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<document id="BC9C65EF970BAB08B9A7B193BA2C1855" ID-CLB-Dataset="31388" ID-DOI="10.11646/zootaxa.4083.4.7" ID-GBIF-Dataset="f2c3d1e3-2e79-4957-9cd2-e6c322bf1654" ID-ISSN="1175-5326" ID-Zenodo-Dep="1053923" ID-ZooBank="1B3E9DB6-7E5F-4AF7-82DF-21E71E2D8349" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510926575616" checkinUser="plazi" docAuthor="Pati, S. K., Thackeray, T. &amp; Khaire, A." docDate="2016" docId="03EC87B9FFD5FFEAFF2F6585FD34D74C" docLanguage="en" docName="zootaxa.4083.4.7.pdf" docOrigin="Zootaxa 4083 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Gubernatoriana alcocki Pati &amp; Thackeray &amp; Khaire 2016, n. sp." docType="treatment" docUuid="F68F84D2-E293-4F63-8300-8641CDBF65EF" docUuidSource="ZooBank" docVersion="8" lastPageNumber="580" masterDocId="FFD5FFC1FFDCFFE1FFB86558FFD3D144" masterDocTitle="Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India" masterLastPageNumber="586" masterPageNumber="569" pageNumber="578" updateTime="1698500881418" updateUser="ExternalLinkService">
<mods:mods id="AD88A90110E89CA8F7DFAB05E89ACF81" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="0A28A6C345A94D9C275818671343AAE6">
<mods:title id="A91A4758B97065CD551096020049C4DE">Five new species of freshwater crabs of the genera Ghatiana Pati &amp; Sharma, 2014, and Gubernatoriana Bott, 1970 (Crustacea, Decapoda, Brachyura: Gecarcinucidae Rathbun, 1904) from the Western Ghats, India</mods:title>
</mods:titleInfo>
<mods:name id="904EC661DFF604351637F3CA63C36B83" type="personal">
<mods:role id="FCD0E3A468C433637337D2729F36F19A">
<mods:roleTerm id="AA064AE3864C443F90B207D6E5B0D363">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="912FAA98ABE4746FB3FFDA5DC64A39EC">Pati, S. K.</mods:namePart>
</mods:name>
<mods:name id="CD68836403D1C22CA7E49964F7C3E7BA" type="personal">
<mods:role id="EB9254670BE5935F95AF16F4016A61CE">
<mods:roleTerm id="B0BB6AD55C69E2441DDE0D4006A9B860">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4D11456C9A49F606DAF388B38BBEAAE5">Thackeray, T.</mods:namePart>
</mods:name>
<mods:name id="04692EB7FABE49FF803005888397032B" type="personal">
<mods:role id="90D76B338BF22A9575560E3657CDEBA0">
<mods:roleTerm id="E6FA1ABEAB57BAE3279CCE2DE530B51B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="8EEBB6F1AFA68A9D47773AA8986FF8A7">Khaire, A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="77F3EA4DD1C99FE8D1FE2995EFBE4243">text</mods:typeOfResource>
<mods:relatedItem id="08745652CD23DD1113E6ECD7A7917981" type="host">
<mods:titleInfo id="2EB3C0E684F5F659CB6FCE62C1C50D78">
<mods:title id="62670BB739CFC04C2DC317F267D18186">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DA3C4A6C7F5F7B43DB95276361E158C1">
<mods:date id="B3B146D12FF6E521C0086EE67DE58619">2016</mods:date>
<mods:detail id="6EFAC45A109C465D6C29269ECB007793" type="pubDate">
<mods:number id="EBD4DEA4B59D63E6C088E35DFEE18B1A">2016-02-23</mods:number>
</mods:detail>
<mods:detail id="0774F0D27CCA3B7D9DB23C89B3ED3BC5" type="volume">
<mods:number id="16F38DE91FECB48AE3A42969704A9EA4">4083</mods:number>
</mods:detail>
<mods:detail id="BCF534917724FB8E3F72C45F8F2CD654" type="issue">
<mods:number id="045792D8802812AD25B5B0CA36FCFD10">4</mods:number>
</mods:detail>
<mods:extent id="3D2D60D3A4A83E06DF8A769D802FA4E2" unit="page">
<mods:start id="D4651357874356BABF27C4DED1ED7638">569</mods:start>
<mods:end id="41C1EA0A53D2BBE86852B2A242DEA4FF">586</mods:end>
</mods:extent>
</mods:part>
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<mods:classification id="330D268AAD018D069848C4337DB9ED00">journal article</mods:classification>
<mods:identifier id="4ECC8C83D6681543FFDCB074426E8631" type="CLB-Dataset">31388</mods:identifier>
<mods:identifier id="43A5A4C8A7AED3F691CFA0BC1E070A21" type="DOI">10.11646/zootaxa.4083.4.7</mods:identifier>
<mods:identifier id="5D609FEE2A5C5A371EBCEBA75C9493C5" type="GBIF-Dataset">f2c3d1e3-2e79-4957-9cd2-e6c322bf1654</mods:identifier>
<mods:identifier id="833E1EF1816C4A25371219AB89C47B3D" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="56EC9ACCDEC2208012F358680E03EF61" type="Zenodo-Dep">1053923</mods:identifier>
<mods:identifier id="266B9E32D8111590AEF55FF3972E572F" type="ZooBank">1B3E9DB6-7E5F-4AF7-82DF-21E71E2D8349</mods:identifier>
</mods:mods>
<treatment id="03EC87B9FFD5FFEAFF2F6585FD34D74C" ID-DOI="http://doi.org/10.5281/zenodo.6069952" ID-GBIF-Taxon="137453398" ID-Zenodo-Dep="6069952" LSID="urn:lsid:zoobank.org:act:F68F84D2-E293-4F63-8300-8641CDBF65EF" httpUri="http://treatment.plazi.org/id/03EC87B9FFD5FFEAFF2F6585FD34D74C" lastPageId="11" lastPageNumber="580" pageId="9" pageNumber="578">
<subSubSection id="C35F6524FFD5FFE8FF2F6585FEC2D05C" pageId="9" pageNumber="578" type="nomenclature">
<paragraph id="8BFA36AFFFD5FFE8FF2F6585FDE8D1B3" blockId="9.[151,571,221,280]" box="[151,571,221,247]" pageId="9" pageNumber="578">
<heading id="D0B281C3FFD5FFE8FF2F6585FDE8D1B3" bold="true" box="[151,571,221,247]" fontSize="11" level="1" pageId="9" pageNumber="578" reason="1">
<emphasis id="B931EABDFFD5FFE8FF2F6585FDE8D1B3" bold="true" box="[151,571,221,247]" pageId="9" pageNumber="578">
<taxonomicName id="4C454D2CFFD5FFE8FF2F6585FE21D1B3" ID-CoL="84FVK" authority="Pati" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[151,498,221,247]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD5FFE8FF2F6585FE6BD1B3" bold="true" box="[151,440,221,247]" italics="true" pageId="9" pageNumber="578">Gubernatoriana alcocki</emphasis>
Pati
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD5FFE8FE416585FDE8D1B3" box="[505,571,221,247]" pageId="9" pageNumber="578" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFA36AFFFD5FFE8FF2F65A7FEC2D05C" blockId="9.[151,571,221,280]" box="[151,273,255,280]" pageId="9" pageNumber="578">
(
<figureCitation id="137E2A2AFFD5FFE8FF2765A7FF3DD053" box="[159,238,255,280]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="9" pageNumber="578">Figs. 7</figureCitation>
,
<figureCitation id="137E2A2AFFD5FFE8FF426458FEDBD05C" box="[250,264,256,280]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="9" pageNumber="578">8</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD5FFE8FF2F641FFBDCD0B4" pageId="9" pageNumber="578" type="materials_examined">
<paragraph id="8BFA36AFFFD5FFE8FF2F641FFBDCD0B4" blockId="9.[151,1437,327,893]" pageId="9" pageNumber="578">
<emphasis id="B931EABDFFD5FFE8FF2F641FFE95D024" bold="true" box="[151,326,327,352]" pageId="9" pageNumber="578">
<typeStatus id="54FE880DFFD5FFE8FF2F641FFF01D024" box="[151,210,327,352]" pageId="9" pageNumber="578">Type</typeStatus>
material.
</emphasis>
<collectingCountry id="F352763FFFD5FFE8FEF76410FE71D024" box="[335,418,328,352]" name="India" pageId="9" pageNumber="578">INDIA</collectingCountry>
: adult male,
<typeStatus id="54FE880DFFD5FFE8FD8E641FFD4BD024" box="[566,664,327,352]" pageId="9" pageNumber="578" type="holotype">holotype</typeStatus>
(cw
<quantity id="4CBD9B4AFFD5FFE8FD6B6410FCE8D01B" box="[723,827,328,352]" metricMagnitude="-2" metricUnit="m" metricValue="1.22" pageId="9" pageNumber="578" unit="mm" value="12.2">12.2 mm</quantity>
, cl
<quantity id="4CBD9B4AFFD5FFE8FCDD641FFC1ED01B" box="[869,973,327,352]" metricMagnitude="-3" metricUnit="m" metricValue="9.66" pageId="9" pageNumber="578" unit="mm" value="9.66">9.66 mm</quantity>
, ch
<quantity id="4CBD9B4AFFD5FFE8FC45641FFB84D01B" box="[1021,1111,327,352]" metricMagnitude="-3" metricUnit="m" metricValue="6.1" pageId="9" pageNumber="578" unit="mm" value="6.1">6.1 mm</quantity>
, fw
<quantity id="4CBD9B4AFFD5FFE8FB33641FFB34D024" box="[1163,1255,327,352]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" pageId="9" pageNumber="578" unit="mm" value="4.6">4.6 mm</quantity>
), Vankusawade, near Ghanbi Vatole, Patan taluk, Satara district,
<collectingRegion id="4981F84DFFD5FFE8FD0F6434FC90D0C1" box="[695,835,364,389]" country="India" name="Maharashtra" pageId="9" pageNumber="578">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE715068FFD5FFE8FCEA6434FC1BD0C0" box="[850,968,364,388]" direction="north" orientation="latitude" pageId="9" pageNumber="578" precision="55" value="17.449">17.449° N</geoCoordinate>
and
<geoCoordinate id="EE715068FFD5FFE8FBB86434FBA7D0C0" box="[1024,1140,364,388]" direction="east" orientation="longitude" pageId="9" pageNumber="578" precision="55" value="73.829">73.829° E</geoCoordinate>
), altitude
<quantity id="4CBD9B4AFFD5FFE8FB5D6434FAEED0C0" box="[1253,1341,364,388]" metricMagnitude="3" metricUnit="m" metricValue="1.082" pageId="9" pageNumber="578" unit="m" value="1082.0">1082 m</quantity>
,
<date id="FFFB106FFFD5FFE8FAF26434FF01D0EC" pageId="9" pageNumber="578" value="2014-07-22">22 July 2014</date>
, coll. S.S. Kamble &amp; Party (ZSI, WRC-C.1513);
<typeStatus id="54FE880DFFD5FFE8FCB964C8FCBCD0EC" box="[769,879,400,424]" pageId="9" pageNumber="578" type="paratype">paratypes</typeStatus>
, two males (cw
<quantity id="4CBD9B4AFFD5FFE8FB9F64D7FB3AD0E3" box="[1063,1257,399,424]" metricMagnitude="-2" metricUnit="m" metricValue="1.424" metricValueMax="1.476" metricValueMin="1.372" pageId="9" pageNumber="578" unit="mm" value="14.24" valueMax="14.76" valueMin="13.72">13.72–14.76 mm</quantity>
, cl
<quantity id="4CBD9B4AFFD5FFE8FAA964C8FF16D088" metricMagnitude="-2" metricUnit="m" metricValue="1.135" metricValueMax="1.178" metricValueMin="1.092" pageId="9" pageNumber="578" unit="mm" value="11.35" valueMax="11.78" valueMin="10.92">10.92–11.78 mm</quantity>
, ch
<quantity id="4CBD9B4AFFD5FFE8FF4964ECFE45D088" box="[241,406,436,461]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" metricValueMax="7.76" metricValueMin="7.24" pageId="9" pageNumber="578" unit="mm" value="7.5" valueMax="7.76" valueMin="7.24">7.24–7.76 mm</quantity>
, fw
<quantity id="4CBD9B4AFFD5FFE8FE7E64ECFDB9D088" box="[454,618,436,461]" metricMagnitude="-3" metricUnit="m" metricValue="5.49" metricValueMax="5.68" metricValueMin="5.3" pageId="9" pageNumber="578" unit="mm" value="5.49" valueMax="5.68" valueMin="5.3">5.30–5.68 mm</quantity>
) and two females (cw
<quantity id="4CBD9B4AFFD5FFE8FCDF64ECFBF4D088" box="[871,1063,436,460]" metricMagnitude="-2" metricUnit="m" metricValue="1.251" metricValueMax="1.372" metricValueMin="1.13" pageId="9" pageNumber="578" unit="mm" value="12.51" valueMax="13.72" valueMin="11.3">11.30–13.72 mm</quantity>
, cl
<quantity id="4CBD9B4AFFD5FFE8FBF564ECFAD3D088" box="[1101,1280,436,460]" metricMagnitude="-3" metricUnit="m" metricValue="9.93" metricValueMax="10.92" metricValueMin="8.94" pageId="9" pageNumber="578" unit="mm" value="9.93" valueMax="10.92" valueMin="8.94">8.94–10.92 mm</quantity>
, ch
<quantity id="4CBD9B4AFFD5FFE8FA9464ECFF16D0AB" metricMagnitude="-3" metricUnit="m" metricValue="6.61" metricValueMax="7.24" metricValueMin="5.98" pageId="9" pageNumber="578" unit="mm" value="6.61" valueMax="7.24" valueMin="5.98">5.98–7.24 mm</quantity>
, fw
<quantity id="4CBD9B4AFFD5FFE8FF4D648FFE4FD0B4" box="[245,412,471,496]" metricMagnitude="-3" metricUnit="m" metricValue="4.93" metricValueMax="5.3" metricValueMin="4.56" pageId="9" pageNumber="578" unit="mm" value="4.93" valueMax="5.3" valueMin="4.56">4.56–5.30 mm</quantity>
), same collection data as
<typeStatus id="54FE880DFFD5FFE8FD01648FFCC8D0B4" box="[697,795,471,496]" pageId="9" pageNumber="578" type="holotype">holotype</typeStatus>
(ZSI, WRC-C.1514).
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD5FFE8FF7F64A4FBC4D238" pageId="9" pageNumber="578" type="diagnosis">
<paragraph id="8BFA36AFFFD5FFE8FF7F64A4FBC4D238" blockId="9.[151,1437,327,893]" pageId="9" pageNumber="578">
<emphasis id="B931EABDFFD5FFE8FF7F64A4FE92D351" bold="true" box="[199,321,508,533]" pageId="9" pageNumber="578">Diagnosis.</emphasis>
Carapace quadrate (cw/cl = 1.3); dorsal surface smooth, olive-brown coloured; anterolateral margin short, smooth, cristiform; posterolateral margin slightly convergent, with fine, oblique striations; front strongly deflexed, narrow (fw/cw = 0.4); epigastric cristae faintly visible; postorbital cristae indistinct; exorbital angle triangular, distinct, with long outer margin, ca. 2 times length of inner margin; epibranchial tooth visible as weak notch; frontal median triangle incomplete; epistomal median lobe broadly triangular with acute apex, lacking median tooth (
<figureCitation id="137E2A2AFFD5FFE8FE8067F7FE6FD38C" box="[312,444,687,712]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="9" pageNumber="578">Figs. 7A, B</figureCitation>
). Finger tips of chelipeds pointed (
<figureCitation id="137E2A2AFFD5FFE8FCF867F7FC49D38C" box="[832,922,687,712]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="9" pageNumber="578">Fig. 7D</figureCitation>
). Suture between male thoracic sternites s2/s3 indistinct, suture between sternites s3/s4 missing except for two short lateral grooves (
<figureCitation id="137E2A2AFFD5FFE8FBE7678CFB32D3A8" box="[1119,1249,724,749]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="9" pageNumber="578">Figs. 7C, F</figureCitation>
). Male abdomen narrowly T-shaped; sixth abdominal somite slightly broader than long, nearly equal to telson length; telson short; male sternoabdominal cavity deep, long, extending beyond level of bases of third maxillipeds (
<figureCitation id="137E2A2AFFD5FFE8FB626644FAB6D270" box="[1242,1381,796,821]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="9" pageNumber="578">Figs. 7C, G</figureCitation>
). G1 with slender, cone-shaped, long terminal article, ca. 0.4 times length of subterminal segment, tip slightly curving inwards (
<figureCitation id="137E2A2AFFD5FFE8FF47663CFE55D238" box="[255,390,868,893]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="9" pageNumber="578">Figs. 8A–C</figureCitation>
). G2 with very short or vestigial terminal article (
<figureCitation id="137E2A2AFFD5FFE8FC08663CFBD9D238" box="[944,1034,868,893]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="9" pageNumber="578">Fig. 8D</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3A6627FFD5FFE8FF2F6205FCE4D6F5" httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="9" pageNumber="578" targetBox="[340,1247,981,1864]" targetPageId="9">
<paragraph id="8BFA36AFFFD5FFE8FF2F6205FCE4D6F5" blockId="9.[151,1436,1885,1969]" pageId="9" pageNumber="578">
<emphasis id="B931EABDFFD5FFE8FF2F6205FEC7D637" bold="true" box="[151,276,1885,1907]" pageId="9" pageNumber="578">FIGURE 7.</emphasis>
<taxonomicName id="4C454D2CFFD5FFE8FEA36205FDC3D637" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[283,528,1885,1907]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="578" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD5FFE8FEA36205FDC3D637" box="[283,528,1885,1907]" italics="true" pageId="9" pageNumber="578">Gubernatoriana alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD5FFE8FDA06206FD81D637" box="[536,594,1886,1907]" pageId="9" pageNumber="578" rank="species">
<emphasis id="B931EABDFFD5FFE8FDA06206FD81D637" bold="true" box="[536,594,1886,1907]" pageId="9" pageNumber="578">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (ZSI, WRC—C.1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D–G), 1 mm (H).
</paragraph>
</caption>
<caption id="DF3A6627FFD6FFEBFF2F667EFBC1D21F" httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="10" pageNumber="579" targetBox="[360,1240,237,754]" targetPageId="10">
<paragraph id="8BFA36AFFFD6FFEBFF2F667EFBC1D21F" blockId="10.[151,1435,806,859]" pageId="10" pageNumber="579">
<emphasis id="B931EABDFFD6FFEBFF2F667EFEC0D278" bold="true" box="[151,275,806,828]" pageId="10" pageNumber="579">FIGURE 8.</emphasis>
<taxonomicName id="4C454D2CFFD6FFEBFEA2667EFDDCD278" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[282,527,806,828]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="579" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD6FFEBFEA2667EFDDCD278" box="[282,527,806,828]" italics="true" pageId="10" pageNumber="579">Gubernatoriana alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD6FFEBFDAE667FFD83D278" box="[534,592,807,828]" pageId="10" pageNumber="579" rank="species">
<emphasis id="B931EABDFFD6FFEBFDAE667FFD83D278" bold="true" box="[534,592,807,828]" pageId="10" pageNumber="579">n. sp.</emphasis>
</taxonomicNameLabel>
, holotype male (ZSI, WRC—C.1513). A, dorsal view of right G1; B, ventral view of right G1; C, dorsal view of right G1 terminal article D, right G2. Scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="C35F6524FFD6FFEBFF2F66DAFE22D6DE" pageId="10" pageNumber="579" type="description">
<paragraph id="8BFA36AFFFD6FFEBFF2F66DAFA90D5C0" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
<emphasis id="B931EABDFFD6FFEBFF2F66DAFE69D2DF" bold="true" box="[151,442,898,923]" pageId="10" pageNumber="579">
Description of
<typeStatus id="54FE880DFFD6FFEBFEF466DAFE66D2DF" box="[332,437,898,923]" pageId="10" pageNumber="579" type="holotype">holotype</typeStatus>
.
</emphasis>
Carapace quadrate (cw/cl = 1.3); dorsal surface smooth; anterolateral carapace gently inflated in frontal view; anterolateral margin short, smooth, cristiform; posterolateral margin slightly convergent, with fine, oblique striations; front strongly deflexed, square cut, nearly flat, frontal margin slightly concave, narrow (fw/cw = 0.4); epigastric cristae faintly visible, smooth; postorbital cristae indistinct; exorbital angle triangular, distinct, with long outer margin, ca. 2 times length of inner margin; epibranchial tooth visible as weak notch; postorbital region shallow; branchial regions gently inflated; subhepatic region slightly rugose; cervical grooves indistinct; mesogastric groove long, with bifurcation posteriorly; H-groove visible, shallow; frontal median triangle incomplete; epistomal median lobe broadly triangular with acute apex, lacking median tooth (
<figureCitation id="137E2A2AFFD6FFEBFB096134FAE6D5C0" box="[1201,1333,1132,1157]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">Figs. 7A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F61D5FBFED472" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
Chelipeds unequal; right chela larger, smooth, fingers with two or three larger teeth, distinct gape when finger tips in contact (
<figureCitation id="137E2A2AFFD6FFEBFEFD61E9FE4CD58E" box="[325,415,1201,1226]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">Fig. 7D</figureCitation>
); length of dactylus greater than length of upper margin of palm; carpal spine small, blunt; outer surface of merus rugose. Ambulatory legs (p2–p5) long with fine, long tuft of brownish bristles on anterior margin of merus, carpus, propodus, dactylus, posterior margin of propodus, dactylus; dactylus (p2–p5) slightly longer than propodus; longest propodus (p3) two times as long as broad (
<figureCitation id="137E2A2AFFD6FFEBFC716045FBF2D472" box="[969,1057,1309,1334]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">Fig. 7E</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F601AFDCAD43A" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
Suture between male thoracic sternites s2/s3 indistinct, suture between sternites s3/s4 missing except for two short lateral grooves (
<figureCitation id="137E2A2AFFD6FFEBFE33603DFDDFD43A" box="[395,524,1381,1406]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">Figs. 7C, F</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F60D1FBD4D74A" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
Male abdomen narrowly T-shaped; fifth abdominal somite distinctly broader than long, with concave lateral margins; sixth abdominal somite slightly broader than long, nearly equal in length to telson, with slightly concave lateral margins; telson short, broadly triangular with almost straight outer margins; male sternoabdominal cavity deep, long, extending beyond level of bases of third maxillipeds (
<figureCitation id="137E2A2AFFD6FFEBFCCC60ADFC29D74A" box="[884,1018,1525,1550]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">Figs. 7C, G</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F6341FA41D73E" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
Mandibular palp with two joints, terminal joint bilobed; exopods of first, second maxillipeds with long flagellum; exopod of third maxilliped lacking flagellum, longer than ischium; ventral sulcus on ischium distinct, subparallel to inner margin; merus with rounded anterior external angle, depression towards inner sides (
<figureCitation id="137E2A2AFFD6FFEBFA936339FA56D73E" box="[1323,1413,1633,1658]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">Fig. 7H</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F63DEFAAFD64E" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
G1 with slender, cone-shaped, long terminal article, ca. 0.4 times length of subterminal segment, tip broad, slightly curving inwards, joint between terminal article, subterminal segment clear; subterminal segment broader than terminal article, basal half broader than distal half (
<figureCitation id="137E2A2AFFD6FFEBFCA46395FC76D7A2" box="[796,933,1741,1766]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="10" pageNumber="579">Figs. 8A–C</figureCitation>
). G2 conspicuously short with very short or vestigial terminal article; proximal half portion of basal segment much stouter than distal half portion (
<figureCitation id="137E2A2AFFD6FFEBFAAE63A9FAA3D64E" box="[1302,1392,1777,1802]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="10" pageNumber="579">Fig. 8D</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F624DFB67D66A" blockId="10.[151,1437,898,2018]" box="[199,1204,1813,1838]" pageId="10" pageNumber="579">
<emphasis id="B931EABDFFD6FFEBFF7F624DFE80D66A" bold="true" box="[199,339,1813,1838]" pageId="10" pageNumber="579">Live colour.</emphasis>
Live crabs are olive-brown but their hands are orange-brown (
<figureCitation id="137E2A2AFFD6FFEBFBAB624DFBA6D66A" box="[1043,1141,1813,1838]" captionStart="FIGURE 2" captionStartId="2.[151,250,1654,1676]" captionTargetBox="[340,1247,758,1632]" captionTargetId="figure@2.[340,1247,758,1632]" captionTargetPageId="2" captionText="FIGURE 2. Live crabs of new species of Ghatiana and Gubernatoriana. A, Ghatiana aurantiaca n. sp. from Amboli; B, Ghatiana splendida n. sp. from Chaukul; C, Gubernatoriana alcocki n. sp. from Vankusawade; D, Gubernatoriana thackerayi n. sp. from Raghuvir Ghat; C, Gubernatoriana waghi n. sp. from Malshej Ghat. Photo credits: A, B, Arjun Kamdar; C, B. V. Jadhav; D, Shailesh Bhosale; E, Rachit Shah." httpUri="https://zenodo.org/record/1053927/files/figure.png" pageId="10" pageNumber="579">Figs. 2C</figureCitation>
,
<figureCitation id="137E2A2AFFD6FFEBFB39624EFB75D66A" box="[1153,1190,1814,1838]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="10" pageNumber="579">7A</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD6FFEBFF7F6261FE22D6DE" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
<emphasis id="B931EABDFFD6FFEBFF7F6261FE95D616" bold="true" box="[199,326,1849,1874]" pageId="10" pageNumber="579">
<typeStatus id="54FE880DFFD6FFEBFF7F6261FE92D616" box="[199,321,1849,1874]" pageId="10" pageNumber="579" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<typeStatus id="54FE880DFFD6FFEBFEF66262FE68D616" box="[334,443,1850,1874]" pageId="10" pageNumber="579" type="paratype">Paratypes</typeStatus>
are identical to the
<typeStatus id="54FE880DFFD6FFEBFD276261FCD2D616" box="[671,769,1849,1874]" pageId="10" pageNumber="579" type="holotype">holotype</typeStatus>
in all morphological features barring females (ZSI, WRC- C.1514), in which chelipeds are almost equal, their fingers have 12 to 14 small, blunt teeth, and the gape between fingers is comparatively small.
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD6FFEBFF7F62FDFE82D6A6" pageId="10" pageNumber="579" type="etymology">
<paragraph id="8BFA36AFFFD6FFEBFF7F62FDFE82D6A6" blockId="10.[151,1437,898,2018]" pageId="10" pageNumber="579">
<emphasis id="B931EABDFFD6FFEBFF7F62FDFE9DD6FA" bold="true" box="[199,334,1957,1982]" pageId="10" pageNumber="579">Etymology.</emphasis>
Species is named in the honour of British physician, naturalist, and carcinologist Major Dr. Alfred William Alcock.
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD7FFEAFF7F65CFFE32D190" pageId="11" pageNumber="580" type="materials_examined">
<paragraph id="8BFA36AFFFD7FFEAFF7F65CFFE32D190" blockId="11.[151,1437,151,1544]" pageId="11" pageNumber="580">
<emphasis id="B931EABDFFD7FFEAFF7F65CFFEBAD1F4" bold="true" box="[199,361,151,176]" pageId="11" pageNumber="580">
<typeStatus id="54FE880DFFD7FFEAFF7F65CFFED1D1F4" box="[199,258,151,176]" pageId="11" pageNumber="580">Type</typeStatus>
locality.
</emphasis>
Vankusawade, near Ghanbi Vatole, Patan taluk, Satara district,
<collectingRegion id="4981F84DFFD7FFEAFBEA65CFFB0DD1F4" box="[1106,1246,151,176]" country="India" name="Maharashtra" pageId="11" pageNumber="580">Maharashtra</collectingRegion>
(
<geoCoordinate id="EE715068FFD7FFEAFB4965C0FABAD1EB" box="[1265,1385,152,176]" direction="north" orientation="latitude" pageId="11" pageNumber="580" precision="55" value="17.449">17.449° N</geoCoordinate>
and
<geoCoordinate id="EE715068FFD7FFEAFF2F65E4FEDBD190" box="[151,264,188,212]" direction="east" orientation="longitude" pageId="11" pageNumber="580" precision="55" value="73.829">73.829° E</geoCoordinate>
) (altitude
<quantity id="4CBD9B4AFFD7FFEAFEC465E4FE06D190" box="[380,469,188,212]" metricMagnitude="3" metricUnit="m" metricValue="1.082" pageId="11" pageNumber="580" unit="m" value="1082.0">1082 m</quantity>
).
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD7FFEAFF7F6587FE63D28C" pageId="11" pageNumber="580" type="discussion">
<paragraph id="8BFA36AFFFD7FFEAFF7F6587FBC4D020" blockId="11.[151,1437,151,1544]" pageId="11" pageNumber="580">
<emphasis id="B931EABDFFD7FFEAFF7F6587FEE8D1BC" bold="true" box="[199,315,223,248]" pageId="11" pageNumber="580">Remarks.</emphasis>
<taxonomicName id="4C454D2CFFD7FFEAFEFB65B8FD80D1BC" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[323,595,224,248]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFEFB65B8FD80D1BC" box="[323,595,224,248]" italics="true" pageId="11" pageNumber="580">Gubernatoriana alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFDE46587FD4FD1BC" box="[604,668,223,248]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFDE46587FD4FD1BC" bold="true" box="[604,668,223,248]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
is similar to its congeners except
<taxonomicName id="4C454D2CFFD7FFEAFB9D65B8FB11D1BC" authorityName="Klaus, Fernandez &amp; Yeo" authorityYear="2014" box="[1061,1218,224,248]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B931EABDFFD7FFEAFB9D65B8FB11D1BC" box="[1061,1218,224,248]" italics="true" pageId="11" pageNumber="580">G. basalticola</emphasis>
</taxonomicName>
by the presence of following suite of morphological characters: carapace quadrate (cw/cl = 1.1–1.3), posterolateral margin nearly straight to slightly convergent, frontal margin narrow (fw/cw = 0.3–0.4), epistomal median lobe with an acute apex, telson short, and G2 terminal article very short or vestigial (
<figureCitation id="137E2A2AFFD7FFEAFC8D6414FC0AD020" box="[821,985,332,357]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="11" pageNumber="580">Figs. 7A–C, G</figureCitation>
,
<figureCitation id="137E2A2AFFD7FFEAFC5C6414FBD9D020" box="[996,1034,332,356]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="11" pageNumber="580">8D</figureCitation>
).
</paragraph>
<paragraph id="8BFA36AFFFD7FFEAFF7F6428FE63D28C" blockId="11.[151,1437,151,1544]" pageId="11" pageNumber="580">
A diagnostic feature of
<taxonomicName id="4C454D2CFFD7FFEAFE606428FD99D0CC" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[472,586,368,392]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFE606428FD99D0CC" box="[472,586,368,392]" italics="true" pageId="11" pageNumber="580">G. alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFDEC6437FD47D0CC" box="[596,660,367,392]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFDEC6437FD47D0CC" bold="true" box="[596,660,367,392]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
is the inwardly curved tip of the G1 terminal article (
<figureCitation id="137E2A2AFFD7FFEAFABD6437FA5CD0CC" box="[1285,1423,367,392]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="11" pageNumber="580">Figs. 8A, C</figureCitation>
). Otherwise,
<taxonomicName id="4C454D2CFFD7FFEAFEA064CCFE79D0E8" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[280,426,404,428]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFEA064CCFE79D0E8" box="[280,426,404,428]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C454D2CFFD7FFEAFE5964CCFD42D0E8" authorityName="Alcock" authorityYear="1909" box="[481,657,404,428]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="gubernatoris">
<emphasis id="B931EABDFFD7FFEAFE5964CCFD42D0E8" box="[481,657,404,428]" italics="true" pageId="11" pageNumber="580">G. gubernatoris</emphasis>
</taxonomicName>
are more similar to
<taxonomicName id="4C454D2CFFD7FFEAFCCD64CCFC36D0E8" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[885,997,404,428]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFCCD64CCFC36D0E8" box="[885,997,404,428]" italics="true" pageId="11" pageNumber="580">G. alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFC5464CCFBF8D0E9" box="[1004,1067,404,429]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFC5464CCFBF8D0E9" bold="true" box="[1004,1067,404,429]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
among congeners due their long G1 terminal article, ca. 0.4–0.5 times the length of the subterminal segment (see
<bibRefCitation id="EFD44B5EFFD7FFEAFB9E64EFFACED094" author="Pati, S. K. &amp; Sharma, R. M." box="[1062,1309,439,464]" journalOrPublisher="India. Journal of Natural History" pageId="11" pageNumber="580" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014" yearSuffix="a">Pati &amp; Sharma 2014a</bibRefCitation>
: fig. 7E for
<taxonomicName id="4C454D2CFFD7FFEAFF2F6484FEF8D0B0" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[151,299,476,500]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFF2F6484FEF8D0B0" box="[151,299,476,500]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFD44B5EFFD7FFEAFE856484FDEFD0B1" author="Pati, S. K. &amp; Sharma, R. M." box="[317,572,476,501]" journalOrPublisher="Occasional Paper, No. 363" pageId="11" pageNumber="580" pagination="1 - 44" refString="Pati, S. K. &amp; Sharma, R. M. (2014 b) Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India, Occasional Paper, No. 363, 1 - 44. [published by the Director, Zoological Survey of India, Kolkata]" title="Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India" type="book chapter" year="2014" yearSuffix="b">Pati &amp; Sharma 2014b</bibRefCitation>
: pl. VII, fig. 7D for
<taxonomicName id="4C454D2CFFD7FFEAFC886484FC31D0B0" authorityName="Alcock" authorityYear="1909" box="[816,994,476,500]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="gubernatoris">
<emphasis id="B931EABDFFD7FFEAFC886484FC31D0B0" box="[816,994,476,500]" italics="true" pageId="11" pageNumber="580">G. gubernatoris</emphasis>
</taxonomicName>
). The terminal article of the G1 of
<taxonomicName id="4C454D2CFFD7FFEAFA3C6484FEC0D35C" authorityName="Klaus, Fernandez &amp; Yeo" authorityYear="2014" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B931EABDFFD7FFEAFA3C6484FEC0D35C" italics="true" pageId="11" pageNumber="580">G. basalticola</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D2CFFD7FFEAFE9C6758FE79D35C" authorityName="Alcock" authorityYear="1909" box="[292,426,512,536]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B931EABDFFD7FFEAFE9C6758FE79D35C" box="[292,426,512,536]" italics="true" pageId="11" pageNumber="580">G. pilosipes</emphasis>
</taxonomicName>
,
<taxonomicName id="4C454D2CFFD7FFEAFE036758FD80D35C" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[443,595,512,536]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="thackerayi" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFE036758FD80D35C" box="[443,595,512,536]" italics="true" pageId="11" pageNumber="580">G. thackerayi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFDE564A7FD4CD35C" box="[605,671,511,536]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFDE564A7FD4CD35C" bold="true" box="[605,671,511,536]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C454D2CFFD7FFEAFD646758FC91D35C" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[732,834,512,536]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="waghi" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFD646758FC91D35C" box="[732,834,512,536]" italics="true" pageId="11" pageNumber="580">G. waghi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFCF464A7FC5DD35C" box="[844,910,511,536]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFCF464A7FC5DD35C" bold="true" box="[844,910,511,536]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
is short, ca. 0.3–0.35 times the length of the subterminal segment (
<figureCitation id="137E2A2AFFD7FFEAFE2E677CFE2ED378" box="[406,509,548,573]" captionStart="FIGURE 10" captionStartId="13.[151,250,809,831]" captionTargetBox="[350,1242,224,761]" captionTargetId="figure@13.[340,1247,193,788]" captionTargetPageId="13" captionText="FIGURE 10. Gubernatoriana thackerayi n. sp., holotype male (ZSI, WRC—C. 1515). A, dorsal view of left G 1; B, ventral view of left G 1; C, dorsal view of left G 1 terminal article; D, left G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053943/files/figure.png" pageId="11" pageNumber="580">Fig. 10A</figureCitation>
for
<taxonomicName id="4C454D2CFFD7FFEAFD8B677CFD18D378" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[563,715,548,572]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="thackerayi">
<emphasis id="B931EABDFFD7FFEAFD8B677CFD18D378" box="[563,715,548,572]" italics="true" pageId="11" pageNumber="580">G. thackerayi</emphasis>
</taxonomicName>
,
<figureCitation id="137E2A2AFFD7FFEAFD65677CFC90D378" box="[733,835,548,573]" captionStart="FIGURE 12" captionStartId="15.[151,250,768,790]" captionTargetBox="[355,1227,268,683]" captionTargetId="figure@15.[340,1247,193,787]" captionTargetPageId="15" captionText="FIGURE 12. Gubernatoriana waghi n. sp., holotype male (ZSI, WRC—C. 1517). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article; D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053947/files/figure.png" pageId="11" pageNumber="580">Fig. 12A</figureCitation>
for
<taxonomicName id="4C454D2CFFD7FFEAFCC1677CFC0CD378" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[889,991,548,572]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="waghi">
<emphasis id="B931EABDFFD7FFEAFCC1677CFC0CD378" box="[889,991,548,572]" italics="true" pageId="11" pageNumber="580">G. waghi</emphasis>
</taxonomicName>
; see
<bibRefCitation id="EFD44B5EFFD7FFEAFB98677CFB3CD378" author="Klaus, S. &amp; Fernandez, K. &amp; Yeo, D. C. J." box="[1056,1263,548,573]" journalOrPublisher="Zoological Scripta" pageId="11" pageNumber="580" pagination="651 - 660" part="43" publicationUrl="http://dx.doi.org/10.1111/zsc.12078" refString="Klaus, S., Fernandez, K. &amp; Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoological Scripta, 43 (6), 651 - 660. http: // dx. doi. org / 10.1111 / zsc. 12078" title="Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae)" type="journal article" year="2014">
Klaus
<emphasis id="B931EABDFFD7FFEAFBD4677DFB7BD378" box="[1132,1192,548,572]" italics="true" pageId="11" pageNumber="580">et al.</emphasis>
2014
</bibRefCitation>
: fig. 3D for
<taxonomicName id="4C454D2CFFD7FFEAFA3C677CFEC0D324" authorityName="Klaus, Fernandez &amp; Yeo" authorityYear="2014" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B931EABDFFD7FFEAFA3C677CFEC0D324" italics="true" pageId="11" pageNumber="580">G. basalticola</emphasis>
</taxonomicName>
;
<bibRefCitation id="EFD44B5EFFD7FFEAFE9B671FFDC9D324" author="Pati, S. K. &amp; Sharma, R. M." box="[291,538,583,608]" journalOrPublisher="Occasional Paper, No. 363" pageId="11" pageNumber="580" pagination="1 - 44" refString="Pati, S. K. &amp; Sharma, R. M. (2014 b) Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India, Occasional Paper, No. 363, 1 - 44. [published by the Director, Zoological Survey of India, Kolkata]" title="Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India" type="book chapter" year="2014" yearSuffix="b">Pati &amp; Sharma 2014b</bibRefCitation>
: pl. VIII, fig. 8D for
<taxonomicName id="4C454D2CFFD7FFEAFCB16710FC5FD324" authorityName="Alcock" authorityYear="1909" box="[777,908,584,608]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="pilosipes">
<emphasis id="B931EABDFFD7FFEAFCB16710FC5FD324" box="[777,908,584,608]" italics="true" pageId="11" pageNumber="580">G. pilosipes</emphasis>
</taxonomicName>
). While the male sternoabdominal cavity of
<taxonomicName id="4C454D2CFFD7FFEAFA3C6710FF3BD3C0" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFA3C6710FF3BD3C0" italics="true" pageId="11" pageNumber="580">G. alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFF576734FEFDD3C1" box="[239,302,620,645]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFF576734FEFDD3C1" bold="true" box="[239,302,620,645]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
and
<taxonomicName id="4C454D2CFFD7FFEAFEDE6734FE2BD3C0" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[358,504,620,644]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFEDE6734FE2BD3C0" box="[358,504,620,644]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
is long, extending beyond the level of the third maxilliped bases (
<figureCitation id="137E2A2AFFD7FFEAFB596734FAEAD3C0" box="[1249,1337,620,645]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="11" pageNumber="580">Fig. 7C</figureCitation>
; see
<bibRefCitation id="EFD44B5EFFD7FFEAFAC86734FE88D3EC" author="Pati, S. K. &amp; Sharma, R. M." journalOrPublisher="India. Journal of Natural History" pageId="11" pageNumber="580" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014" yearSuffix="a">Pati &amp; Sharma 2014a</bibRefCitation>
: fig. 6C for
<taxonomicName id="4C454D2CFFD7FFEAFE5C67C8FDA4D3EC" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[484,631,656,680]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFE5C67C8FDA4D3EC" box="[484,631,656,680]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
), that of the
<taxonomicName id="4C454D2CFFD7FFEAFCB167C8FC69D3EC" authorityName="Alcock" authorityYear="1909" box="[777,954,656,680]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="gubernatoris">
<emphasis id="B931EABDFFD7FFEAFCB167C8FC69D3EC" box="[777,954,656,680]" italics="true" pageId="11" pageNumber="580">G. gubernatoris</emphasis>
</taxonomicName>
is short, extending beyond the level of the cheliped bases (see
<bibRefCitation id="EFD44B5EFFD7FFEAFECB67ECFDB4D389" author="Pati, S. K. &amp; Sharma, R. M." box="[371,615,692,717]" journalOrPublisher="Occasional Paper, No. 363" pageId="11" pageNumber="580" pagination="1 - 44" refString="Pati, S. K. &amp; Sharma, R. M. (2014 b) Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India, Occasional Paper, No. 363, 1 - 44. [published by the Director, Zoological Survey of India, Kolkata]" title="Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India" type="book chapter" year="2014" yearSuffix="b">Pati &amp; Sharma 2014b</bibRefCitation>
: pl. VII, fig. 7C). The distal portion of the G1 terminal article is distinctly broader than its proximal portion, and the subterminal segment appears slimmer in
<taxonomicName id="4C454D2CFFD7FFEAFB8E6780FB76D3B4" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[1078,1189,728,752]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFB8E6780FB76D3B4" box="[1078,1189,728,752]" italics="true" pageId="11" pageNumber="580">G. alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFB14678FFB39D3B4" box="[1196,1258,727,752]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFB14678FFB39D3B4" bold="true" box="[1196,1258,727,752]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
(
<figureCitation id="137E2A2AFFD7FFEAFB41678FFA82D3B4" box="[1273,1361,727,752]" captionStart="FIGURE 8" captionStartId="10.[151,250,806,828]" captionTargetBox="[360,1240,237,754]" captionTargetId="figure@10.[340,1247,193,785]" captionTargetPageId="10" captionText="FIGURE 8. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view of right G 1; B, ventral view of right G 1; C, dorsal view of right G 1 terminal article D, right G 2. Scale bar = 0.5 mm." httpUri="https://zenodo.org/record/1053939/files/figure.png" pageId="11" pageNumber="580">Fig. 8A</figureCitation>
) when compared to the distinctly narrow distal portion of the G1 terminal article, and stout, triangular appearance of the subterminal segment of
<taxonomicName id="4C454D2CFFD7FFEAFE1C6678FDE5D27C" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[420,566,800,824]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFE1C6678FDE5D27C" box="[420,566,800,824]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFD44B5EFFD7FFEAFDC96647FCB6D27C" author="Pati, S. K. &amp; Sharma, R. M." box="[625,869,799,824]" journalOrPublisher="India. Journal of Natural History" pageId="11" pageNumber="580" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014" yearSuffix="a">Pati &amp; Sharma 2014a</bibRefCitation>
: fig. 7E for
<taxonomicName id="4C454D2CFFD7FFEAFC526678FBA8D27C" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[1002,1147,800,824]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFC526678FBA8D27C" box="[1002,1147,800,824]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
). Furthermore,
<taxonomicName id="4C454D2CFFD7FFEAFA946678FA4FD27C" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[1324,1436,800,824]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFA946678FA4FD27C" box="[1324,1436,800,824]" italics="true" pageId="11" pageNumber="580">G. alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFF2F661CFF08D219" box="[151,219,836,861]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFF2F661CFF08D219" bold="true" box="[151,219,836,861]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
can be distinguished from
<taxonomicName id="4C454D2CFFD7FFEAFD90661CFD13D218" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[552,704,836,860]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="triangulus">
<emphasis id="B931EABDFFD7FFEAFD90661CFD13D218" box="[552,704,836,860]" italics="true" pageId="11" pageNumber="580">G. triangulus</emphasis>
</taxonomicName>
by the triangular, distinct exorbital angle (
<figureCitation id="137E2A2AFFD7FFEAFB7D661CFAF1D218" box="[1221,1314,836,861]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="11" pageNumber="580">Fig. 7A</figureCitation>
) [vs. low, indistinct exorbital angle (see
<bibRefCitation id="EFD44B5EFFD7FFEAFE5E663FFD09D2C4" author="Pati, S. K. &amp; Sharma, R. M." box="[486,730,871,896]" journalOrPublisher="India. Journal of Natural History" pageId="11" pageNumber="580" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014" yearSuffix="a">Pati &amp; Sharma 2014a</bibRefCitation>
: fig. 6A)] and from
<taxonomicName id="4C454D2CFFD7FFEAFC0F6630FBB5D2C4" authorityName="Alcock" authorityYear="1909" box="[951,1126,872,896]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="gubernatoris">
<emphasis id="B931EABDFFD7FFEAFC0F6630FBB5D2C4" box="[951,1126,872,896]" italics="true" pageId="11" pageNumber="580">G. gubernatoris</emphasis>
</taxonomicName>
by the long outer margin of the exorbital angle (
<figureCitation id="137E2A2AFFD7FFEAFEC166D4FE01D2E0" box="[377,466,908,933]" captionStart="FIGURE 7" captionStartId="9.[151,250,1885,1907]" captionTargetBox="[340,1247,981,1864]" captionTargetId="figure@9.[340,1247,981,1864]" captionTargetPageId="9" captionText="FIGURE 7. Gubernatoriana alcocki n. sp., holotype male (ZSI, WRC—C. 1513). A, dorsal view; B, frontal view; C, ventral view; D, right or major cheliped; E, right ambulatory leg (p 3); F, anterior male thoracic sternites; G, male abdomen; H, left third maxilliped. Scale bars = 5 mm (A, C), 2 mm (B, D – G), 1 mm (H)." httpUri="https://zenodo.org/record/1053937/files/figure.png" pageId="11" pageNumber="580">Fig. 7A</figureCitation>
) [vs. short outer margin of the exorbital angle (see
<bibRefCitation id="EFD44B5EFFD7FFEAFBA066D4FAC3D2E1" author="Pati, S. K. &amp; Sharma, R. M." box="[1048,1296,908,933]" journalOrPublisher="Occasional Paper, No. 363" pageId="11" pageNumber="580" pagination="1 - 44" refString="Pati, S. K. &amp; Sharma, R. M. (2014 b) Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India, Occasional Paper, No. 363, 1 - 44. [published by the Director, Zoological Survey of India, Kolkata]" title="Freshwater crabs (Crustacea: Decapoda: Brachyura: Gecarcinucidae) in the collection of the Western Regional Centre, Pune. Records of the Zoological Survey of India" type="book chapter" year="2014" yearSuffix="b">Pati &amp; Sharma 2014b</bibRefCitation>
: pl. VII, fig. 7A for
<taxonomicName id="4C454D2CFFD7FFEAFF5066E8FE45D28C" authorityName="Alcock" authorityYear="1909" box="[232,406,944,968]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="gubernatoris">
<emphasis id="B931EABDFFD7FFEAFF5066E8FE45D28C" box="[232,406,944,968]" italics="true" pageId="11" pageNumber="580">G. gubernatoris</emphasis>
</taxonomicName>
)].
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD7FFEAFF7F668CFD7FD4FB" pageId="11" pageNumber="580" type="reference_group">
<paragraph id="8BFA36AFFFD7FFEAFF7F668CFAA8D539" blockId="11.[151,1437,151,1544]" pageId="11" pageNumber="580">
<taxonomicName id="4C454D2CFFD7FFEAFF7F668CFDD0D2A8" ID-CoL="7WDYS" authorityName="Klaus, Fernandez &amp; Yeo" authorityYear="2014" box="[199,515,980,1004]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="basalticola">
<emphasis id="B931EABDFFD7FFEAFF7F668CFDD0D2A8" box="[199,515,980,1004]" italics="true" pageId="11" pageNumber="580">Gubernatoriana basalticola</emphasis>
</taxonomicName>
has some unique morphological features [strongly transverse carapace (cw/cl = 1.8), distinctly convergent posterolateral margin, conspicuously broad frontal margin (fw/cw = 0.7), massive major chela, and elongated G2 terminal article (see
<bibRefCitation id="EFD44B5EFFD7FFEAFD136144FCAAD570" author="Klaus, S. &amp; Fernandez, K. &amp; Yeo, D. C. J." box="[683,889,1052,1077]" journalOrPublisher="Zoological Scripta" pageId="11" pageNumber="580" pagination="651 - 660" part="43" publicationUrl="http://dx.doi.org/10.1111/zsc.12078" refString="Klaus, S., Fernandez, K. &amp; Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoological Scripta, 43 (6), 651 - 660. http: // dx. doi. org / 10.1111 / zsc. 12078" title="Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae)" type="journal article" year="2014">
Klaus
<emphasis id="B931EABDFFD7FFEAFD4F6145FCFFD570" box="[759,812,1052,1076]" italics="true" pageId="11" pageNumber="580">et al</emphasis>
. 2014
</bibRefCitation>
: figs. 3A–C)] that neither its congeners nor its related genera (
<taxonomicName id="4C454D2CFFD7FFEAFEED6118FD02D51C" ID-CoL="7PGS5" authority="Pati &amp; Sharma, 2014" authorityName="Pati &amp; Sharma" authorityYear="2014" box="[341,721,1087,1112]" class="Malacostraca" family="Gecarcinucidae" genus="Ghatiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B931EABDFFD7FFEAFEED6118FE6CD51C" box="[341,447,1088,1112]" italics="true" pageId="11" pageNumber="580">Ghatiana</emphasis>
<bibRefCitation id="EFD44B5EFFD7FFEAFE766167FD02D51C" author="Pati, S. K. &amp; Sharma, R. M." box="[462,721,1087,1112]" journalOrPublisher="India. Journal of Natural History" pageId="11" pageNumber="580" pagination="1279 - 1298" part="48" publicationUrl="http://dx.doi.org/10.1080/00222933.2013.859315" refString="Pati, S. K. &amp; Sharma, R. M. (2014 a) Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains, India. Journal of Natural History, 48 (21 - 22), 1279 - 1298. http: // dx. doi. org / 10.1080 / 00222933.2013.859315" title="Description of Ghatiana, a new genus of freshwater crab, with two new species and a new species of Gubernatoriana (Crustacea: Decapoda: Brachyura: Gecarcinucidae) from the Western Ghat Mountains" type="journal article" year="2014">Pati &amp; Sharma, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C454D2CFFD7FFEAFCA06118FB9ED51C" ID-CoL="7PLMY" authority="Bott, 1970" authorityName="Bott" authorityYear="1970" box="[792,1101,1088,1112]" class="Malacostraca" family="Gecarcinucidae" genus="Inglethelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B931EABDFFD7FFEAFCA06118FC6DD51C" box="[792,958,1088,1112]" italics="true" pageId="11" pageNumber="580">Inglethelphusa</emphasis>
<bibRefCitation id="EFD44B5EFFD7FFEAFC756118FB9ED51C" author="Bott, R." box="[973,1101,1088,1112]" journalOrPublisher="Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft" pageId="11" pageNumber="580" pagination="1 - 338" part="526" refString="Bott, R. (1970) Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda). Abhandlungen der Senckenbergischen Naturforschenden Gesellschaft, 526, 1 - 338." title="Die Susswasserkrabben von Europa, Asien, Australien und ihre Stammesgeschichte. Eine Revision der Potamoidea und der Parathelphusoidea (Crustacea, Decapoda)" type="journal article" year="1970">Bott, 1970</bibRefCitation>
</taxonomicName>
) possess, and this supports recognizing the taxon as a new genus (
<bibRefCitation id="EFD44B5EFFD7FFEAFDF0613CFCDDD538" author="Klaus, S. &amp; Fernandez, K. &amp; Yeo, D. C. J." box="[584,782,1124,1149]" journalOrPublisher="Zoological Scripta" pageId="11" pageNumber="580" pagination="651 - 660" part="43" publicationUrl="http://dx.doi.org/10.1111/zsc.12078" refString="Klaus, S., Fernandez, K. &amp; Yeo, D. C. J. (2014) Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae). Zoological Scripta, 43 (6), 651 - 660. http: // dx. doi. org / 10.1111 / zsc. 12078" title="Phylogeny of the freshwater crabs of the Western Ghats (Brachyura, Gecarcinucidae)" type="journal article" year="2014">
Klaus
<emphasis id="B931EABDFFD7FFEAFD2A613DFD10D538" box="[658,707,1124,1148]" italics="true" pageId="11" pageNumber="580">et al</emphasis>
. 2014
</bibRefCitation>
). Molecular studies are suggested to solve the problem.
</paragraph>
<paragraph id="8BFA36AFFFD7FFEAFF7561DFFD7FD4FB" blockId="11.[151,1437,151,1544]" pageId="11" pageNumber="580">
<emphasis id="B931EABDFFD7FFEAFF7561DFFE47D5E4" bold="true" box="[205,404,1159,1184]" pageId="11" pageNumber="580">Ecological notes.</emphasis>
Specimens of
<taxonomicName id="4C454D2CFFD7FFEAFD8F61D0FD71D5E4" ID-CoL="84FVK" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[567,674,1160,1184]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFD8F61D0FD71D5E4" box="[567,674,1160,1184]" italics="true" pageId="11" pageNumber="580">G. alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFD1461DFFD38D5E4" box="[684,747,1159,1184]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFD1461DFFD38D5E4" bold="true" box="[684,747,1159,1184]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
were collected beneath medium sized (
<quantity id="4CBD9B4AFFD7FFEAFB1961DFFACFD5E4" box="[1185,1308,1159,1184]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="3.5" metricValueMin="2.5" pageId="11" pageNumber="580" unit="mm" value="30.0" valueMax="35.0" valueMin="25.0">25–35 mm</quantity>
) pebbles in ephemeral streams and from the adjacent rock formations on a high mountain plateau. This species might be confined to high altitude areas of the
<collectingRegion id="4981F84DFFD7FFEAFD8E6188FD42D5A3" box="[566,657,1232,1255]" country="Sri Lanka" name="Western" pageId="11" pageNumber="580">Western</collectingRegion>
Ghats since the species is not so far recorded from mountain crests or slopes (S.K. Pati, unpublished data). Individuals were found in association with
<collectingCountry id="F352763FFFD7FFEAFB9161ACFB5BD449" box="[1065,1160,1268,1293]" name="India" pageId="11" pageNumber="580">Bombay</collectingCountry>
Swampeel,
<taxonomicName id="4C454D2CFFD7FFEAFAAC61ADFA48D448" ID-CoL="8NSM3" box="[1300,1435,1269,1292]" class="Insecta" family="Acrididae" genus="Monopterus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Orthoptera" pageId="11" pageNumber="580" phylum="Arthropoda" rank="genus">
<emphasis id="B931EABDFFD7FFEAFAAC61ADFA48D448" box="[1300,1435,1269,1292]" italics="true" pageId="11" pageNumber="580">Monopterus</emphasis>
</taxonomicName>
cf.
<emphasis id="B931EABDFFD7FFEAFF076040FEC3D474" box="[191,272,1304,1328]" italics="true" pageId="11" pageNumber="580">indicus</emphasis>
(Silas &amp; Dawson, 1961). The new species and the eel are abundant at Vankusawade, which is the breeding ground for both. On one occasion, an eel was observed preying upon a crab (B.V. Jadhav, personal communication). Due to the small size of juvenile eels, crabs might also be predating on these young eels. Crabs are generally found in large numbers on the mountain plateau during the monsoon (June to September), but large populations were seen even during dry seasons.
</paragraph>
</subSubSection>
<subSubSection id="C35F6524FFD7FFEAFF7F6094FD34D74C" pageId="11" pageNumber="580" type="description">
<paragraph id="8BFA36AFFFD7FFEAFF7F6094FD34D74C" blockId="11.[151,1437,151,1544]" pageId="11" pageNumber="580">
<emphasis id="B931EABDFFD7FFEAFF7F6094FDD7D4A1" bold="true" box="[199,516,1484,1509]" pageId="11" pageNumber="580">Geographical distribution.</emphasis>
<taxonomicName id="4C454D2CFFD7FFEAFDB26094FCC4D4A0" authority="Pati &amp; Thackeray &amp; Khaire, 2016" authorityName="Pati &amp; Thackeray &amp; Khaire" authorityYear="2016" box="[522,791,1484,1508]" class="Malacostraca" family="Gecarcinucidae" genus="Gubernatoriana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="580" phylum="Arthropoda" rank="species" species="alcocki" status="sp. nov.">
<emphasis id="B931EABDFFD7FFEAFDB26094FCC4D4A0" box="[522,791,1484,1508]" italics="true" pageId="11" pageNumber="580">Gubernatoriana alcocki</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20257C6FFD7FFEAFCA56094FC8FD4A1" box="[797,860,1484,1509]" pageId="11" pageNumber="580" rank="species">
<emphasis id="B931EABDFFD7FFEAFCA56094FC8FD4A1" bold="true" box="[797,860,1484,1509]" pageId="11" pageNumber="580">n. sp.</emphasis>
</taxonomicNameLabel>
is known only from the
<typeStatus id="54FE880DFFD7FFEAFBD46095FB4FD4A1" box="[1132,1180,1485,1509]" pageId="11" pageNumber="580">type</typeStatus>
locality, Vankusawade near Ghanbi Vatole in Satara district of
<collectingRegion id="4981F84DFFD7FFEAFDEB60B7FD30D74C" box="[595,739,1519,1544]" country="India" name="Maharashtra" pageId="11" pageNumber="580">Maharashtra</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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