features of
<taxonomicName id="428A4D588E349F45024739627D12F81D" authority="Makbun, 2022" authorityName="Makbun" authorityYear="2022" box="[587,797,1907,1930]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="349" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="B7FEEAC98E349F45024739627D12F81D" box="[587,797,1907,1930]" italics="true" pageId="3" pageNumber="349">Macromia siamensis</emphasis>
</taxonomicName>
<emphasis id="B7FEEAC98E349F45032E39637D7EF81D" bold="true" box="[802,881,1906,1930]" pageId="3" pageNumber="349">
<taxonomicNameLabel id="ACCD57B28E349F45032E39637D7EF81D" box="[802,881,1906,1930]" pageId="3" pageNumber="349" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype: (2) head, frontal view; (3) head, dorsal view; (4) synthorax, oblique frontal view; (5) synthorax, lateral view; (6) secondary genitalia, lateral view; (7) fore- and hindwing, lateral view.
</paragraph>
</caption>
<paragraph id="853536DB8E339F4200CB3E867F9FFEF2" blockId="4.[151,1437,151,465]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4200CB3E867FA7FF26" box="[199,424,151,177]" italics="true" pageId="4" pageNumber="350">Caudal appendages</emphasis>
blackish brown, pale at the base, longer than S10 (
<figureCitation id="1DB12A5E8E339F4203E03E867A69FF26" box="[1004,1126,151,177]" captionStart="FIGURES 811" captionStartId="4.[151,264,1370,1394]" captionTargetBox="[151,1436,489,1346]" captionTargetId="figure-153@4.[151,1436,489,1346]" captionTargetPageId="4" captionText="FIGURES 811. Abdomen (8, 10) and caudal appendages (9, 11) of Macromia siamensis sp. nov. holotype: (8, 9) lateral view; (10, 11) dorsal view." figureDoi="http://doi.org/10.5281/zenodo.6522532" httpUri="https://zenodo.org/record/6522532/files/figure.png" pageId="4" pageNumber="350">Figs. 9, 11</figureCitation>
). Cerci slightly longer than paraprocts. In lateral view, cerci broad basally and slightly tapering to its apex, ventral margin of posterior half possesses small blunt spines. Their tips acute and pointing upward. Paraprocts curved, broadest at middle, with pointed apex, directed inward. In dorsal view, cerci slightly diverge from base and gently curved inward at their half-length, their outer margin furnished with lateral spines with tips pointing outward. Paraprocts triangle-shaped with a small hook pointing inward.
</paragraph>
<paragraph id="853536DB8E339F4200CB3F7E7D93FE46" blockId="4.[151,1437,151,465]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4200CB3F7E7FA7FE1E" box="[199,424,367,393]" italics="true" pageId="4" pageNumber="350">Secondary genitalia</emphasis>
(
<figureCitation id="1DB12A5E8E339F4201BA3F7E7FF6FE1D" box="[438,505,367,394]" captionStart="FIGURES 27" captionStartId="3.[151,264,1906,1930]" captionTargetBox="[151,1436,597,1883]" captionTargetId="figure-235@3.[151,1436,597,1883]" captionTargetPageId="3" captionText="FIGURES 27. Morphological features of Macromia siamensis sp. nov. holotype: (2) head, frontal view; (3) head, dorsal view; (4) synthorax, oblique frontal view; (5) synthorax, lateral view; (6) secondary genitalia, lateral view; (7) fore- and hindwing, lateral view." figureDoi="http://doi.org/10.5281/zenodo.6522530" httpUri="https://zenodo.org/record/6522530/files/figure.png" pageId="4" pageNumber="350">Fig. 6</figureCitation>
). Vesicle triangle-shaped, covered densely with thick short setae. Posterior hamulus basally pale and dark apically, broad at base and distinctly tapered to its apex furnished with a hook on its posterior side and expanded on its anterior side, forming a hammer-like shape.
</paragraph>
<caption id="D1F566538E339F42009B3B4B7F67FA01" ID-DOI="http://doi.org/10.5281/zenodo.6522532" ID-Zenodo-Dep="6522532" httpUri="https://zenodo.org/record/6522532/files/figure.png" pageId="4" pageNumber="350" startId="4.[151,264,1370,1394]" targetBox="[151,1436,489,1346]" targetPageId="4">
<paragraph id="853536DB8E339F42009B3B4B7F67FA01" blockId="4.[151,1436,1370,1430]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F42009B3B4B7F49FAE5" bold="true" box="[151,326,1370,1394]" pageId="4" pageNumber="350">FIGURES 811.</emphasis>
Abdomen (8, 10) and caudal appendages (9, 11) of
<taxonomicName id="428A4D588E339F4203433B4A7A2FFAE5" authority="Makbun, 2022" authorityName="Makbun" authorityYear="2022" box="[847,1056,1371,1394]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="350" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="B7FEEAC98E339F4203433B4A7A2FFAE5" box="[847,1056,1371,1394]" italics="true" pageId="4" pageNumber="350">Macromia siamensis</emphasis>
</taxonomicName>
<emphasis id="B7FEEAC98E339F42042A3B4B7A79FAE5" bold="true" box="[1062,1142,1370,1394]" pageId="4" pageNumber="350">
<taxonomicNameLabel id="ACCD57B28E339F42042A3B4B7A79FAE5" box="[1062,1142,1370,1394]" pageId="4" pageNumber="350" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
holotype: (8, 9) lateral view; (10, 11) dorsal view.
</paragraph>
</caption>
<paragraph id="853536DB8E339F4200CB3BD57F54F994" blockId="4.[151,1437,1476,1971]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4200CB3BD57F79FA49" bold="true" box="[199,374,1476,1502]" pageId="4" pageNumber="350">Measurements</emphasis>
(in mm). HW 39.0, abdomen including anal appendages 45.0, total length (with head and anal appendages) 63.5
</paragraph>
<paragraph id="853536DB8E339F4200CB381D7B51F904" blockId="4.[151,1437,1476,1971]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4200CB381D7C22F9B1" bold="true" box="[199,557,1548,1574]" pageId="4" pageNumber="350">
Variations in male
<typeStatus id="5A3188798E339F4201A3381D7C28F9B1" box="[431,551,1548,1574]" pageId="4" pageNumber="350" type="paratype">paratypes</typeStatus>
.
</emphasis>
All
<typeStatus id="5A3188798E339F42026D381C7CCEF9B0" box="[609,705,1549,1575]" pageId="4" pageNumber="350" type="paratype">paratype</typeStatus>
males are smaller than the
<typeStatus id="5A3188798E339F42040E381D7A6BF9B1" box="[1026,1124,1548,1574]" pageId="4" pageNumber="350" type="holotype">holotype</typeStatus>
(HW: 36.837.8, abdomen including caudal appendages: 43.545.9, total length: 61.163.0). One specimen has a pale spot in middle of the anteclypeus. One specimen with two cells in triangle on left HW. Ax with
<quantity id="42729B3E8E339F4203DF38457A03F9F9" box="[979,1036,1620,1646]" metricMagnitude="-1" metricUnit="m" metricValue="3.302" pageId="4" pageNumber="350" unit="in" value="13.0">13 in</quantity>
FW and
<quantity id="42729B3E8E339F42047E38457AB6F9F9" box="[1138,1209,1620,1646]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" metricValueMax="2.032" metricValueMin="1.778" pageId="4" pageNumber="350" unit="in" value="7.5" valueMax="8.0" valueMin="7.0">78 in</quantity>
HW, Px with
<quantity id="42729B3E8E339F42055A38457B93F9F9" box="[1366,1436,1620,1647]" metricMagnitude="-1" metricUnit="m" metricValue="1.524" metricValueMax="1.778" metricValueMin="1.27" pageId="4" pageNumber="350" unit="in" value="6.0" valueMax="7.0" valueMin="5.0">57 in</quantity>
FW and
<quantity id="42729B3E8E339F4200FB38697F43F905" box="[247,332,1656,1682]" metricMagnitude="-1" metricUnit="m" metricValue="2.159" metricValueMax="2.54" metricValueMin="1.778" pageId="4" pageNumber="350" unit="in" value="8.5" valueMax="10.0" valueMin="7.0">710 in</quantity>
HW. Cubital crossveins 45 on FW and 3 on HW. Supratriangle with 12 crossveins on FW.
</paragraph>
<paragraph id="853536DB8E339F4200CB388D7F98F921" blockId="4.[151,1437,1476,1971]" box="[199,407,1692,1718]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4200CB388D7F2CF921" box="[199,291,1692,1718]" italics="true" pageId="4" pageNumber="350">Female.</emphasis>
Unknown
</paragraph>
</subSubSection>
<subSubSection id="CD9065508E339F4000CB38D17B70F826" lastPageId="6" lastPageNumber="352" pageId="4" pageNumber="350" type="diagnosis">
<paragraph id="853536DB8E339F4200CB38D17F04F824" blockId="4.[151,1437,1476,1971]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4200CB38D17FC2F94D" bold="true" box="[199,461,1728,1754]" pageId="4" pageNumber="350">Differential diagnosis.</emphasis>
The new species is easily separated from all
<taxonomicName id="428A4D588E339F4203B138D07A87F94D" box="[957,1160,1728,1754]" pageId="4" pageNumber="350">
<emphasis id="B7FEEAC98E339F4203B138D07A3EF94D" box="[957,1073,1729,1754]" italics="true" pageId="4" pageNumber="350">Macromia</emphasis>
species
</taxonomicName>
so far known from
<collectingCountry id="FD9D764B8E339F42056C38D17EC8F969" name="Thailand" pageId="4" pageNumber="350">Thailand</collectingCountry>
by the antehumeral stripe reaching the antealar sinus. So far, this long and broad stripe is unique among the species in the Indomalayan realm reported in
<bibRefCitation id="E11B4B2A8E339F42029339197D5FF8B5" author="Laidlaw, F. F." box="[671,848,1800,1826]" pageId="4" pageNumber="350" pagination="218 - 229" refId="ref4244" refString="Laidlaw, F. F. (1922) Some notes on oriental dragonflies: the genus Macromia. Journal of the Straits Branch of the Royal Asiatic Society, 85, 218 - 229." type="journal article" year="1922">Laidlaw (1922)</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E339F42035139187DFFF8B5" author="Fraser, F. C." box="[861,1008,1800,1827]" pageId="4" pageNumber="350" pagination="423 - 522" refId="ref3711" refString="Fraser, F. C. (1924) A survey of the Odonata fauna of western India with special remarks on the genera Macromia and Idionyx and descriptions of thirty new species. Records Indian Museum, 26 (5), 423 - 522." type="journal article" year="1924">Fraser (1924</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E339F4203F039197A37F8B4" author="Fraser, F. C." box="[1020,1080,1800,1827]" pageId="4" pageNumber="350" refId="ref3758" refString="Fraser, F. C. (1936) The Fauna of British India, including Ceylon and Burma. Odonata. Vol. III. Taylor &amp; Francis, London. xi + 461 pp." type="book" year="1936">1936</bibRefCitation>
),
<bibRefCitation id="E11B4B2A8E339F42044139197B0FF8B5" author="Lieftinck, M. A." box="[1101,1280,1800,1826]" pageId="4" pageNumber="350" pagination="59 - 108" refId="ref4280" refString="Lieftinck, M. A. (1929) A revision of the known Malaysian dragonflies of the genus Macromia Rambur, with comparative notes on species from neighbouring countries and descriptions of new species. Tijdschrift voor Entomologie, 72, 59 - 108." type="journal article" year="1929">Lieftinck (1929</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E339F42050139197B47F8B4" author="Lieftinck, M. A." box="[1293,1352,1800,1827]" pageId="4" pageNumber="350" pagination="251 - 277" refId="ref4326" refString="Lieftinck, M. A. (1955) Further inquiries into the Old World species of Macromia Rambur (Odonata). Zoologische Mededelingen, 33 (25), 251 - 277." type="journal article" year="1955">1955</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E339F42055839197B80F8B5" author="Lieftinck, M. A." box="[1364,1423,1800,1826]" pageId="4" pageNumber="350" pagination="1 - 63" refId="ref4361" refString="Lieftinck, M. A. (1971) Studies in Oriental Corduliidae (Odonata) I. Tijdschrift voor Entomologie, 114, 1 - 63." type="journal article" year="1971">1971</bibRefCitation>
),
<bibRefCitation id="E11B4B2A8E339F42009B393D7F39F8D1" author="Muraki, A." box="[151,310,1836,1862]" pageId="4" pageNumber="350" pagination="39 - 45" refId="ref4463" refString="Muraki, A. (2009) Records of the dragonflies genus Macromia from Taiping of Malay Peninsula. Aeschna, 45, 39 - 45. [in Japanese with English summary]" type="journal article" year="2009">Muraki (2009</bibRefCitation>
, 2010, 2014, 2015, 2017),
<bibRefCitation id="E11B4B2A8E339F420265393D7DCEF8D1" author="Yokoi, N. &amp; Souphanthong, V." box="[617,961,1836,1862]" pageId="4" pageNumber="350" refId="ref4995" refString="Yokoi, N. &amp; Souphanthong, V. (2014) A List of Lao Dragonflies. Kyoei Printing Co Ltd., Koriyama, 110 pp." type="book" year="2014">Yokoi &amp; Souphanthong (2014</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E339F4203C1393D7A06F8D1" author="Yokoi, N. &amp; Souphanthong, V." box="[973,1033,1836,1862]" pageId="4" pageNumber="350" pagination="17 - 24" refId="ref5024" refString="Yokoi, N. &amp; Souphanthong, V. (2019) Description of Macromia murakii sp. nov. (Anisoptera: Macromiidae) from northern Laos, with a consideration on the taxonomic relationship between similar species M. chui and M. daimoji. Tombo, 61, 17 - 24." type="journal article" year="2019">2019</bibRefCitation>
),
<bibRefCitation id="E11B4B2A8E339F420412393D7AF2F8D1" author="Sumanapala, A. P." box="[1054,1277,1836,1862]" pageId="4" pageNumber="350" pagination="169 - 177" refId="ref4804" refString="Sumanapala, A. P. (2021) Macromia weerakooni sp. nov. (Odonata: Anisoptera: Macromiidae), a new dragonfly species from Sri Lanka. International Journal of Odonatology, 24, 169 - 177. https: // doi. org / 10.23797 / 2159 - 6719 _ 24 _ 13" type="journal article" year="2021">Sumanapala (2021)</bibRefCitation>
. Furthermore, there are two
<taxonomicName id="428A4D588E339F42013B39407FDCF8FD" box="[311,467,1872,1898]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="350" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B7FEEAC98E339F42013B39407FA4F8FD" box="[311,427,1873,1898]" italics="true" pageId="4" pageNumber="350">Macromia</emphasis>
sp.
</taxonomicName>
species from
<collectingRegion id="474EF8398E339F42027E39417D50F8FD" box="[626,863,1872,1898]" country="Thailand" name="Chiang Mai" pageId="4" pageNumber="350">Chiang Mai province</collectingRegion>
listed in the book “Atlas of the dragonflies of
<collectingCountry id="FD9D764B8E339F42056C39417EC8F819" name="Thailand" pageId="4" pageNumber="350">Thailand</collectingCountry>
distribution maps by provinces” (
<bibRefCitation id="E11B4B2A8E339F42025439657DBEF819" author="Hamalainen, M. &amp; Pinratana, A." box="[600,945,1908,1934]" pageId="4" pageNumber="350" refId="ref3854" refString="Hamalainen, M., &amp; Pinratana, A. (1999) &quot; Atlas of the dragonflies of Thailand - distribution maps by provinces. Brothers of St. Gabriel in Thailand, Bangkok, vi + 176 pp." type="book" year="1999">Hämäläinen &amp; Pinratana 1999</bibRefCitation>
, p. 95). One of those species is probably
<taxonomicName id="428A4D588E339F42057239647F0BF825" authorityName="Makbun" authorityYear="2022" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="350" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B7FEEAC98E339F42057239647F0BF825" italics="true" pageId="4" pageNumber="350">M. siamensis</emphasis>
</taxonomicName>
.
</paragraph>
<caption id="D1F566538E329F43009B396C7D9CF801" ID-DOI="http://doi.org/10.5281/zenodo.6522534" ID-Zenodo-Dep="6522534" box="[151,915,1917,1942]" httpUri="https://zenodo.org/record/6522534/files/figure.png" pageId="5" pageNumber="351" startId="5.[151,264,1917,1941]" targetBox="[151,1436,181,1894]" targetPageId="5">
<paragraph id="853536DB8E329F43009B396C7D9CF801" blockId="5.[151,915,1917,1942]" box="[151,915,1917,1942]" pageId="5" pageNumber="351">
<emphasis id="B7FEEAC98E329F43009B396C7F5BF801" bold="true" box="[151,340,1917,1942]" pageId="5" pageNumber="351">FIGURES 1213.</emphasis>
Habitat of
<taxonomicName id="428A4D588E329F4301CA396F7C97F802" authority="Makbun, 2022" authorityName="Makbun" authorityYear="2022" box="[454,664,1918,1941]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="5" pageNumber="351" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="B7FEEAC98E329F4301CA396F7C97F802" box="[454,664,1918,1941]" italics="true" pageId="5" pageNumber="351">Macromia siamensis</emphasis>
</taxonomicName>
<emphasis id="B7FEEAC98E329F430292396F7CE0F801" bold="true" box="[670,751,1918,1942]" pageId="5" pageNumber="351">
<taxonomicNameLabel id="ACCD57B28E329F430292396F7CE0F801" box="[670,751,1918,1942]" pageId="5" pageNumber="351" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
in type locality.
</paragraph>
</caption>
<paragraph id="853536DB8E319F4000CB3E867C42FC82" blockId="6.[151,1437,150,789]" pageId="6" pageNumber="352">
Using the diagnostic characters and distribution records of the
<taxonomicName id="428A4D588E319F40039D3E867A52FF26" box="[913,1117,151,177]" pageId="6" pageNumber="352">
<emphasis id="B7FEEAC98E319F40039D3E867A0AFF27" box="[913,1029,151,176]" italics="true" pageId="6" pageNumber="352">Macromia</emphasis>
species
</taxonomicName>
in Asia in
<bibRefCitation id="E11B4B2A8E319F4004D03E867B98FF26" author="Lieftinck, M. A." box="[1244,1431,150,177]" pageId="6" pageNumber="352" pagination="251 - 277" refId="ref4326" refString="Lieftinck, M. A. (1955) Further inquiries into the Old World species of Macromia Rambur (Odonata). Zoologische Mededelingen, 33 (25), 251 - 277." type="journal article" year="1955">Lieftinck (1955)</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E319F40009B3EAA7F49FF41" author="Asahina, S." box="[151,326,187,214]" pageId="6" pageNumber="352" pagination="89 - 98" refId="ref3505" refString="Asahina, S. (1968) Taiwanese Odonata taken by Mr. Y. I. Chui. Kontyu, 36 (1), 89 - 98." type="journal article" year="1968">Asahina (1968)</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E319F40015D3EAA7FD6FF42" author="Vick, G. S." box="[337,473,187,213]" pageId="6" pageNumber="352" pagination="141 - 146" refId="ref4899" refString="Vick, G. S. (1988) A new species of Macromia from Nepal (Anisoptera: Corduliidae). Odonatologica, 17 (2), 141 - 146." type="journal article" year="1988">Vick (1988)</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E319F4001E83EAA7CBFFF42" author="Zhou, W. B. &amp; Wang, Z. D. &amp; Shuai, X. F. &amp; Liu, S. D." box="[484,688,187,213]" pageId="6" pageNumber="352" pagination="149 - 157" refId="ref5079" refString="Zhou, W. B., Wang, Z. D., Shuai, X. F. &amp; Liu, S. D. (1994) Notes on Chinese Macromia and Idionyx, with descriptions of M. kiautai sp. n., M. macula sp. n. and I. yunnanensis sp. n. (Anisoptera: Corduliidae). Odonatologica, 23 (2), 149 - 157." type="journal article" year="1994">
Zhou
<emphasis id="B7FEEAC98E319F40022A3EAA7C52FF42" box="[550,605,187,213]" italics="true" pageId="6" pageNumber="352">et al.</emphasis>
(1994)
</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E319F4002B03EAA7D6DFF42" author="Karube, H." box="[700,866,187,213]" pageId="6" pageNumber="352" pagination="168369" refId="ref3923" refString="Karube, H. (2020) Macromia amphigena. The IUCN Red List of Threatened Species, 2020, e. T 168369 A 140604805. https: // doi. org / 10.2305 / IUCN. UK. 2020 - 3. RLTS. T 168369 A 140604805. en" type="journal article" year="2020">Karube (2020)</bibRefCitation>
,
<bibRefCitation id="E11B4B2A8E319F4003603EAA7A00FF42" author="Wilson, K. D. P." box="[876,1039,187,213]" pageId="6" pageNumber="352" pagination="167109" refId="ref4932" refString="Wilson, K. D. P. (2020) Macromia clio (amended version of 2013 assessment). The IUCN Red List of Threatened Species, 2020, e. T 167109 A 176631533. https: // doi. org / 10.2305 / IUCN. UK. 2020 - 3. RLTS. T 167109 A 176631533. en" type="journal article" year="2020">Wilson (2020)</bibRefCitation>
,
<taxonomicName id="428A4D588E319F4004163EAA7AA3FF43" authorityName="Makbun" authorityYear="2022" box="[1050,1196,187,212]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B7FEEAC98E319F4004163EAA7AA3FF43" box="[1050,1196,187,212]" italics="true" pageId="6" pageNumber="352">M. siamensis</emphasis>
</taxonomicName>
is found to be closest to the following congeners:
<taxonomicName id="428A4D588E319F4001D43ECE7D07FF6E" authority="Selys, 1871" authorityName="Selys" authorityYear="1871" box="[472,776,223,249]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="amphigena">
<emphasis id="B7FEEAC98E319F4001D43ECE7C74FF6E" box="[472,635,223,249]" italics="true" pageId="6" pageNumber="352">M. amphigena</emphasis>
Selys, 1871
</taxonomicName>
from
<collectingCountry id="FD9D764B8E319F40035C3ECE7DD9FF6E" box="[848,982,223,249]" name="Kazakhstan" pageId="6" pageNumber="352">Kazakhstan</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F4003EE3ECE7A20FF6E" box="[994,1071,223,249]" name="Russia" pageId="6" pageNumber="352">Russia</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F4004303ECE7A8DFF6E" box="[1084,1154,223,249]" name="China" pageId="6" pageNumber="352">China</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F4004833ECE7AF0FF6E" box="[1167,1279,223,249]" name="Mongolia" pageId="6" pageNumber="352">Mongolia</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F4005073ECE7B41FF6E" box="[1291,1358,223,249]" name="Japan" pageId="6" pageNumber="352">Japan</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F4005573ECE7ED4FE8A" name="North Korea" pageId="6" pageNumber="352">North Korea</collectingCountry>
and
<collectingCountry id="FD9D764B8E319F4001193F127FA9FE8A" box="[277,422,259,285]" name="South Korea" pageId="6" pageNumber="352">South Korea</collectingCountry>
,
<taxonomicName id="428A4D588E319F4001BF3F127CA0FE8A" authority="Vick, 1988" authorityName="Vick" authorityYear="1988" box="[435,687,259,285]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="sombui">
<emphasis id="B7FEEAC98E319F4001BF3F127C24FE8A" box="[435,555,259,285]" italics="true" pageId="6" pageNumber="352">M. sombui</emphasis>
<bibRefCitation id="E11B4B2A8E319F40023F3F127CA0FE8A" author="Vick, G. S." box="[563,687,259,285]" pageId="6" pageNumber="352" pagination="141 - 146" refId="ref4899" refString="Vick, G. S. (1988) A new species of Macromia from Nepal (Anisoptera: Corduliidae). Odonatologica, 17 (2), 141 - 146." type="journal article" year="1988">Vick, 1988</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="FD9D764B8E319F4002FB3F127D32FE8A" box="[759,829,259,285]" name="Nepal" pageId="6" pageNumber="352">Nepal</collectingCountry>
,
<taxonomicName id="428A4D588E319F4003453F127A7DFE8A" authority="Laidlaw, 1922" authorityName="Laidlaw" authorityYear="1922" box="[841,1138,259,285]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="cydippe">
<emphasis id="B7FEEAC98E319F4003453F127DC8FE8A" box="[841,967,259,285]" italics="true" pageId="6" pageNumber="352">M. cydippe</emphasis>
<bibRefCitation id="E11B4B2A8E319F4003DC3F127A7DFE8A" author="Laidlaw, F. F." box="[976,1138,259,285]" pageId="6" pageNumber="352" pagination="218 - 229" refId="ref4244" refString="Laidlaw, F. F. (1922) Some notes on oriental dragonflies: the genus Macromia. Journal of the Straits Branch of the Royal Asiatic Society, 85, 218 - 229." type="journal article" year="1922">Laidlaw, 1922</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="FD9D764B8E319F4004B53F127B26FE8A" box="[1209,1321,259,285]" name="Indonesia" pageId="6" pageNumber="352">Indonesia</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F40053A3F127B92FE8A" box="[1334,1437,259,285]" name="Malaysia" pageId="6" pageNumber="352">Malaysia</collectingCountry>
and
<collectingCountry id="FD9D764B8E319F4000CA3F367F23FED6" box="[198,300,295,321]" name="Thailand" pageId="6" pageNumber="352">Thailand</collectingCountry>
,
<taxonomicName id="428A4D588E319F4001343F367D26FED6" authority="Yokoi et Mitamura, 2002" authorityName="Yokoi et Mitamura" authorityYear="2002" box="[312,809,295,321]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="vangviengensis">
<emphasis id="B7FEEAC98E319F4001343F367C07FED7" box="[312,520,295,320]" italics="true" pageId="6" pageNumber="352">M. vangviengensis</emphasis>
Yokoi et Mitamura, 2002
</taxonomicName>
from
<collectingCountry id="FD9D764B8E319F4003613F367DDEFED6" box="[877,977,295,321]" name="Thailand" pageId="6" pageNumber="352">Thailand</collectingCountry>
and
<collectingCountry id="FD9D764B8E319F40040B3F367A4FFED6" box="[1031,1088,295,321]" name="Laos" pageId="6" pageNumber="352">Laos</collectingCountry>
,
<taxonomicName id="428A4D588E319F4004403F367B01FED6" authority="Ris, 1916" authorityName="Ris" authorityYear="1916" box="[1100,1294,295,321]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="B7FEEAC98E319F4004403F367A94FED6" box="[1100,1179,295,321]" italics="true" pageId="6" pageNumber="352">M. clio</emphasis>
Ris, 1916
</taxonomicName>
from
<collectingCountry id="FD9D764B8E319F40055E3F367B97FED6" box="[1362,1432,295,321]" name="China" pageId="6" pageNumber="352">China</collectingCountry>
,
<collectingCountry id="FD9D764B8E319F40009B3F5A7EE3FEF2" box="[151,236,331,357]" name="Taiwan" pageId="6" pageNumber="352">Taiwan</collectingCountry>
, and
<collectingCountry id="FD9D764B8E319F4001253F5A7F63FEF2" box="[297,364,331,357]" name="Japan" pageId="6" pageNumber="352">Japan</collectingCountry>
, and also
<taxonomicName id="428A4D588E319F4001D23F5A7D3EFEF2" authority="Lieftinck, 1955" authorityName="Lieftinck" authorityYear="1955" box="[478,817,330,357]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="malleifera">
<emphasis id="B7FEEAC98E319F4001D23F5A7C77FEF2" box="[478,632,331,357]" italics="true" pageId="6" pageNumber="352">M. malleifera</emphasis>
<bibRefCitation id="E11B4B2A8E319F40028C3F5A7D3EFEF2" author="Lieftinck, M. A." box="[640,817,330,357]" pageId="6" pageNumber="352" pagination="251 - 277" refId="ref4326" refString="Lieftinck, M. A. (1955) Further inquiries into the Old World species of Macromia Rambur (Odonata). Zoologische Mededelingen, 33 (25), 251 - 277." type="journal article" year="1955">Lieftinck, 1955</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="428A4D588E319F4003623F5A7B52FEF2" authority="Zhou, Wang, Shuai &amp; Liu, 1994" authorityName="Zhou, Wang, Shuai &amp; Liu" authorityYear="1994" box="[878,1373,331,357]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="macula">
<emphasis id="B7FEEAC98E319F4003623F5A7DE8FEF2" box="[878,999,331,357]" italics="true" pageId="6" pageNumber="352">M. macula</emphasis>
<bibRefCitation id="E11B4B2A8E319F4003E33F5A7B52FEF2" author="Zhou, W. B. &amp; Wang, Z. D. &amp; Shuai, X. F. &amp; Liu, S. D." box="[1007,1373,331,357]" pageId="6" pageNumber="352" pagination="149 - 157" refId="ref5079" refString="Zhou, W. B., Wang, Z. D., Shuai, X. F. &amp; Liu, S. D. (1994) Notes on Chinese Macromia and Idionyx, with descriptions of M. kiautai sp. n., M. macula sp. n. and I. yunnanensis sp. n. (Anisoptera: Corduliidae). Odonatologica, 23 (2), 149 - 157." type="journal article" year="1994">Zhou, Wang, Shuai &amp; Liu, 1994</bibRefCitation>
</taxonomicName>
from
<collectingCountry id="FD9D764B8E319F40009B3F7E7ED1FE1E" box="[151,222,367,393]" name="China" pageId="6" pageNumber="352">China</collectingCountry>
.All these species share the hammer-shaped tip of posterior hamulus. However,
<taxonomicName id="428A4D588E319F4004433F7E7AEFFE1F" authorityName="Makbun" authorityYear="2022" box="[1103,1248,367,392]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B7FEEAC98E319F4004433F7E7AEFFE1F" box="[1103,1248,367,392]" italics="true" pageId="6" pageNumber="352">M. siamensis</emphasis>
</taxonomicName>
can be separated from all of these by a combination of characters: a broad antehumeral stripe reaching antealar sinus (a short and narrower stripe in
<taxonomicName id="428A4D588E319F40016E3FA67C0DFE46" authorityName="Selys" authorityYear="1871" box="[354,514,439,465]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="amphigena">
<emphasis id="B7FEEAC98E319F40016E3FA67C0DFE46" box="[354,514,439,465]" italics="true" pageId="6" pageNumber="352">M. amphigena</emphasis>
</taxonomicName>
,
<emphasis id="B7FEEAC98E319F40021C3FA67D6BFE46" box="[528,868,439,465]" italics="true" pageId="6" pageNumber="352">
<taxonomicName id="428A4D588E319F40021C3FA67C6FFE46" authorityName="Ris" authorityYear="1916" box="[528,608,439,465]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="clio">M. clio</taxonomicName>
,
<taxonomicName id="428A4D588E319F4002673FA67CEBFE46" authorityName="Zhou, Wang, Shuai &amp; Liu" authorityYear="1994" box="[619,740,439,465]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="macula">M. macula</taxonomicName>
,
<taxonomicName id="428A4D588E319F4002E23FA67D6BFE46" authorityName="Vick" authorityYear="1988" box="[750,868,439,465]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="sombui">M. sombui</taxonomicName>
</emphasis>
; absent in
<emphasis id="B7FEEAC98E319F4003D63FA67B41FE47" box="[986,1358,439,465]" italics="true" pageId="6" pageNumber="352">
<taxonomicName id="428A4D588E319F4003D63FA67A7AFE46" authorityName="Lieftinck" authorityYear="1955" box="[986,1141,439,465]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="malleifera">M. malleifera</taxonomicName>
,
<taxonomicName id="428A4D588E319F4004733FA67B41FE47" authorityName="Yokoi et Mitamura" authorityYear="2002" box="[1151,1358,439,464]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="vangviengensis">M. vangviengensis</taxonomicName>
</emphasis>
), postfrons yellow with black upper margin (dark brown to black with green/blue iridescent reflection in
<taxonomicName id="428A4D588E319F4004FF3FCA7B9AFE62" authorityName="Selys" authorityYear="1871" box="[1267,1429,475,501]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="amphigena">
<emphasis id="B7FEEAC98E319F4004FF3FCA7B9AFE62" box="[1267,1429,475,501]" italics="true" pageId="6" pageNumber="352">M. amphigena</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40009B3FEE7EE8FD8E" authorityName="Ris" authorityYear="1916" box="[151,231,511,537]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="B7FEEAC98E319F40009B3FEE7EE8FD8E" box="[151,231,511,537]" italics="true" pageId="6" pageNumber="352">M. clio</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F4000F93FEE7F7EFD8E" authorityName="Laidlaw" authorityYear="1922" box="[245,369,511,537]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="cydippe">
<emphasis id="B7FEEAC98E319F4000F93FEE7F7EFD8E" box="[245,369,511,537]" italics="true" pageId="6" pageNumber="352">M. cydippe</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F4001733FEE7C17FD8E" authorityName="Lieftinck" authorityYear="1955" box="[383,536,511,537]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="malleifera">
<emphasis id="B7FEEAC98E319F4001733FEE7C17FD8E" box="[383,536,511,537]" italics="true" pageId="6" pageNumber="352">M. malleifera</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40022A3FEE7C93FD8E" authorityName="Vick" authorityYear="1988" box="[550,668,511,537]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="sombui">
<emphasis id="B7FEEAC98E319F40022A3FEE7C93FD8E" box="[550,668,511,537]" italics="true" pageId="6" pageNumber="352">M. sombui</emphasis>
</taxonomicName>
; black with blue iridescent reflection with two small yellow spots in
<taxonomicName id="428A4D588E319F40009B3C327F02FDAA" authorityName="Zhou, Wang, Shuai &amp; Liu" authorityYear="1994" box="[151,269,547,573]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="macula">
<emphasis id="B7FEEAC98E319F40009B3C327F02FDAA" box="[151,269,547,573]" italics="true" pageId="6" pageNumber="352">M. macula</emphasis>
</taxonomicName>
; dark brown in
<taxonomicName id="428A4D588E319F4001B03C327C84FDAB" authorityName="Yokoi et Mitamura" authorityYear="2002" box="[444,651,547,572]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="vangviengensis">
<emphasis id="B7FEEAC98E319F4001B03C327C84FDAB" box="[444,651,547,572]" italics="true" pageId="6" pageNumber="352">M. vangviengensis</emphasis>
</taxonomicName>
), S10 without distinct protuberance (the same in
<taxonomicName id="428A4D588E319F4004A63C327B2FFDAA" authorityName="Zhou, Wang, Shuai &amp; Liu" authorityYear="1994" box="[1194,1312,547,573]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="macula">
<emphasis id="B7FEEAC98E319F4004A63C327B2FFDAA" box="[1194,1312,547,573]" italics="true" pageId="6" pageNumber="352">M. macula</emphasis>
</taxonomicName>
; a strongly raised protuberance with pointed apex in
<taxonomicName id="428A4D588E319F4002643C567CD0FDF6" authorityName="Vick" authorityYear="1988" box="[616,735,583,609]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="sombui">
<emphasis id="B7FEEAC98E319F4002643C567CD0FDF6" box="[616,735,583,609]" italics="true" pageId="6" pageNumber="352">M. sombui</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F4002E13C567D89FDF6" authorityName="Lieftinck" authorityYear="1955" box="[749,902,583,609]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="malleifera">
<emphasis id="B7FEEAC98E319F4002E13C567D89FDF6" box="[749,902,583,609]" italics="true" pageId="6" pageNumber="352">M. malleifera</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F4003983C567A6AFDF7" authorityName="Yokoi et Mitamura" authorityYear="2002" box="[916,1125,583,608]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="vangviengensis">
<emphasis id="B7FEEAC98E319F4003983C567A6AFDF7" box="[916,1125,583,608]" italics="true" pageId="6" pageNumber="352">M. vangviengensis</emphasis>
</taxonomicName>
; a strongly raised protuberance with blunt apex in
<taxonomicName id="428A4D588E319F4001913C7A7FE4FD12" authorityName="Ris" authorityYear="1916" box="[413,491,619,645]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="B7FEEAC98E319F4001913C7A7FE4FD12" box="[413,491,619,645]" italics="true" pageId="6" pageNumber="352">M. clio</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F4001F43C7A7C7CFD12" authorityName="Laidlaw" authorityYear="1922" box="[504,627,619,645]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="cydippe">
<emphasis id="B7FEEAC98E319F4001F43C7A7C7CFD12" box="[504,627,619,645]" italics="true" pageId="6" pageNumber="352">M. cydippe</emphasis>
</taxonomicName>
; absent in
<taxonomicName id="428A4D588E319F4002E53C7A7D86FD12" authorityName="Selys" authorityYear="1871" box="[745,905,619,645]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="amphigena">
<emphasis id="B7FEEAC98E319F4002E53C7A7D86FD12" box="[745,905,619,645]" italics="true" pageId="6" pageNumber="352">M. amphigena</emphasis>
</taxonomicName>
), S46 with a large basal ring (S36 with dorsal spots in
<taxonomicName id="428A4D588E319F4000FF3C9E7F9CFD3E" authorityName="Selys" authorityYear="1871" box="[243,403,655,681]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="amphigena">
<emphasis id="B7FEEAC98E319F4000FF3C9E7F9CFD3E" box="[243,403,655,681]" italics="true" pageId="6" pageNumber="352">M. amphigena</emphasis>
</taxonomicName>
,
<taxonomicName id="428A4D588E319F4001AD3C9E7FE0FD3E" authorityName="Ris" authorityYear="1916" box="[417,495,655,681]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="clio">
<emphasis id="B7FEEAC98E319F4001AD3C9E7FE0FD3E" box="[417,495,655,681]" italics="true" pageId="6" pageNumber="352">M. clio</emphasis>
</taxonomicName>
,
<emphasis id="B7FEEAC98E319F4001F03C9E7D95FD3E" box="[508,922,655,681]" italics="true" pageId="6" pageNumber="352">
<taxonomicName id="428A4D588E319F4001F03C9E7C7AFD3E" authorityName="Zhou, Wang, Shuai &amp; Liu" authorityYear="1994" box="[508,629,655,681]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="macula">M. macula</taxonomicName>
,
<taxonomicName id="428A4D588E319F40028C3C9E7D15FD3E" authorityName="Lieftinck" authorityYear="1955" box="[640,794,655,681]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="malleifera">M. malleifera</taxonomicName>
,
<taxonomicName id="428A4D588E319F4003283C9E7D95FD3E" authorityName="Vick" authorityYear="1988" box="[804,922,655,681]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="sombui">M. sombui</taxonomicName>
</emphasis>
; S35 with dorsal spots in
<taxonomicName id="428A4D588E319F4004C93C9E7B9BFD3F" authorityName="Yokoi et Mitamura" authorityYear="2002" box="[1221,1428,655,680]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="vangviengensis">
<emphasis id="B7FEEAC98E319F4004C93C9E7B9BFD3F" box="[1221,1428,655,680]" italics="true" pageId="6" pageNumber="352">M. vangviengensis</emphasis>
</taxonomicName>
; S36 unmarked in
<taxonomicName id="428A4D588E319F4001653CA27FEAFD5A" authorityName="Laidlaw" authorityYear="1922" box="[361,485,691,717]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="cydippe">
<emphasis id="B7FEEAC98E319F4001653CA27FEAFD5A" box="[361,485,691,717]" italics="true" pageId="6" pageNumber="352">M. cydippe</emphasis>
</taxonomicName>
), postclypeus entirely yellow (the same in
<taxonomicName id="428A4D588E319F4003CD3CA27A6DFD5A" authorityName="Selys" authorityYear="1871" box="[961,1122,691,717]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="amphigena">
<emphasis id="B7FEEAC98E319F4003CD3CA27A6DFD5A" box="[961,1122,691,717]" italics="true" pageId="6" pageNumber="352">M. amphigena</emphasis>
</taxonomicName>
,
<emphasis id="B7FEEAC98E319F4004633CA27B4DFD5A" box="[1135,1346,691,717]" italics="true" pageId="6" pageNumber="352">
<taxonomicName id="428A4D588E319F4004633CA27ACFFD5A" authorityName="Ris" authorityYear="1916" box="[1135,1216,691,717]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="clio">M. clio</taxonomicName>
,
<taxonomicName id="428A4D588E319F4004C73CA27B4DFD5A" authorityName="Zhou, Wang, Shuai &amp; Liu" authorityYear="1994" box="[1227,1346,691,717]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="macula">M. macula</taxonomicName>
</emphasis>
; yellow with dark brown lower margin in
<emphasis id="B7FEEAC98E319F4002023CC67D8BFD67" box="[526,900,727,753]" italics="true" pageId="6" pageNumber="352">
<taxonomicName id="428A4D588E319F4002023CC67CA6FD66" authorityName="Lieftinck" authorityYear="1955" box="[526,681,727,753]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="malleifera">M. malleifera</taxonomicName>
,
<taxonomicName id="428A4D588E319F4002B83CC67D8BFD67" authorityName="Yokoi et Mitamura" authorityYear="2002" box="[692,900,727,752]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="vangviengensis">M. vangviengensis</taxonomicName>
</emphasis>
; entirely dark brown in
<taxonomicName id="428A4D588E319F40049E3CC67B01FD66" authorityName="Laidlaw" authorityYear="1922" box="[1170,1294,727,753]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="cydippe">
<emphasis id="B7FEEAC98E319F40049E3CC67B01FD66" box="[1170,1294,727,753]" italics="true" pageId="6" pageNumber="352">M. cydippe</emphasis>
</taxonomicName>
; dark brown with wedge-shape spots in
<taxonomicName id="428A4D588E319F4001CA3CEA7C32FC82" authorityName="Vick" authorityYear="1988" box="[454,573,763,789]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="sombui">
<emphasis id="B7FEEAC98E319F4001CA3CEA7C32FC82" box="[454,573,763,789]" italics="true" pageId="6" pageNumber="352">M. sombui</emphasis>
</taxonomicName>
).
</paragraph>
<caption id="D1F566538E319F40009B388C7D12F922" ID-DOI="http://doi.org/10.5281/zenodo.6522536" ID-Zenodo-Dep="6522536" box="[151,797,1693,1717]" httpUri="https://zenodo.org/record/6522536/files/figure.png" pageId="6" pageNumber="352" startId="6.[151,250,1693,1717]" targetBox="[151,1436,813,1670]" targetPageId="6">
<paragraph id="853536DB8E319F40009B388C7D12F922" blockId="6.[151,797,1693,1717]" box="[151,797,1693,1717]" pageId="6" pageNumber="352">
<emphasis id="B7FEEAC98E319F40009B388C7F2FF922" bold="true" box="[151,288,1693,1717]" pageId="6" pageNumber="352">FIGURE 14.</emphasis>
Distribution map of
<taxonomicName id="428A4D588E319F4001F8388F7CC9F922" authority="Makbun, 2022" authorityName="Makbun" authorityYear="2022" box="[500,710,1694,1717]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="siamensis" status="sp. nov.">
<emphasis id="B7FEEAC98E319F4001F8388F7CC9F922" box="[500,710,1694,1717]" italics="true" pageId="6" pageNumber="352">Macromia siamensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACCD57B28E319F4002C0388C7D12F922" box="[716,797,1693,1717]" pageId="6" pageNumber="352" rank="species">sp. nov.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="853536DB8E319F4000CB38F27B70F826" blockId="6.[151,1436,1763,2006]" pageId="6" pageNumber="352">
<emphasis id="B7FEEAC98E319F4000CB38F27FCCF96A" bold="true" box="[199,451,1763,1789]" pageId="6" pageNumber="352">Habitat and Ecology.</emphasis>
<taxonomicName id="428A4D588E319F4001C638F57CBEF96A" authorityName="Makbun" authorityYear="2022" box="[458,689,1764,1789]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="siamensis">
<emphasis id="B7FEEAC98E319F4001C638F57CBEF96A" box="[458,689,1764,1789]" italics="true" pageId="6" pageNumber="352">Macromia siamensis</emphasis>
</taxonomicName>
was found patrolling above a shallow and small stream in an open area at 580 meters above sea level (
<figureCitation id="1DB12A5E8E319F40022539167CBCF8B6" box="[553,691,1799,1825]" captionStart="FIGURES 1213" captionStartId="5.[151,264,1917,1941]" captionTargetBox="[151,1436,181,1894]" captionTargetId="figure-19@5.[151,1436,181,1894]" captionTargetPageId="5" captionText="FIGURES 1213. Habitat of Macromia siamensis sp. nov. in type locality." figureDoi="http://doi.org/10.5281/zenodo.6522534" httpUri="https://zenodo.org/record/6522534/files/figure.png" pageId="6" pageNumber="352">Figs. 1213</figureCitation>
). I found this species a few times and only one or
<specimenCount id="938CFD528E319F4004E739197B51F8B6" box="[1259,1374,1799,1826]" pageId="6" pageNumber="352" type="male">two males</specimenCount>
were encountered each time.
<taxonomicName id="428A4D588E319F4001A4393D7C69F8D2" authorityName="Fraser" authorityYear="1922" box="[424,614,1835,1861]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="flavocolorata">
<emphasis id="B7FEEAC98E319F4001A4393D7C69F8D2" box="[424,614,1835,1861]" italics="true" pageId="6" pageNumber="352">M. flavocolorata</emphasis>
</taxonomicName>
and
<taxonomicName id="428A4D588E319F4002AD393D7D86F8D2" authorityName="Hamalainen" authorityYear="1986" box="[673,905,1835,1861]" class="Insecta" family="Corduliidae" genus="Macromia" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="chaiyaphumensis">
<emphasis id="B7FEEAC98E319F4002AD393D7D86F8D2" box="[673,905,1835,1861]" italics="true" pageId="6" pageNumber="352">M. chaiyaphumensis</emphasis>
</taxonomicName>
are also present in this stream but at different periods in a year. Other interesting species found at this stream were
<taxonomicName id="428A4D588E319F40039C395E7B97F8FE" authority="Makbun &amp; Fleck, 2018" authorityName="Makbun &amp; Fleck" authorityYear="2018" box="[912,1432,1871,1897]" class="Insecta" family="Gomphidae" genus="Microgomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="farrelli">
<emphasis id="B7FEEAC98E319F40039C395E7A83F8FE" box="[912,1164,1871,1897]" italics="true" pageId="6" pageNumber="352">Microgomphus farrelli</emphasis>
Makbun &amp; Fleck, 2018
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40009B39627F75F819" box="[151,378,1907,1934]" class="Insecta" family="Gomphidae" genus="Lamelligomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B7FEEAC98E319F40009B39627F5DF81A" box="[151,338,1907,1933]" italics="true" pageId="6" pageNumber="352">Lamelligomphus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40018A39627D8EF81A" authority="(Fraser, 1927)" baseAuthorityName="Fraser" baseAuthorityYear="1927" box="[390,897,1907,1933]" class="Insecta" family="Gomphidae" genus="Nihonogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="pulcherrimus">
<emphasis id="B7FEEAC98E319F40018A39627CD5F81A" box="[390,730,1907,1933]" italics="true" pageId="6" pageNumber="352">Nihonogomphus pulcherrimus</emphasis>
(Fraser, 1927)
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40038039627A5DF819" box="[908,1106,1907,1934]" class="Insecta" family="Gomphidae" genus="Stylogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B7FEEAC98E319F40038039627A26F81A" box="[908,1065,1907,1933]" italics="true" pageId="6" pageNumber="352">Stylogomphus</emphasis>
sp.
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40045239627F6CF825" authority="(Fraser, 1926)" baseAuthorityName="Fraser" baseAuthorityYear="1926" class="Insecta" family="Gomphidae" genus="Phaenandrogomphus" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="tokinicus">
<emphasis id="B7FEEAC98E319F40045239627EB3F826" italics="true" pageId="6" pageNumber="352">Phaenandrogomphus tokinicus</emphasis>
(Fraser, 1926)
</taxonomicName>
,
<taxonomicName id="428A4D588E319F40016339867C54F826" authorityName="Hagen in Selys" authorityYear="1859" box="[367,603,1943,1969]" class="Insecta" family="Lestoideidae" genus="Philoganga" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="B7FEEAC98E319F40016339867C54F826" box="[367,603,1943,1969]" italics="true" pageId="6" pageNumber="352">Philoganga montana</emphasis>
</taxonomicName>
(Hagen in Selys, 1859), and
<taxonomicName id="428A4D588E319F4003AD39867B73F826" authority="Hoess, 2007" authorityName="Hoess" authorityYear="2007" box="[929,1404,1943,1970]" class="Insecta" family="Protoneuridae" genus="Prodasineura" kingdom="Animalia" order="Odonata" pageId="6" pageNumber="352" phylum="Arthropoda" rank="species" species="doisuthepensis">
<emphasis id="B7FEEAC98E319F4003AD39867AE9F826" box="[929,1254,1943,1969]" italics="true" pageId="6" pageNumber="352">Prodasineura doisuthepensis</emphasis>
Hoess, 2007
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="CD9065508E319F4000CB39AA7B96F842" box="[199,1433,1979,2006]" pageId="6" pageNumber="352" type="distribution">
<paragraph id="853536DB8E319F4000CB39AA7B96F842" blockId="6.[151,1436,1763,2006]" box="[199,1433,1979,2006]" pageId="6" pageNumber="352">
<emphasis id="B7FEEAC98E319F4000CB39AA7F6EF842" bold="true" box="[199,353,1979,2005]" pageId="6" pageNumber="352">Distribution.</emphasis>
To date, it is only known from the
<typeStatus id="5A3188798E319F4002E739AD7D14F841" box="[747,795,1980,2006]" pageId="6" pageNumber="352">type</typeStatus>
locality,
<collectingRegion id="474EF8398E319F40038E39AA7A7CF842" box="[898,1139,1979,2005]" country="Thailand" name="Chiang Mai" pageId="6" pageNumber="352">Chiang Mai province</collectingRegion>
, North
<collectingCountry id="FD9D764B8E319F4004CA39AA7B25F842" box="[1222,1322,1979,2005]" name="Thailand" pageId="6" pageNumber="352">Thailand</collectingCountry>
(
<figureCitation id="1DB12A5E8E319F40053539AA7B83F842" box="[1337,1420,1979,2005]" captionStart="FIGURE 14" captionStartId="6.[151,250,1693,1717]" captionTargetBox="[151,1436,813,1670]" captionTargetId="figure-355@6.[151,1436,813,1670]" captionTargetPageId="6" captionText="FIGURE 14. Distribution map of Macromia siamensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.6522536" httpUri="https://zenodo.org/record/6522536/files/figure.png" pageId="6" pageNumber="352">Fig. 14</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>