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<document id="2438001EB3FEBA50BF3519B8CD3ABC54" ID-DOI="10.11646/zootaxa.4748.2.11" ID-GBIF-Dataset="fd9edc0b-4e36-4fa6-b4e9-33599b22f984" ID-ISSN="1175-5326" ID-Zenodo-Dep="3698739" ID-ZooBank="B3F989CF-A760-46B2-8619-B0A633DAA0B6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1583482537524" checkinUser="plazi" docAuthor="Anichtchenko, Alexander &amp; Medina, Milton Norman" docDate="2020" docId="03EC87CD7B1BFFFFBDE7FF76FF19FAAE" docLanguage="en" docName="zootaxa.4748.2.11.pdf" docOrigin="Zootaxa 4748 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Thopeutica angulihumerosa Des. F. Cassola 1997" docType="treatment" docVersion="5" lastPageNumber="391" masterDocId="FFD5FFB57B1AFFFDBD70FF97FFD3FFF8" masterDocTitle="Contribution to the knowledge of three species of the genus Thopeutica Schaum, 1861 from the Philippines (Coleoptera: Carabidae: Cicindelinae)" masterLastPageNumber="395" masterPageNumber="389" pageNumber="390" updateTime="1698725122065" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B600ED96AF362E8F8056F36C876BB642">Contribution to the knowledge of three species of the genus Thopeutica Schaum, 1861 from the Philippines (Coleoptera: Carabidae: Cicindelinae)</mods:title>
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<treatment id="03EC87CD7B1BFFFFBDE7FF76FF19FAAE" ID-DOI="http://doi.org/10.5281/zenodo.3705122" ID-GBIF-Taxon="162462214" ID-Zenodo-Dep="3705122" LSID="urn:lsid:plazi:treatment:03EC87CD7B1BFFFFBDE7FF76FF19FAAE" httpUri="http://treatment.plazi.org/id/03EC87CD7B1BFFFFBDE7FF76FF19FAAE" lastPageId="2" lastPageNumber="391" pageId="1" pageNumber="390">
<subSubSection id="C35F65507B1BFFFCBDE7FF76FCC0FF03" box="[151,787,225,252]" pageId="1" pageNumber="390" type="nomenclature">
<paragraph id="8BFA36DB7B1BFFFCBDE7FF76FCC0FF03" blockId="1.[151,787,225,287]" box="[151,787,225,252]" pageId="1" pageNumber="390">
<heading id="D0B281B77B1BFFFCBDE7FF76FCC0FF03" bold="true" box="[151,787,225,252]" fontSize="11" level="1" pageId="1" pageNumber="390" reason="1">
<taxonomicName id="4C454D587B1BFFFCBDE7FF76FCC0FF03" ID-CoL="56JZ3" authority="(W. Horn, 1929)" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[151,787,225,252]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1BFFFCBDE7FF76FCC0FF03" bold="true" box="[151,787,225,252]" pageId="1" pageNumber="390">
<emphasis id="B931EAC97B1BFFFCBDE7FF76FEF3FF04" bold="true" box="[151,288,225,252]" italics="true" pageId="1" pageNumber="390">Thopeutica</emphasis>
(s.str.)
<emphasis id="B931EAC97B1BFFFCBC0CFF76FD90FF04" bold="true" box="[380,579,225,252]" italics="true" pageId="1" pageNumber="390">angulihumerosa</emphasis>
(W.
<bibRefCitation id="EFD44B2A7B1BFFFCBF0DFF75FCD8FF03" author="Horn, W." box="[637,779,225,252]" pageId="1" pageNumber="390" pagination="4 - 8" refId="ref3248" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae - Genera. Entomologisches Nachrichtenblatt, 3 (1), 4 - 8." type="journal article" year="1929">Horn, 1929</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B1BFFFCBDE7FE92FEC1FEE7" box="[151,274,261,287]" pageId="1" pageNumber="390" type="description">
<paragraph id="8BFA36DB7B1BFFFCBDE7FE92FEC1FEE7" blockId="1.[151,787,225,287]" box="[151,274,261,287]" pageId="1" pageNumber="390">
(
<figureCitation id="137E2A5E7B1BFFFCBDEFFE92FEDAFEE7" box="[159,265,261,287]" captionStart="FIGURES 1–2" captionStartId="1.[151,264,1997,2022]" captionTargetBox="[250,1334,738,1957]" captionTargetId="figure@1.[244,1343,730,1972]" captionTargetPageId="1" captionText="FIGURES 1–2. Habitus of paralectotypes of Thopeutica (s.str.) angulihumerosa: 1—male; 2—female." figureDoi="http://doi.org/10.5281/zenodo.3698741" httpUri="https://zenodo.org/record/3698741/files/figure.png" pageId="1" pageNumber="390">Figs. 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B1BFFFCBDE7FEDDFD29FE78" pageId="1" pageNumber="390" type="reference_group">
<paragraph id="8BFA36DB7B1BFFFCBDE7FEDDFD87FE9A" blockId="1.[151,762,330,385]" box="[151,596,330,354]" pageId="1" pageNumber="390">
=
<taxonomicName id="4C454D587B1BFFFCBDDCFEDDFD87FE9A" ID-CoL="5YT4R" authority="W. Horn, 1929" authorityName="W. Horn" authorityYear="1929" box="[172,596,330,354]" class="Insecta" family="Carabidae" genus="Cicindela" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1BFFFCBDDCFEDDFE6AFE9A" box="[172,441,330,354]" italics="true" pageId="1" pageNumber="390">Cicindela angulihumerosa</emphasis>
W.
<bibRefCitation id="EFD44B2A7B1BFFFCBC90FEDDFD87FE9A" author="Horn, W." box="[480,596,330,354]" pageId="1" pageNumber="390" pagination="4 - 8" refId="ref3248" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae - Genera. Entomologisches Nachrichtenblatt, 3 (1), 4 - 8." type="journal article" year="1929">Horn, 1929</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="8BFA36DB7B1BFFFCBDE7FEFEFD29FE78" blockId="1.[151,762,330,385]" box="[151,762,360,385]" pageId="1" pageNumber="390">
=
<taxonomicName id="4C454D587B1BFFFCBDDCFEFFFD44FE78" ID-CoL="8LP27" authority="Schule, 2017" authorityName="Schule" authorityYear="2017" box="[172,663,360,385]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="juergenwiesneri">
<emphasis id="B931EAC97B1BFFFCBDDCFEFFFECDFE78" box="[172,286,360,384]" italics="true" pageId="1" pageNumber="390">Thopeutica</emphasis>
(s.str.)
<emphasis id="B931EAC97B1BFFFCBC1AFEFEFDDFFE78" box="[362,524,361,384]" italics="true" pageId="1" pageNumber="390">juergenwiesneri</emphasis>
<bibRefCitation id="EFD44B2A7B1BFFFCBF62FEFFFD44FE78" author="Schule, P." box="[530,663,360,385]" pageId="1" pageNumber="390" pagination="119 - 120" refId="ref3326" refString="Schule, P. (2017) Thopeutica juergenwiesneri, a new species of tiger beetles from the Philippines (Coleoptera: Carabidae, Cicindelinae). Entomologische Zeitschrift, Schwanfeld, 127 (2), 119 - 120." type="journal article" year="2017">Schüle, 2017</bibRefCitation>
</taxonomicName>
<emphasis id="B931EAC97B1BFFFCBFEDFEFFFD29FE78" bold="true" box="[669,762,360,384]" pageId="1" pageNumber="390">
<taxonomicNameLabel id="A20257B27B1BFFFCBFEDFEFFFD29FE78" box="[669,762,360,384]" pageId="1" pageNumber="390">syn. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B1BFFFFBDE7FE38FDE2FCC2" lastPageId="2" lastPageNumber="391" pageId="1" pageNumber="390" type="materials_examined">
<paragraph id="8BFA36DB7B1BFFFCBDE7FE38FAECFD3D" blockId="1.[151,1437,430,709]" pageId="1" pageNumber="390">
<emphasis id="B931EAC97B1BFFFCBDE7FE38FE94FE31" bold="true" box="[151,327,430,457]" pageId="1" pageNumber="390">Type material.</emphasis>
<materialsCitation id="3B2D3C867B1BFFFCBC21FE38FD22FDCD" ID-GBIF-Occurrence="2574624211" collectionCode="V, DEI, SDEI" collectorName="Ehlers &amp; Type W. Horn" pageId="1" pageNumber="390" specimenCount="3" typeStatus="paralectotype">
<typeStatus id="54FE88797B1BFFFCBC21FE38FE3FFE31" box="[337,492,431,457]" pageId="1" pageNumber="390" type="paralectotype">Paralectotype</typeStatus>
, male—“Ins. Philipp. [hand-written] coll.
<collectorName id="26B0530D7B1BFFFCBEA2FE38FBC9FE31" box="[978,1050,431,457]" pageId="1" pageNumber="390">Ehlers</collectorName>
<collectionCode id="ED54AE1E7B1BFFFCB952FE38FBE5FE31" box="[1058,1078,431,457]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="390" type="Museum">V</collectionCode>
. de Poll [typed]” [white label], “
<collectorName id="26B0530D7B1BFFFCBDD1FE44FE96FE15" box="[161,325,467,493]" pageId="1" pageNumber="390">Type W. Horn</collectorName>
” [typed, white label], “
<typeStatus id="54FE88797B1BFFFCBF3CFE44FD6AFE15" box="[588,697,467,493]" pageId="1" pageNumber="390" type="syntype">Syntypus</typeStatus>
” [red label], “
<collectionCode id="ED54AE1E7B1BFFFCBE28FE44FC5AFE15" box="[856,905,467,493]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="1" pageNumber="390" type="Other type of research institution/biorepository">DEI</collectionCode>
sub
<emphasis id="B931EAC97B1BFFFCBEB0FE44FBEBFE15" box="[960,1080,467,493]" italics="true" pageId="1" pageNumber="390">angulihum</emphasis>
.” [typed, white label], “PARA-
<typeStatus id="54FE88797B1BFFFCBDE7FE60FE9AFDE9" box="[151,329,503,529]" pageId="1" pageNumber="390" type="lectotype">LECTOTYPUS</typeStatus>
<taxonomicName id="4C454D587B1BFFFCBC24FE60FB69FDE9" authority="(W. Horn, 1929) Des. F. Cassola, 1997" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[340,1210,503,529]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1BFFFCBC24FE60FE01FDE9" box="[340,466,503,529]" italics="true" pageId="1" pageNumber="390">Thopeutica</emphasis>
(s. str.)
<emphasis id="B931EAC97B1BFFFCBF46FE60FD38FDE9" box="[566,747,503,529]" italics="true" pageId="1" pageNumber="390">angulihumerosa</emphasis>
(W.
<bibRefCitation id="EFD44B2A7B1BFFFCBE57FE60FC78FDE9" author="Horn, W." box="[807,939,503,529]" pageId="1" pageNumber="390" pagination="4 - 8" refId="ref3248" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae - Genera. Entomologisches Nachrichtenblatt, 3 (1), 4 - 8." type="journal article" year="1929">Horn, 1929</bibRefCitation>
) Des. F. Cassola, 1997
</taxonomicName>
” [typed, red label], “
<collectionCode id="ED54AE1E7B1BFFFCBDD1FD8CFF01FDCD" box="[161,210,539,565]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="1" pageNumber="390" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg Col—12139” [green label] (
<collectionCode id="ED54AE1E7B1BFFFCBFD5FD8CFD3AFDCD" box="[677,745,539,565]" pageId="1" pageNumber="390">SDEI</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B2D3C867B1BFFFCBF8BFD8CFAE8FD3D" ID-GBIF-Occurrence="2574624209" collectionCode="DEI, SDEI" collectorName="Type W. Horn &amp; Coll. W. Horn" pageId="1" pageNumber="390" specimenCount="3" typeStatus="paralectotype">
<typeStatus id="54FE88797B1BFFFCBF8BFD8CFC70FDCD" box="[763,931,539,565]" pageId="1" pageNumber="390" type="paralectotype">Paralectotypes</typeStatus>
: female—same labels, but “
<collectionCode id="ED54AE1E7B1BFFFCB9A7FD8CFADBFDCD" box="[1239,1288,539,565]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="1" pageNumber="390" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg Col—12140” [green label] (
<collectionCode id="ED54AE1E7B1BFFFCBCA8FDA8FDCFFDA1" box="[472,540,575,601]" pageId="1" pageNumber="390">SDEI</collectionCode>
); female—“Island Samar Baker”[typed, white label], “
<collectorName id="26B0530D7B1BFFFCB9E4FDA8FAE8FDA1" box="[1172,1339,575,601]" pageId="1" pageNumber="390">Type W. Horn</collectorName>
” [typed, white label], “Mus. Washington” [handwritten], “
<collectorName id="26B0530D7B1BFFFCBFB0FDF4FCB3FD85" box="[704,864,611,637]" pageId="1" pageNumber="390">Coll. W. Horn</collectorName>
<collectionCode id="ED54AE1E7B1BFFFCBE16FDF4FC46FD85" box="[870,917,611,637]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="1" pageNumber="390" type="Other type of research institution/biorepository">DEI</collectionCode>
Eberswalde” [typed, white label], “
<typeStatus id="54FE88797B1BFFFCB855FDF4FA41FD85" box="[1317,1426,611,637]" pageId="1" pageNumber="390" type="syntype">Syntypus</typeStatus>
” [red label], “
<emphasis id="B931EAC97B1BFFFCBC5BFD10FE3AFD59" box="[299,489,647,673]" italics="true" pageId="1" pageNumber="390">anguli-humerosa</emphasis>
W.H.” [hand-written], “
<typeStatus id="54FE88797B1BFFFCBF8EFD10FC2FFD59" box="[766,1020,647,673]" pageId="1" pageNumber="390" type="paralectotype">PARALECTOTYPUS</typeStatus>
<taxonomicName id="4C454D587B1BFFFCB976FD10FD99FD3D" authority="(W. Horn, 1929) Des. F. Cassola, 1997" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1BFFFCB976FD10FB57FD59" box="[1030,1156,647,673]" italics="true" pageId="1" pageNumber="390">Thopeutica</emphasis>
(s. str.)
<emphasis id="B931EAC97B1BFFFCB997FD10FA4FFD59" box="[1255,1436,647,673]" italics="true" pageId="1" pageNumber="390">angulihumerosa</emphasis>
(W.
<bibRefCitation id="EFD44B2A7B1BFFFCBDB6FD3CFE94FD3D" author="Horn, W." box="[198,327,683,709]" pageId="1" pageNumber="390" pagination="4 - 8" refId="ref3248" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae - Genera. Entomologisches Nachrichtenblatt, 3 (1), 4 - 8." type="journal article" year="1929">Horn, 1929</bibRefCitation>
) Des. F. Cassola, 1997
</taxonomicName>
” [red label], “
<collectionCode id="ED54AE1E7B1BFFFCBF97FD3CFCCBFD3D" box="[743,792,683,709]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33211" lsid="urn:lsid:biocol.org:col:33211" name="Senckenberg Deutsches Entomologisches Institut" pageId="1" pageNumber="390" type="Other type of research institution/biorepository">DEI</collectionCode>
Müncheberg Col—12138” [green label] (
<collectionCode id="ED54AE1E7B1BFFFCB99FFD3CFAE0FD3D" box="[1263,1331,683,709]" pageId="1" pageNumber="390">SDEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<caption id="DF3A66537B1BFFFCBDE7F85AFB78F81E" ID-DOI="http://doi.org/10.5281/zenodo.3698741" ID-Zenodo-Dep="3698741" box="[151,1195,1997,2022]" httpUri="https://zenodo.org/record/3698741/files/figure.png" pageId="1" pageNumber="390" startId="1.[151,264,1997,2022]" targetBox="[250,1334,738,1957]" targetPageId="1">
<paragraph id="8BFA36DB7B1BFFFCBDE7F85AFB78F81E" blockId="1.[151,1195,1997,2022]" box="[151,1195,1997,2022]" pageId="1" pageNumber="390">
<emphasis id="B931EAC97B1BFFFCBDE7F85AFEE9F81E" bold="true" box="[151,314,1997,2022]" pageId="1" pageNumber="390">FIGURES 1–2.</emphasis>
Habitus of paralectotypes of
<taxonomicName id="4C454D587B1BFFFCBF15F859FC14F81E" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[613,967,1998,2022]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="390" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B1BFFFCBF15F859FD04F81E" box="[613,727,1998,2022]" italics="true" pageId="1" pageNumber="390">Thopeutica</emphasis>
(s.str.)
<emphasis id="B931EAC97B1BFFFCBE53F859FC14F81E" box="[803,967,1998,2022]" italics="true" pageId="1" pageNumber="390">angulihumerosa</emphasis>
</taxonomicName>
: 1—male; 2—female.
</paragraph>
</caption>
<paragraph id="8BFA36DB7B18FFFFBDB7FF0FFE0EFF03" blockId="2.[151,1437,152,1366]" pageId="2" pageNumber="391">
<emphasis id="B931EAC97B18FFFFBDB7FF0FFE6EFF4B" bold="true" box="[199,445,152,179]" pageId="2" pageNumber="391">Additional material:</emphasis>
“
<materialsCitation id="3B2D3C867B18FFFFBCBFFF0EFB1CFF4B" ID-GBIF-Occurrence="2574624206" box="[463,1231,152,179]" collectingDate="2017-03" location="Lope de Vega" municipality="Eastern Visayas" pageId="2" pageNumber="391" specimenCount="1" specimenCount-female="1" stateProvince="Samar">
<collectingRegion id="4981F8397B18FFFFBCBFFF0EFDC8FF4B" box="[463,539,153,179]" country="Philippines" name="Samar" pageId="2" pageNumber="391">Samar</collectingRegion>
,
<collectingMunicipality id="6B9EACA17B18FFFFBF57FF0EFD0EFF4A" box="[551,733,152,179]" pageId="2" pageNumber="391">Eastern Visayas</collectingMunicipality>
,
<location id="8E9A60007B18FFFFBF99FF0EFC54FF4B" LSID="urn:lsid:plazi:treatment:03EC87CD7B1BFFFFBDE7FF76FF19FAAE:8E9A60007B18FFFFBF99FF0EFC54FF4B" box="[745,903,152,179]" municipality="Eastern Visayas" name="Lope de Vega" pageId="2" pageNumber="391" stateProvince="Samar">Lope de Vega</location>
,
<date id="FFFB101B7B18FFFFBEE3FF0EFC23FF4B" box="[915,1008,153,179]" pageId="2" pageNumber="391" value="2017-03">
<collectingDate id="EFBFE9F37B18FFFFBEE3FF0EFC23FF4B" box="[915,1008,153,179]" pageId="2" pageNumber="391" value="2017-03">III.2017</collectingDate>
</date>
” (
<specimenCount id="9D43FD527B18FFFFB97EFF0EFBBDFF4A" box="[1038,1134,152,179]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS)
</materialsCitation>
;
<materialsCitation id="3B2D3C867B18FFFFB9ACFF0FFE1CFF2F" ID-GBIF-Occurrence="2574624207" collectingDate="2016-05" collectionCode="V" location="Lope de Vega" municipality="Eastern Visayas" pageId="2" pageNumber="391" specimenCount="4" specimenCount-female="2" specimenCount-male="2" stateProvince="Samar">
<emphasis id="B931EAC97B18FFFFB9ACFF0FFAC1FF4A" box="[1244,1298,152,178]" italics="true" pageId="2" pageNumber="391">idem</emphasis>
, but
<date id="FFFB101B7B18FFFFB83DFF0FFA4EFF4A" box="[1357,1437,152,178]" pageId="2" pageNumber="391" value="2016-05">
<collectingDate id="EFBFE9F37B18FFFFB83DFF0FFA4EFF4A" box="[1357,1437,152,178]" pageId="2" pageNumber="391" value="2016-05">
<collectionCode id="ED54AE1E7B18FFFFB83DFF0FFAB2FF4A" box="[1357,1377,152,178]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="2" pageNumber="391" type="Museum">V</collectionCode>
.2016
</collectingDate>
</date>
(
<specimenCount id="9D43FD527B18FFFFBDD0FF2AFF2AFF2E" box="[160,249,188,215]" pageId="2" pageNumber="391" type="male">2 males</specimenCount>
,
<specimenCount id="9D43FD527B18FFFFBC74FF2AFEA3FF2E" box="[260,368,188,215]" pageId="2" pageNumber="391" type="female">2 females</specimenCount>
in cRS)
</materialsCitation>
;
<materialsCitation id="3B2D3C867B18FFFFBCAAFF2BFC1EFF2F" ID-GBIF-Occurrence="2574624203" box="[474,973,188,215]" collectingDate="2016-06" collectionCode="VI" location="Lope de Vega" municipality="Eastern Visayas" pageId="2" pageNumber="391" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Samar">
<emphasis id="B931EAC97B18FFFFBCAAFF2BFDC3FF2E" box="[474,528,188,214]" italics="true" pageId="2" pageNumber="391">idem</emphasis>
, but
<date id="FFFB101B7B18FFFFBF38FF2BFD76FF2E" box="[584,677,188,214]" pageId="2" pageNumber="391" value="2016-06">
<collectingDate id="EFBFE9F37B18FFFFBF38FF2BFD76FF2E" box="[584,677,188,214]" pageId="2" pageNumber="391" value="2016-06">
<collectionCode id="ED54AE1E7B18FFFFBF38FF2BFDB5FF2E" box="[584,614,188,214]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="2" pageNumber="391">VI</collectionCode>
.2016
</collectingDate>
</date>
(
<specimenCount id="9D43FD527B18FFFFBFC7FF2AFCD0FF2E" box="[695,771,188,215]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
,
<specimenCount id="9D43FD527B18FFFFBE7EFF2AFCBCFF2E" box="[782,879,188,215]" pageId="2" pageNumber="391" type="female">1 female</specimenCount>
in cRS)
</materialsCitation>
;
<materialsCitation id="3B2D3C867B18FFFFBEA9FF2BFA80FF2F" ID-GBIF-Occurrence="2574624205" box="[985,1363,188,215]" collectingDate="2018-02" collectionCode="VI" location="Lope de Vega" municipality="Eastern Visayas" pageId="2" pageNumber="391" specimenCount="1" specimenCount-male="1" stateProvince="Samar">
<emphasis id="B931EAC97B18FFFFBEA9FF2BFBDCFF2E" box="[985,1039,188,214]" italics="true" pageId="2" pageNumber="391">idem</emphasis>
, but
<date id="FFFB101B7B18FFFFB938FF2AFB49FF2F" box="[1096,1178,189,215]" pageId="2" pageNumber="391" value="2018-02">
<collectingDate id="EFBFE9F37B18FFFFB938FF2AFB49FF2F" box="[1096,1178,189,215]" pageId="2" pageNumber="391" value="2018-02">II.2018</collectingDate>
</date>
(
<specimenCount id="9D43FD527B18FFFFB9DBFF2AFB27FF2E" box="[1195,1268,188,215]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
in cRS)
</materialsCitation>
;
<materialsCitation id="3B2D3C867B18FFFFB82FFF2BFE0AFF03" ID-GBIF-Occurrence="2574624208" collectingDate="2018-11" collectionCode="VI" location="Lope de Vega" municipality="Eastern Visayas" pageId="2" pageNumber="391" specimenCount="1" specimenCount-male="1" stateProvince="Samar">
<emphasis id="B931EAC97B18FFFFB82FFF2BFA46FF2E" box="[1375,1429,188,214]" italics="true" pageId="2" pageNumber="391">idem</emphasis>
, but
<date id="FFFB101B7B18FFFFBDB2FF76FECDFF03" box="[194,286,225,251]" pageId="2" pageNumber="391" value="2018-11">
<collectingDate id="EFBFE9F37B18FFFFBDB2FF76FECDFF03" box="[194,286,225,251]" pageId="2" pageNumber="391" value="2018-11">XI.2018</collectingDate>
</date>
(
<specimenCount id="9D43FD527B18FFFFBC41FF76FEA9FF02" box="[305,378,224,251]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
in cRS)
</materialsCitation>
.
</paragraph>
<paragraph id="8BFA36DB7B18FFFFBDB7FE93FDE2FCC2" blockId="2.[151,1437,152,1366]" pageId="2" pageNumber="391">
Species was described by W.
<bibRefCitation id="EFD44B2A7B18FFFFBF7BFE92FD46FEE7" author="Horn, W." box="[523,661,261,287]" pageId="2" pageNumber="391" pagination="4 - 8" refId="ref3248" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae - Genera. Entomologisches Nachrichtenblatt, 3 (1), 4 - 8." type="journal article" year="1929">Horn (1929)</bibRefCitation>
based on the specimens collected by Baker in
<collectingRegion id="4981F8397B18FFFFB9E5FE92FB0EFEE7" box="[1173,1245,261,287]" country="Philippines" name="Samar" pageId="2" pageNumber="391">Samar</collectingRegion>
Island. The story with the allocation of types was carefully studied and described in detail by Cassola and Ward (2004). Recently a closely related species,
<taxonomicName id="4C454D587B18FFFFBCD5FEDAFDA6FE9E" authorityName="Schule" authorityYear="2017" box="[421,629,333,358]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="juergenwiesneri">
<emphasis id="B931EAC97B18FFFFBCD5FEDAFDA6FE9E" box="[421,629,333,358]" italics="true" pageId="2" pageNumber="391">T. juergenwiesneri</emphasis>
</taxonomicName>
, was described by Peter
<bibRefCitation id="EFD44B2A7B18FFFFBEE5FEDBFBE4FE9F" author="Schule, P." box="[917,1079,332,359]" pageId="2" pageNumber="391" pagination="119 - 120" refId="ref3326" refString="Schule, P. (2017) Thopeutica juergenwiesneri, a new species of tiger beetles from the Philippines (Coleoptera: Carabidae, Cicindelinae). Entomologische Zeitschrift, Schwanfeld, 127 (2), 119 - 120." type="journal article" year="2017">Schüle (2017)</bibRefCitation>
, from
<collectingRegion id="4981F8397B18FFFFB9F3FEDAFB18FE9F" box="[1155,1227,333,359]" country="Philippines" name="Samar" pageId="2" pageNumber="391">Samar</collectingRegion>
island on a single specimen (
<typeStatus id="54FE88797B18FFFFBC62FEE7FEACFE72" box="[274,383,368,394]" pageId="2" pageNumber="391" type="holotype">Holotype</typeStatus>
, male: „
<collectingCountry id="F352764B7B18FFFFBCACFEE6FD54FE73" box="[476,647,369,395]" name="Philippines" pageId="2" pageNumber="391">PHILIPPINES</collectingCountry>
,
<collectingRegion id="4981F8397B18FFFFBFE4FEE6FD08FE73" box="[660,731,369,395]" country="Philippines" name="Samar" pageId="2" pageNumber="391">Samar</collectingRegion>
\ Central Visayas, Lope de Vega\
<date id="FFFB101B7B18FFFFB92DFEE7FB1FFE72" box="[1117,1228,368,394]" pageId="2" pageNumber="391" value="2016-07">July 2016</date>
\ local collector”). In differencial diagnosis was indicated that new species is similar to
<taxonomicName id="4C454D587B18FFFFBEC1FE02FB57FE56" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[945,1156,404,430]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBEC1FE02FB57FE56" box="[945,1156,404,430]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
in body size and elytral pattern, with humeral macula ochre and ivory-white dorsal maculae, but easily distinguished by the much longer antennae in male which is surpassing the basal half in
<taxonomicName id="4C454D587B18FFFFBF8DFE4AFB33FE0F" authority="(Cassola &amp; Ward 2004)" baseAuthorityName="Cassola &amp; Ward" baseAuthorityYear="2004" box="[765,1248,476,503]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBF8DFE4AFC1DFE0E" box="[765,974,476,502]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
(Cassola &amp; Ward 2004)
</taxonomicName>
but reaching the apical fourth in
<taxonomicName id="4C454D587B18FFFFBC37FD96FDC6FDE2" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[327,533,513,538]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="juergenwiesneri" status="sp. nov.">
<emphasis id="B931EAC97B18FFFFBC37FD96FDC6FDE2" box="[327,533,513,538]" italics="true" pageId="2" pageNumber="391">T. juergenwiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B18FFFFBF6BFD96FD8AFDE3" bold="true" box="[539,601,513,539]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A20257B27B18FFFFBF6BFD96FD8AFDE3" box="[539,601,513,539]" pageId="2" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, and differently shaped aedeagus which is broader in lateral view, and with apex straighter in
<taxonomicName id="4C454D587B18FFFFBC10FDB2FDFCFDC6" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[352,559,549,574]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="juergenwiesneri" status="sp. nov.">
<emphasis id="B931EAC97B18FFFFBC10FDB2FDFCFDC6" box="[352,559,549,574]" italics="true" pageId="2" pageNumber="391">T. juergenwiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B18FFFFBF46FDB2FDA6FDC7" bold="true" box="[566,629,549,575]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A20257B27B18FFFFBF46FDB2FDA6FDC7" box="[566,629,549,575]" pageId="2" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
”. We examined
<typeStatus id="54FE88797B18FFFFBE5EFDB3FC03FDC6" box="[814,976,548,574]" pageId="2" pageNumber="391" type="paralectotype">paralectotypes</typeStatus>
of
<taxonomicName id="4C454D587B18FFFFBE85FDB2FB15FDC6" baseAuthorityName="Cassola &amp; Ward" baseAuthorityYear="2004" box="[1013,1222,548,574]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBE85FDB2FB15FDC6" box="[1013,1222,548,574]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
(one male and two females) stored in the
<collectionCode id="ED54AE1E7B18FFFFBCEAFDDEFE0BFD9B" box="[410,472,585,611]" pageId="2" pageNumber="391">SDEI</collectionCode>
Museum. After careful examination, we came to the conclusion that the mentioned distinguishing feature of the length of the antennae is erroneous (
<figureCitation id="137E2A5E7B18FFFFBE1EFDFBFC08FD7F" box="[878,987,620,647]" captionStart="FIGURES 3–4" captionStartId="3.[151,264,1944,1969]" captionTargetBox="[399,1187,190,1907]" captionTargetId="figure@3.[384,1203,181,1919]" captionTargetPageId="3" captionText="FIGURES 3–4. Comparative length of antennae of males of Thopeutica spp.: 3—holotype of T. (s.str.) juergenwiesneri syn." figureDoi="http://doi.org/10.5281/zenodo.3698743" httpUri="https://zenodo.org/record/3698743/files/figure.png" pageId="2" pageNumber="391">Figs. 3–4</figureCitation>
). Length of antennae is identical in both species. We noticed slight differences in the shape of apex of the aedeagus between the single known specimen of
<taxonomicName id="4C454D587B18FFFFBDE7FD22FEB6FD36" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[151,357,693,718]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="juergenwiesneri">
<emphasis id="B931EAC97B18FFFFBDE7FD22FEB6FD36" box="[151,357,693,718]" italics="true" pageId="2" pageNumber="391">T. juergenwiesneri</emphasis>
</taxonomicName>
, and a
<typeStatus id="54FE88797B18FFFFBCC4FD23FD98FD36" box="[436,587,692,718]" pageId="2" pageNumber="391" type="paralectotype">paralectotype</typeStatus>
of
<taxonomicName id="4C454D587B18FFFFBF1EFD22FCEDFD36" baseAuthorityName="Cassola &amp; Ward" baseAuthorityYear="2004" box="[622,830,692,718]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBF1EFD22FCEDFD36" box="[622,830,692,718]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
. However, these differences are weaker than indicated in the description (Figs. 7–9). Both species were described from
<collectingRegion id="4981F8397B18FFFFBEF2FD4EFC18FD0B" box="[898,971,729,755]" country="Philippines" name="Samar" pageId="2" pageNumber="391">Samar</collectingRegion>
, and the coloration of body and legs are identical. Hence, the new synonymy
<taxonomicName id="4C454D587B18FFFFBF48FD6AFC3CFCEF" authority="Schule, 2017" authorityName="Schule" authorityYear="2017" box="[568,1007,764,791]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="juergenwiesneri">
<emphasis id="B931EAC97B18FFFFBF48FD6AFD9EFCEE" box="[568,589,765,790]" italics="true" pageId="2" pageNumber="391">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B18FFFFBFD2FD6AFC86FCEE" box="[674,853,765,790]" italics="true" pageId="2" pageNumber="391">juergenwiesneri</emphasis>
<bibRefCitation id="EFD44B2A7B18FFFFBE2DFD6BFC3CFCEF" author="Schule, P." box="[861,1007,764,791]" pageId="2" pageNumber="391" pagination="119 - 120" refId="ref3326" refString="Schule, P. (2017) Thopeutica juergenwiesneri, a new species of tiger beetles from the Philippines (Coleoptera: Carabidae, Cicindelinae). Entomologische Zeitschrift, Schwanfeld, 127 (2), 119 - 120." type="journal article" year="2017">Schüle, 2017</bibRefCitation>
</taxonomicName>
<emphasis id="B931EAC97B18FFFFBE87FD6AFBA5FCEF" bold="true" box="[1015,1142,765,791]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A20257B27B18FFFFBE87FD6AFB8CFCEF" box="[1015,1119,765,791]" pageId="2" pageNumber="391">syn. nov.</taxonomicNameLabel>
=
</emphasis>
<taxonomicName id="4C454D587B18FFFFB90EFD6AFE9CFCC3" authority="(W. Horn, 1929)" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFB90EFD6AFB40FCEE" box="[1150,1171,765,790]" italics="true" pageId="2" pageNumber="391">T.</emphasis>
(s.str.)
<emphasis id="B931EAC97B18FFFFB997FD6BFA4FFCEE" box="[1255,1436,764,790]" italics="true" pageId="2" pageNumber="391">angulihumerosa</emphasis>
(W.
<bibRefCitation id="EFD44B2A7B18FFFFBDB6FCB6FE94FCC3" author="Horn, W." box="[198,327,801,827]" pageId="2" pageNumber="391" pagination="4 - 8" refId="ref3248" refString="Horn, W. (1929) Ueber einige orientalische und aethiopische Cicindelinae - Genera. Entomologisches Nachrichtenblatt, 3 (1), 4 - 8." type="journal article" year="1929">Horn, 1929</bibRefCitation>
)
</taxonomicName>
is hereby proposed.
</paragraph>
</subSubSection>
<subSubSection id="C35F65507B18FFFFBDB7FCD3FF19FAAE" pageId="2" pageNumber="391" type="discussion">
<paragraph id="8BFA36DB7B18FFFFBDB7FCD3FB3FFBCF" blockId="2.[151,1437,152,1366]" pageId="2" pageNumber="391">
<emphasis id="B931EAC97B18FFFFBDB7FCD3FE9CFCA7" bold="true" box="[199,335,836,863]" pageId="2" pageNumber="391">Comments.</emphasis>
We would like to draw attention on the structure of the labrum. This trait is traditionally used in the taxonomy of
<taxonomicName id="4C454D587B18FFFFBC25FCFFFE36FC7A" authorityName="Latreille" authorityYear="1802" box="[341,485,872,898]" class="Insecta" family="Carabidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="subFamily" subFamily="Cicindelinae">Cicindelinae</taxonomicName>
, and often used as one of the main traits in distinguishing between species. However, many species exhibit strong variability in its structure. The number of setae, the shape of the anterior margin, and even the number of teeth can vary. This was found by us, for example, in species of genera
<taxonomicName id="4C454D587B18FFFFB9F7FC27FAD6FC32" authorityName="Schaum" authorityYear="1861" box="[1159,1285,944,970]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC97B18FFFFB9F7FC27FAD6FC32" box="[1159,1285,944,970]" italics="true" pageId="2" pageNumber="391">Thopeutica</emphasis>
</taxonomicName>
and
<taxonomicName id="4C454D587B18FFFFB84AFC27FEECFC16" authority="Latreille, 1816" authorityName="Latreille" authorityYear="1816" class="Insecta" family="Carabidae" genus="Therates" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="genus">
<emphasis id="B931EAC97B18FFFFB84AFC27FA4FFC32" box="[1338,1436,944,970]" italics="true" pageId="2" pageNumber="391">Therates</emphasis>
Latreille, 1816
</taxonomicName>
. The value of this feature is clearly overestimated and it should be used with caution. Following the tradition, we provided description of the structure of the labrum (
<figureCitation id="137E2A5E7B18FFFFBEF7FC6FFBC2FBEB" box="[903,1041,1016,1043]" captionStart="FIGURES 10–12" captionStartId="4.[151,264,1855,1880]" captionTargetBox="[170,1418,992,1825]" captionTargetId="figure@4.[163,1425,988,1830]" captionTargetPageId="4" captionText="FIGURES 10–12. Labrum of Thopeutica spp. (a—males, b—females): 10—paralectotype of T. (s.str.) angulihumerosa; 11— holotype of T. (s.str.) juergenwiesneri syn. nov.; 12—holotype and paratype of T. (s.str.) wiesneri sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3698747" httpUri="https://zenodo.org/record/3698747/files/figure.png" pageId="2" pageNumber="391">Figs. 10–12</figureCitation>
), however, we emphasized that its shape varies in the males and females in both species (
<taxonomicName id="4C454D587B18FFFFBF8AFB8AFC18FBCE" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[762,971,1052,1078]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBF8AFB8AFC18FBCE" box="[762,971,1052,1078]" italics="true" pageId="2" pageNumber="391">T. angulihumerosa</emphasis>
</taxonomicName>
and
<taxonomicName id="4C454D587B18FFFFB971FB8AFBAFFBCE" authority="Anichtchenko &amp; Medina, 2020" authorityName="Anichtchenko &amp; Medina" authorityYear="2020" box="[1025,1148,1053,1078]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="wiesneri" status="sp. nov.">
<emphasis id="B931EAC97B18FFFFB971FB8AFBAFFBCE" box="[1025,1148,1053,1078]" italics="true" pageId="2" pageNumber="391">T. wiesneri</emphasis>
</taxonomicName>
<emphasis id="B931EAC97B18FFFFB9F3FB8AFB0FFBCF" bold="true" box="[1155,1244,1053,1079]" pageId="2" pageNumber="391">
<taxonomicNameLabel id="A20257B27B18FFFFB9F3FB8AFB0FFBCF" box="[1155,1244,1053,1079]" pageId="2" pageNumber="391" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
).
</paragraph>
<paragraph id="8BFA36DB7B18FFFFBDB7FBD7FF19FAAE" blockId="2.[151,1437,152,1366]" pageId="2" pageNumber="391">
<taxonomicName id="4C454D587B18FFFFBDB7FBD7FDD3FBA2" authorityName="Des. F. Cassola" authorityYear="1997" baseAuthorityName="W. Horn" baseAuthorityYear="1929" box="[199,512,1088,1114]" class="Insecta" family="Carabidae" genus="Thopeutica" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="391" phylum="Arthropoda" rank="species" species="angulihumerosa">
<emphasis id="B931EAC97B18FFFFBDB7FBD7FDD3FBA2" box="[199,512,1088,1114]" italics="true" pageId="2" pageNumber="391">Thopeutica angulihumerosa</emphasis>
</taxonomicName>
was cited by Cassola and Ward (2004) from
<collectingRegion id="4981F8397B18FFFFBE80FBD6FBEAFBA3" box="[1008,1081,1089,1115]" country="Philippines" name="Samar" pageId="2" pageNumber="391">Samar</collectingRegion>
, Mindanao and
<collectingRegion id="4981F8397B18FFFFB999FBD6FAFFFBA3" box="[1257,1324,1089,1115]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="391">Leyte</collectingRegion>
. This data was later included in the checklist of Tiger beetles of the
<collectingCountry id="F352764B7B18FFFFBE52FBF3FC73FB86" box="[802,928,1124,1150]" name="Philippines" pageId="2" pageNumber="391">Philippines</collectingCountry>
(
<bibRefCitation id="EFD44B2A7B18FFFFBEC0FBF3FB56FB87" author="Cabras, A. &amp; Cabigas, E. &amp; Wiesner, J." box="[944,1157,1124,1151]" pageId="2" pageNumber="391" pagination="188 - 201" refId="ref3090" refString="Cabras, A., Cabigas, E. &amp; Wiesner, J. (2016) Updated Checklist of Tiger Beetles in the Philippines (Coleoptera, Carabidae, Cicindelinae) (131 st. Contribution towards the knowledge of Cicindelinae). Lambillionea, 116 (3), 188 - 201." type="journal article" year="2016">
Cabras
<emphasis id="B931EAC97B18FFFFB977FBF2FB92FB86" box="[1031,1089,1124,1150]" italics="true" pageId="2" pageNumber="391">et al.</emphasis>
2016
</bibRefCitation>
). The following material used by Cassola and Ward (2004) requires re-verification of determination: “Philippine Is., Mindanao,
<collectingRegion id="4981F8397B18FFFFB856FB1EFABCFB5B" box="[1318,1391,1161,1187]" country="Philippines" name="Davao" pageId="2" pageNumber="391">Davao</collectingRegion>
Pr., Todaya, June 3,1974, Fr. E. Schoenig” (1 fem., CMNH). “Boston, Mt. Agtuuganon, Camp 55, 29.V–7.VI.
<quantity id="4CBD9B3E7B18FFFFB82CFB3BFF3EFB13" metricMagnitude="3" metricUnit="m" metricValue="1.996102" pageId="2" pageNumber="391" unit="m" value="1996.102">1996, 1020 m</quantity>
, light trap, R.A. Müller, A. Buenafe &amp; L. Gorostica” (2 fem., RMNH), and
<collectingRegion id="4981F8397B18FFFFB941FB46FBA6FB13" box="[1073,1141,1233,1259]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="391">Leyte</collectingRegion>
: “
<quantity id="4CBD9B3E7B18FFFFB9FBFB46FB17FB12" box="[1163,1220,1232,1259]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="2" pageNumber="391" unit="km" value="8.0">8 km</quantity>
N Baybay, ViSCA, Pacubasan River, Bato,
<quantity id="4CBD9B3E7B18FFFFBCD2FB63FE38FAF7" box="[418,491,1268,1295]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="391" unit="m" value="150.0">150 m</quantity>
,77–19.Y.1.994, P. Schütz” (
<specimenCount id="9D43FD527B18FFFFBE56FB62FCA1FAF6" box="[806,882,1268,1295]" pageId="2" pageNumber="391" type="male">1 male</specimenCount>
, AOC). The presence of this species in Mindanao and
<collectingRegion id="4981F8397B18FFFFBDB6FA8EFED5FACB" box="[198,262,1305,1331]" country="Philippines" name="Southern Leyte" pageId="2" pageNumber="391">Leyte</collectingRegion>
remains questionable. Most likely, that quotation from Mindanao correspond to the newly described species.
</paragraph>
</subSubSection>
</treatment>
</document>
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