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<mods:title id="71D0368BADB0DE99903FB50B5FF139B0">Review of the genus Cruziohyla (Anura: Phyllomedusidae), with description of a new species</mods:title>
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<treatment id="03EC87FDFFFFFF9BC2B2F8B043EAFB9D" ID-DOI="http://doi.org/10.5281/zenodo.5953564" ID-GBIF-Taxon="148462438" ID-Zenodo-Dep="5953564" LSID="urn:lsid:plazi:treatment:03EC87FDFFFFFF9BC2B2F8B043EAFB9D" httpUri="http://treatment.plazi.org/id/03EC87FDFFFFFF9BC2B2F8B043EAFB9D" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407">
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<paragraph id="8BFA36EBFFFFFF9AC2B2F8B04356F8CB" blockId="6.[151,677,1858,1953]" box="[151,635,1858,1884]" pageId="6" pageNumber="407">
<heading id="D0B28187FFFFFF9AC2B2F8B04356F8CB" bold="true" box="[151,635,1858,1884]" fontSize="11" level="1" pageId="6" pageNumber="407" reason="1">
<taxonomicName id="4C454D68FFFFFF9AC2B2F8B04356F8CB" authority="(Boulenger, 1902)" baseAuthorityName="Boulenger" baseAuthorityYear="1902" box="[151,635,1858,1884]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="407" phylum="Chordata" rank="species" species="calcarifer">
<emphasis id="B931EAF9FFFFFF9AC2B2F8B04356F8CB" bold="true" box="[151,635,1858,1884]" pageId="6" pageNumber="407">
<emphasis id="B931EAF9FFFFFF9AC2B2F8B040B4F8CB" bold="true" box="[151,409,1858,1884]" italics="true" pageId="6" pageNumber="407">Cruziohyla calcarifer</emphasis>
(
<bibRefCitation id="EFD44B1AFFFFFF9AC38DF8B0435FF8CB" box="[424,626,1858,1884]" pageId="6" pageNumber="407" refId="ref11310" refString="Boulenger, G. A. (1902) Descriptions of new batrachians and reptiles from north-western Ecuador. Annals and Magazine of Natural History, Series 7, 9, 51 - 57. https: // doi. org / 10.1080 / 00222930208678538" type="journal article">Boulenger, 1902</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BFA36EBFFFFFF9AC2B2F89640F1F8EA" blockId="6.[151,677,1858,1953]" box="[151,476,1892,1917]" pageId="6" pageNumber="407">
(
<figureCitation id="137E2A6EFFFFFF9AC2BAF89641D1F8EA" box="[159,252,1892,1917]" captionStart="FIGURE 1" captionStartId="5.[151,250,1375,1397]" captionTargetBox="[168,1419,278,1340]" captionTargetId="figure@5.[151,1436,265,1354]" captionTargetPageId="5" captionText="FIGURE 1. Members of the genus Cruziohyla: (a) C. sylviae sp. n.; (b) C. calcarifer (Boulenger, 1902); (c) C. craspedopus (Funkhouser, 1957)." httpUri="https://zenodo.org/record/1444876/files/figure.png" pageId="6" pageNumber="407">Figs. 1b</figureCitation>
,
<figureCitation id="137E2A6EFFFFFF9AC32DF8964035F8EA" box="[264,280,1892,1917]" captionStart="FIGURE 5" captionStartId="10.[151,250,1973,1995]" captionTargetBox="[215,1375,378,1953]" captionTargetId="figure@10.[211,1379,373,1961]" captionTargetPageId="10" captionText="FIGURE 5. Dark pigmented ventral markings of C. calcarifer (a) Juvenile originating Esmeraldas, Ecuador (b) Ventral marking detail (c) Ventral marking x 40 magnification." httpUri="https://zenodo.org/record/1444884/files/figure.png" pageId="6" pageNumber="407">5</figureCitation>
,
<figureCitation id="137E2A6EFFFFFF9AC300F896406EF8EA" box="[293,323,1892,1917]" captionStart="FIGURE 6" captionStartId="11.[151,250,1500,1522]" captionTargetBox="[203,1348,225,1429]" captionTargetId="figure@11.[151,1436,193,1478]" captionTargetPageId="11" captionText="FIGURE 6. Hands and feet of Cruziohyla: (a) C. sylviae sp. n.; (b) C. calcarifer; (c) C. craspedopus." httpUri="https://zenodo.org/record/1444886/files/figure.png" pageId="6" pageNumber="407">6b</figureCitation>
,
<figureCitation id="137E2A6EFFFFFF9AC36BF8964040F8EA" box="[334,365,1892,1917]" captionStart="FIGURE 7" captionStartId="12.[151,250,1644,1666]" captionTargetBox="[182,1401,366,1534]" captionTargetId="figure@12.[151,1436,337,1622]" captionTargetPageId="12" captionText="FIGURE 7. Selected morphological characters of Cruziohyla: (a) C. sylviae sp. n. (MM1029); (b) C. calcarifer (MM1020); (c) C. craspedopus (MM1016)." httpUri="https://zenodo.org/record/1444888/files/figure.png" pageId="6" pageNumber="407">7b</figureCitation>
,
<figureCitation id="137E2A6EFFFFFF9AC35DF89640BAF8EA" box="[376,407,1892,1917]" captionStart="FIGURE 9" captionStartId="13.[151,250,1531,1553]" captionTargetBox="[164,1422,1152,1499]" captionTargetId="figure@13.[151,1436,1144,1509]" captionTargetPageId="13" captionText="FIGURE 9. Audiospectragrams of the advertisement call of members of the genus Cruziohyla: (a) C. sylviae sp. n.; (b) C. calcarifer (courtesy AMNH); (c) C. craspedopus (courtesy Morley Read)." httpUri="https://zenodo.org/record/1444892/files/figure.png" pageId="6" pageNumber="407">9b</figureCitation>
,
<figureCitation id="137E2A6EFFFFFF9AC387F89640E2F8EA" box="[418,463,1892,1917]" captionStart="FIGURE 11" captionStartId="16.[151,250,1793,1815]" captionTargetBox="[191,1366,506,1733]" captionTargetId="figure@16.[151,1436,481,1771]" captionTargetPageId="16" captionText="FIGURE 11. Size comparison of two adult female Cruziohyla specimens from Panama: (a). C. sylviae sp. n. [KU 77415] from Laguna, Darien (Courtesy William E. Duellman) (b). C. calcarifer [AMNH A107238] from Chanquinola, Bocas del Toro (Courtesy Charles W. Myers)." httpUri="https://zenodo.org/record/1444896/files/figure.png" pageId="6" pageNumber="407">11b</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6560FFFFFF9BC2B2F87A4213FE34" lastPageId="7" lastPageNumber="408" pageId="6" pageNumber="407" type="description">
<paragraph id="8BFA36EBFFFFFF9AC2B2F87A4388F836" blockId="6.[151,677,1858,1953]" box="[151,677,1928,1953]" pageId="6" pageNumber="407">Splendid Tree Frog / Rana de arból Esplendida</paragraph>
<paragraph id="8BFA36EBFFFFFF9AC2B2F822436FF871" blockId="6.[151,578,2000,2022]" box="[151,578,2000,2022]" pageId="6" pageNumber="407">
<emphasis id="B931EAF9FFFFFF9AC2B2F82240AAF871" box="[151,391,2000,2022]" italics="true" pageId="6" pageNumber="407">
<taxonomicName id="4C454D68FFFFFF9AC2B2F8224025F871" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[151,264,2000,2022]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="407" phylum="Chordata" rank="genus">Agalychnis</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFFFF9AC3ADF8224312F871" box="[392,575,2000,2022]" pageId="6" pageNumber="407" refId="ref11310" refString="Boulenger, G. A. (1902) Descriptions of new batrachians and reptiles from north-western Ecuador. Annals and Magazine of Natural History, Series 7, 9, 51 - 57. https: // doi. org / 10.1080 / 00222930208678538" type="journal article">Boulenger (1902)</bibRefCitation>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FF6A427BFF39" blockId="7.[151,964,152,419]" box="[151,854,152,174]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FF6A408BFF39" box="[151,422,152,174]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FF6A400BFF39" authorityName=", Wagler" authorityYear="1830" box="[151,294,152,174]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Phyllomedusa</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC382FF6B430EFF39" box="[423,547,152,174]" pageId="7" pageNumber="408" refId="ref13774" refString="Lutz, B. (1950) Anfibios anuros da colecao Adolpho Lutz do Instituto Oswaldo Cruz. V / Frogs in the Adolpho Lutz collection of the Instituto Oswaldo Cruz. V. Memorias do Instituto Oswaldo Cruz, Rio de Janeiro, 48, 599 - 637." type="journal article">Lutz (1950)</bibRefCitation>
: Transferred from
<taxonomicName id="4C454D68FFFEFF9BC0FAFF6A427DFF39" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[735,848,152,174]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
<emphasis id="B931EAF9FFFEFF9BC0FAFF6A427DFF39" box="[735,848,152,174]" italics="true" pageId="7" pageNumber="408">Agalychnis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FF4442A5FF5B" blockId="7.[151,964,152,419]" box="[151,904,182,204]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FF44408BFF5B" box="[151,422,182,204]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FF44400BFF5B" authorityName=", Wagler" authorityYear="1830" box="[151,294,182,204]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Phyllomedusa</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC382FF444341FF5B" box="[423,620,182,204]" pageId="7" pageNumber="408" refId="ref13009" refString="Funkhouser, A. (1957) Review of the neotropical frogs of the genus Phyllomedusa. Occasional papers of the Natural History Museum of Stanford University, 5, 1 - 90." type="journal article">Funkhouser (1957)</bibRefCitation>
: in part [BM.1902.5.27.26].
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FF2742A6FF7C" blockId="7.[151,964,152,419]" box="[151,907,213,235]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FF2740AAFF7C" box="[151,391,213,235]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FF274025FF7C" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[151,264,213,235]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Agalychnis</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC3ADFF274314FF7C" box="[392,569,213,235]" pageId="7" pageNumber="408" refId="ref12111" refString="Duellman, W. E. (1968) The genera of phyllomedusine frogs (Anura: Hylidae). University of Kansas Publications, Museum of Natural History, Kansas, 18, 1 - 10. https: // doi. org / 10.5962 / bhl. part. 19989" type="journal article">Duellman (1968)</bibRefCitation>
: Transferred from
<taxonomicName id="4C454D68FFFEFF9BC0D3FF2742A8FF7C" authorityName=", Wagler" authorityYear="1830" box="[758,901,213,235]" class="Amphibia" family="Hylidae" genus="Phyllomedusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
<emphasis id="B931EAF9FFFEFF9BC0D3FF2742A8FF7C" box="[758,901,213,235]" italics="true" pageId="7" pageNumber="408">Phyllomedusa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FF0643BFFE9D" blockId="7.[151,964,152,419]" box="[151,658,244,266]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FF0640AAFE9D" box="[151,391,244,266]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FF064025FE9D" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[151,264,244,266]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Agalychnis</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC3ADFF0643A2FE9D" box="[392,655,244,266]" pageId="7" pageNumber="408" refId="ref11733" refString="Cochran, D. M. &amp; Goin, C. J. (1970) Frogs of Colombia. Smithsonian Institution Press, Washington D. C., 655 pp." type="book">Cochran and Goin (1970)</bibRefCitation>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FEE04383FEBF" blockId="7.[151,964,152,419]" box="[151,686,274,296]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FEE040AAFEBF" box="[151,391,274,296]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FEE04025FEBF" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[151,264,274,296]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Agalychnis</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC3ADFEE04386FEBF" box="[392,683,274,296]" pageId="7" pageNumber="408" refId="ref14352" refString="Myers, C. W. &amp; Duellman, W. E. (1982) A new species of Hyla from Cerro Colorado, and other tree frog records and geographical notes from western Panama. American Museum Novitates, 2752, 1 - 32." type="journal article">Duellman and Myers (1982)</bibRefCitation>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FEC34282FED0" blockId="7.[151,964,152,419]" box="[151,943,305,327]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FEC340A5FED0" box="[151,392,305,327]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FEC34025FED0" box="[151,264,305,327]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Cruziohyla</taxonomicName>
calcarifer—
</emphasis>
Faivorvich
<emphasis id="B931EAF9FFFEFF9BC027FEC04302FED0" box="[514,559,305,327]" italics="true" pageId="7" pageNumber="408">et al</emphasis>
(2005): Transferred from
<taxonomicName id="4C454D68FFFEFF9BC11CFEC34287FED0" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[825,938,305,327]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
<emphasis id="B931EAF9FFFEFF9BC11CFEC34287FED0" box="[825,938,305,327]" italics="true" pageId="7" pageNumber="408">Agalychnis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FEA24312FEF1" blockId="7.[151,964,152,419]" box="[151,575,336,358]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FEA240A5FEF1" box="[151,392,336,358]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FEA24025FEF1" box="[151,264,336,358]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Cruziohyla</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC3ADFEA34316FEF1" box="[392,571,336,358]" pageId="7" pageNumber="408" refId="ref12836" refString="Frost, D. R., Grant, T., Faivorvich, J., Bain, R. H., Haas, A., Haddad, C. F. B., de Sa, R. O., Channing, A., Wilkinson, M., Donnellan, S. C., Raxworthy, C. J., Campbell, J. A., Blotto, B. L., Moler, P., Drewes, R. C., Nussbaum, R. A., Lynch, J. D., Green, D. M. &amp; Wheeler, W. C. (2006) THE AMPHIBIAN TREE OF LIFE. Bulletin of the American Museum of Natural History, 297, 1 - 291. https: // doi. org / 10.1206 / 0003 - 0090 (2006) 297 [0001: TATOL] 2.0. CO; 2" type="journal article">
Frost
<emphasis id="B931EAF9FFFEFF9BC3E7FEA340C2FEF1" box="[450,495,336,358]" italics="true" pageId="7" pageNumber="408">et al</emphasis>
(2006)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FE9C42E9FE13" blockId="7.[151,964,152,419]" box="[151,964,366,389]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FE9C40A5FE13" box="[151,392,366,388]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FE9C4025FE13" box="[151,264,366,388]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Cruziohyla</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC3ADFE9C4340FE13" box="[392,621,366,389]" pageId="7" pageNumber="408" refId="ref12541" refString="Faivovich, J., Haddad, C. F. B., Baeta, D., Jungfer, K-H., Alvares, G. F. R., Brandao, R. A., Sheil. C, Barrientos, L. S. Barrio- Amoro, C. L., Cruz, C. A. G. &amp; Wheeler, W. C. (2010) The phylogenetic relationships of the charismatic poster frogs, Phyllomedusinae (Anura, Hylidae). Cladistics, 26 (3), 227 - 261. https: // doi. org / 10.1111 / j. 1096 - 0031.2009.00287. x" type="journal article">
Faivovich
<emphasis id="B931EAF9FFFEFF9BC3D1FE824303FE13" box="[500,558,366,389]" italics="true" pageId="7" pageNumber="408">et al (</emphasis>
2010)
</bibRefCitation>
: in part [
<accessionNumber id="9416AB08FFFEFF9BC0E1FE9C4217FE13" box="[708,826,366,388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ366229" pageId="7" pageNumber="408">GQ366229</accessionNumber>
,
<accessionNumber id="9416AB08FFFEFF9BC161FE9C4297FE13" box="[836,954,366,388]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/GQ366230" pageId="7" pageNumber="408">GQ366230</accessionNumber>
].
</paragraph>
<paragraph id="8BFA36EBFFFEFF9BC2B2FE7F4213FE34" blockId="7.[151,964,152,419]" box="[151,830,397,419]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2B2FE7F40A5FE34" box="[151,392,397,419]" italics="true" pageId="7" pageNumber="408">
<taxonomicName id="4C454D68FFFEFF9BC2B2FE7F4025FE34" box="[151,264,397,419]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">Cruziohyla</taxonomicName>
calcarifer—
</emphasis>
<bibRefCitation id="EFD44B1AFFFEFF9BC3ADFE7F4344FE34" box="[392,617,397,419]" pageId="7" pageNumber="408" refId="ref11907" refString="Crawford, A. J., Lips, K. R. &amp; Bermingham, E. (2010) Epidemic disease decimates amphibian abundance, species diversity, and evolutionary history in the highlands of central Panama. Proceedings of the National Academy of Sciences, 107 (31), 13777 - 13782. https: // doi. org / 10.1073 / pnas. 0914115107" type="journal article">
Crawford
<emphasis id="B931EAF9FFFEFF9BC3CAFE7C4331FE34" box="[495,540,397,419]" italics="true" pageId="7" pageNumber="408">et al</emphasis>
(2010)
</bibRefCitation>
: in part [KRL 0800].
</paragraph>
</subSubSection>
<subSubSection id="C35F6560FFFEFF9BC2B2FE3D442DFE7F" box="[151,1280,463,488]" pageId="7" pageNumber="408" type="materials_examined">
<paragraph id="8BFA36EBFFFEFF9BC2B2FE3D442DFE7F" blockId="7.[151,1437,463,1034]" box="[151,1280,463,488]" pageId="7" pageNumber="408">
<materialsCitation id="3B2D3CB6FFFEFF9BC2B2FE3D442DFE7F" ID-GBIF-Occurrence="1914123842" box="[151,1280,463,488]" country="Ecuador" location="Female" municipality="Rio Durango" pageId="7" pageNumber="408" specimenCount="1" typeStatus="holotype">
<emphasis id="B931EAF9FFFEFF9BC2B2FE3D4020FE7F" bold="true" box="[151,269,463,488]" pageId="7" pageNumber="408">
<typeStatus id="54FE8849FFFEFF9BC2B2FE3D4025FE7F" box="[151,264,463,488]" pageId="7" pageNumber="408" type="holotype">Holotype</typeStatus>
:
</emphasis>
BM.1902.5.27.26 [BM.1942.2.24.22].
<location id="8E9A6030FFFEFF9BC0E3FE3D4234FE7F" LSID="urn:lsid:plazi:treatment:03EC87FDFFFFFF9BC2B2F8B043EAFB9D:8E9A6030FFFEFF9BC0E3FE3D4234FE7F" box="[710,793,463,488]" country="Ecuador" municipality="Rio Durango" name="Female" pageId="7" pageNumber="408">Female</location>
from
<collectingMunicipality id="6B9EAC91FFFEFF9BC178FE3D42DFFE7F" box="[861,1010,463,488]" pageId="7" pageNumber="408">Rio Durango</collectingMunicipality>
, Northwestern
<collectingCountry id="F352767BFFFEFF9BC6B9FE3D45D6FE7F" box="[1180,1275,463,488]" name="Ecuador" pageId="7" pageNumber="408">Ecuador</collectingCountry>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35F6560FFFEFF9BC2E2FE06408FFCC1" pageId="7" pageNumber="408" type="diagnosis">
<paragraph id="8BFA36EBFFFEFF9BC2E2FE06408FFCC1" blockId="7.[151,1437,463,1034]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2E2FE06406CFD9A" bold="true" box="[199,321,500,525]" pageId="7" pageNumber="408">Diagnosis.</emphasis>
<taxonomicName id="4C454D68FFFEFF9BC36DFE064316FD9B" baseAuthorityName="Boulenger" baseAuthorityYear="1902" box="[328,571,500,524]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="species" species="calcarifer">
<emphasis id="B931EAF9FFFEFF9BC36DFE064316FD9B" box="[328,571,500,524]" italics="true" pageId="7" pageNumber="408">Cruziohyla calcarifer</emphasis>
</taxonomicName>
is a moderate to large species of the genus
<taxonomicName id="4C454D68FFFEFF9BC606FE0645B2FD9B" box="[1059,1183,500,524]" class="Amphibia" family="Hylidae" genus="Cruziohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="408" phylum="Chordata" rank="genus">
<emphasis id="B931EAF9FFFEFF9BC606FE0645B2FD9B" box="[1059,1183,500,524]" italics="true" pageId="7" pageNumber="408">Cruziohyla</emphasis>
</taxonomicName>
, attaining a maximum SVL in males of
<quantity id="4CBD9B0EFFFEFF9BC37EFDE540EDFDB8" box="[347,448,535,560]" metricMagnitude="-2" metricUnit="m" metricValue="5.64" pageId="7" pageNumber="408" unit="mm" value="56.4">56.4 mm</quantity>
to
<quantity id="4CBD9B0EFFFEFF9BC3C0FDE54360FDB8" box="[485,589,535,560]" metricMagnitude="-2" metricUnit="m" metricValue="8.05" pageId="7" pageNumber="408" unit="mm" value="80.5">80.5 mm</quantity>
, and
<quantity id="4CBD9B0EFFFEFF9BC0AFFDE543F7FDB8" box="[650,730,535,560]" metricMagnitude="-2" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="408" unit="mm" value="57.0">57 mm</quantity>
to
<quantity id="4CBD9B0EFFFEFF9BC0DAFDEA4248FDB8" box="[767,869,536,560]" metricMagnitude="-2" metricUnit="m" metricValue="8.879999999999999" pageId="7" pageNumber="408" unit="mm" value="88.8">88.8 mm</quantity>
in females, distinguished by a combination of the following characters: (1) prominent dark brown ventral thigh markings present either side of the vent in both sexes; (2) small tympanum, diameter 4060% that of the eye; (3) fingers and toes moderately webbed, approximately two-thirds: modal webbing formula (fingers):
<date id="FFFB102BFFFEFF9BC0BEFD7443C9FD08" box="[667,740,646,671]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC0BEFD74438BFD08" bold="true" box="[667,678,646,671]" pageId="7" pageNumber="408">I</emphasis>
2—2
</date>
-
<date id="FFFB102BFFFEFF9BC0D5FD744236FD08" box="[752,795,646,671]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC0D5FD74422BFD08" bold="true" box="[752,774,646,671]" pageId="7" pageNumber="408">II</emphasis>
1
</date>
+—
<date id="FFFB102BFFFEFF9BC164FD754258FD08" box="[833,885,646,671]" pageId="7" pageNumber="408">
2
<emphasis id="B931EAF9FFFEFF9BC171FD744258FD08" bold="true" box="[852,885,646,671]" pageId="7" pageNumber="408">III</emphasis>
</date>
1 1/2—
<date id="FFFB102BFFFEFF9BC19FFD7542DDFD08" box="[954,1008,643,671]" bridgedPair="-" pageId="7" pageNumber="408">
2-
<emphasis id="B931EAF9FFFEFF9BC1F6FD7442DDFD08" bold="true" box="[979,1008,646,671]" pageId="7" pageNumber="408">IV</emphasis>
</date>
; modal webbing formula (toes):
<date id="FFFB102BFFFEFF9BC745FD744188FD51" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC745FD744446FD08" bold="true" box="[1376,1387,646,671]" pageId="7" pageNumber="408">I</emphasis>
1— 1
</date>
+
<date id="FFFB102BFFFEFF9BC297FD5F4067FD51" box="[178,330,685,710]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC297FD5F41E5FD51" bold="true" box="[178,200,685,710]" pageId="7" pageNumber="408">II</emphasis>
1—2
<emphasis id="B931EAF9FFFEFF9BC337FD5F4067FD51" bold="true" box="[274,330,685,710]" pageId="7" pageNumber="408">III 1</emphasis>
</date>
+—3
<date id="FFFB102BFFFEFF9BC3A0FD5F40CBFD51" box="[389,486,685,710]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC3A0FD5F4089FD51" bold="true" box="[389,420,685,710]" pageId="7" pageNumber="408">IV</emphasis>
3—2
</date>
-
<emphasis id="B931EAF9FFFEFF9BC3D5FD5F432FFD51" bold="true" box="[496,514,685,710]" pageId="7" pageNumber="408">V</emphasis>
; (4) in life, upper surfaces uniform green, often having small white or pale blue spots; (5) in life, yellow-coloured spot on the posterior rim of the eye when open; (6) short black bars to lateral surfaces of the flanks; (7) colour of undersides, hands, feet, concealed surfaces of thighs and flanks, orange; (8) short sloping snout; (9) calcar on heel, often terminating in two dermal processes with one projecting inward and one projecting outward.
</paragraph>
</subSubSection>
<subSubSection id="C35F6560FFFEFF9BC2E2FC93440DFC55" pageId="7" pageNumber="408" type="description">
<paragraph id="8BFA36EBFFFEFF9BC2E2FC93440DFC55" blockId="7.[151,1437,463,1034]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2E2FC934012FCED" bold="true" box="[199,319,865,890]" pageId="7" pageNumber="408">Variation.</emphasis>
Variation is noted in the extent of the outward facing calcar found on the heel: in all specimens from
<collectingCountry id="F352767BFFFEFF9BC2B2FC744039FC09" box="[151,276,901,926]" name="Costa Rica" pageId="7" pageNumber="408">Costa Rica</collectingCountry>
and
<collectingRegion id="4981F809FFFEFF9BC36AFC744085FC0A" box="[335,424,902,925]" country="Panama" name="Panama" pageId="7" pageNumber="408">
<collectingCountry id="F352767BFFFEFF9BC36AFC744085FC0A" box="[335,424,902,925]" name="Panama" pageId="7" pageNumber="408">Panama</collectingCountry>
</collectingRegion>
this feature is large, extensive, and marked with dark pigmentation; in some specimens from
<collectingCountry id="F352767BFFFEFF9BC2F1FC5B406FFC55" box="[212,322,937,962]" name="Colombia" pageId="7" pageNumber="408">Colombia</collectingCountry>
and
<collectingCountry id="F352767BFFFEFF9BC35CFC5B40FBFC55" box="[377,470,937,962]" name="Ecuador" pageId="7" pageNumber="408">Ecuador</collectingCountry>
the outward-facing calcar on the heel is highly reduced and more rounded.
</paragraph>
</subSubSection>
<subSubSection id="C35F6560FFFEFF9BC2E2FC3F43EAFB9D" pageId="7" pageNumber="408" type="distribution">
<paragraph id="8BFA36EBFFFEFF9BC2E2FC3F43EAFB9D" blockId="7.[151,1437,463,1034]" pageId="7" pageNumber="408">
<emphasis id="B931EAF9FFFEFF9BC2E2FC3F404FFC71" bold="true" box="[199,354,973,998]" pageId="7" pageNumber="408">Distribution:</emphasis>
Highly restricted populations occurring in northwest
<collectingCountry id="F352767BFFFEFF9BC1C3FC3F4568FC71" box="[998,1093,973,998]" name="Ecuador" pageId="7" pageNumber="408">Ecuador</collectingCountry>
, western
<collectingCountry id="F352767BFFFEFF9BC69FFC3F4406FC71" box="[1210,1323,973,998]" name="Colombia" pageId="7" pageNumber="408">Colombia</collectingCountry>
,
<collectingRegion id="4981F809FFFEFF9BC719FC3C44B5FC72" box="[1340,1432,974,997]" country="Panama" name="Panama" pageId="7" pageNumber="408">
<collectingCountry id="F352767BFFFEFF9BC719FC3C44B5FC72" box="[1340,1432,974,997]" name="Panama" pageId="7" pageNumber="408">Panama</collectingCountry>
</collectingRegion>
, southeastern
<collectingCountry id="F352767BFFFEFF9BC30FFC004089FB9D" box="[298,420,1009,1034]" name="Costa Rica" pageId="7" pageNumber="408">Costa Rica</collectingCountry>
(
<figureCitation id="137E2A6EFFFEFF9BC396FC03432BFB9D" box="[435,518,1009,1034]" captionStart="FIGURE 10" captionStartId="14.[151,250,1116,1138]" captionTargetBox="[176,1362,244,1071]" captionTargetId="figure@14.[151,1436,188,1097]" captionTargetPageId="14" captionText="FIGURE 10. Geographical distribution of Cruziohyla sylviae sp. n. and Cruziohyla calcarifer." httpUri="https://zenodo.org/record/1444894/files/figure.png" pageId="7" pageNumber="408">Fig. 10</figureCitation>
). See discussion.
</paragraph>
</subSubSection>
</treatment>
</document>