258 lines
33 KiB
XML
258 lines
33 KiB
XML
<document id="FE4AFA7177E793806068727F6BD1BF5C" ID-DOI="10.11646/zootaxa.2048.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="5323018" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629617541061" checkinUser="felipe" docAuthor="Hendrich, Lars & Watts, Chris H. S." docDate="2009" docId="03EC87A5FFFD5819B59EFF6AFF474C7E" docLanguage="en" docName="zt02048p030.pdf" docOrigin="Zootaxa 2048 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.2048.1.1" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Carabhydrus turaki Hendrich & Watts, 2009, sp.n." docType="treatment" docVersion="4" lastPageNumber="25" masterDocId="FFD5FFDDFFEA5801B509FFFDFFA94A11" masterDocTitle="Taxonomic revision of the Australian predaceous water beetle genus Carabhydrus Watts, 1978 (Col. Dytiscidae, Hydroporinae, Hydroporini)" masterLastPageNumber="30" masterPageNumber="1" pageNumber="24" updateTime="1699083959834" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:mods id="EBCC1E062F8662276F1B39740617CE07" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="2330BC13DF2319402342A351FA43C619">
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<mods:title id="1D01A8F1E3EDB790813187F82ECA95C6">Taxonomic revision of the Australian predaceous water beetle genus Carabhydrus Watts, 1978 (Col. Dytiscidae, Hydroporinae, Hydroporini)</mods:title>
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<mods:name id="D547F2A9B0180F314FD205B79F584792" type="personal">
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<mods:roleTerm id="FBCA9C4748427447C9EC41480A9E407C">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F2A4A13FB7C49CE5307CC3DE4DDCB592">Hendrich, Lars</mods:namePart>
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<mods:affiliation id="E2E891319CE5DC8DFFC2140ABB444870">Zoologische Staatssammlung, Münchhausenstrasse 21, D- 81247 München, Germany. E-mail: hendrich 1 @ aol. com</mods:affiliation>
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<mods:name id="AA397860889907706E7952980C2EBADE" type="personal">
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<mods:roleTerm id="B4DFAC2B4BA2F7B478479564212D6762">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="2E7F41AE639C266260AF7DD4140F2326">Watts, Chris H. S.</mods:namePart>
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<mods:affiliation id="50B8FB04D123C81489779BE1ABAA8300">South Australian Museum, North Terrace, Adelaide, South Australia 5000, Australia. E-mail: watts. chris @ saugov. sa. gov. au</mods:affiliation>
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</mods:name>
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<mods:typeOfResource id="576324F0EB6660D541A82776BA006BF9">text</mods:typeOfResource>
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<mods:title id="AF8188B18F53A5FC95F165B8028372F8">Zootaxa</mods:title>
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<mods:part id="DD87967FE66499353310E9F2DE9C79FF">
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<mods:date id="4307692BAF06F7AF12D66CAB2E5E853E">2009</mods:date>
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<mods:number id="4F2A878F8C56052E43B792F40C0F1A89">2009-03-23</mods:number>
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<mods:detail id="B67A75F001453B4B0A5C8CC480710D3E" type="volume">
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<mods:number id="A31587CF3F5BA45974D1CBE36E4A8311">2048</mods:number>
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</mods:detail>
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<mods:number id="3BDA3411A1B5AAEC2B28CA9A62D7D302">1</mods:number>
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<mods:extent id="12914CAFBC1CBCBFDAA66D79C138021B" unit="page">
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<mods:start id="1F8FC433F506811FFA8F41E9597F7857">1</mods:start>
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<mods:end id="7026501113154C2AB29858814C217138">30</mods:end>
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<mods:url id="E4B95A62AD32F4A67A6A91DD1CBE431A">https://biotaxa.org/Zootaxa/article/view/zootaxa.2048.1.1</mods:url>
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</mods:location>
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<mods:classification id="447E31D4308A043779C5DF192D8B8BA4">journal article</mods:classification>
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<mods:identifier id="64339F942AB9DE1196ABDF0DF641F9C4" type="DOI">10.11646/zootaxa.2048.1.1</mods:identifier>
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<mods:identifier id="306F0C5BCBA81F2FE4EFE3F0C2275840" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="9F7B0E246E206B90C72E3361CD9DC810" type="Zenodo-Dep">5323018</mods:identifier>
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<treatment id="03EC87A5FFFD5819B59EFF6AFF474C7E" ID-DOI="http://doi.org/10.5281/zenodo.5334864" ID-Zenodo-Dep="5334864" LSID="urn:lsid:plazi:treatment:03EC87A5FFFD5819B59EFF6AFF474C7E" httpUri="http://treatment.plazi.org/id/03EC87A5FFFD5819B59EFF6AFF474C7E" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">
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<subSubSection id="C35F6538FFFD5816B59EFF6AFE614AA0" box="[151,456,151,177]" pageId="23" pageNumber="24" type="nomenclature">
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<paragraph id="8BFA36B3FFFD5816B59EFF6AFE614AA0" blockId="23.[151,456,151,212]" box="[151,456,151,177]" pageId="23" pageNumber="24">
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<heading id="D0B281DFFFFD5816B59EFF6AFE614AA0" bold="true" box="[151,456,151,177]" fontSize="11" level="1" pageId="23" pageNumber="24" reason="1">
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<emphasis id="B931EAA1FFFD5816B59EFF6AFE614AA0" bold="true" box="[151,456,151,177]" pageId="23" pageNumber="24">
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<taxonomicName id="4C454D30FFFD5816B59EFF6AFE2C4AA0" ID-CoL="QTZX" authorityName="Hendrich & Watts" authorityYear="2009" box="[151,389,151,177]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="turaki" status="sp. nov.">
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<emphasis id="B931EAA1FFFD5816B59EFF6AFE2C4AA0" bold="true" box="[151,389,151,177]" italics="true" pageId="23" pageNumber="24">Carabhydrus turaki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20257DAFFFD5816B485FF6AFE614AA0" box="[396,456,151,177]" pageId="23" pageNumber="24" rank="species">sp.n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6538FFFD5816B59EFF47FEDE4AC5" box="[151,375,186,212]" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BFA36B3FFFD5816B59EFF47FEDE4AC5" blockId="23.[151,456,151,212]" box="[151,375,186,212]" pageId="23" pageNumber="24">
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(
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<figureCitation id="137E2A36FFFD5816B596FF47FF434AC5" box="[159,234,186,212]" captionStart="FIGURES 5–9" captionStartId="5.[151,275,1938,1962]" captionTargetBox="[201,1382,187,1918]" captionTargetId="figure-53@5.[195,1391,172,1923]" captionTargetPageId="5" captionText="FIGURES 5–9. Habitus of Carabhydrus. 5—Carabhydrus turaki sp.n. (holotype, female); 6—C. stephanieae (paratype); 7—C. mubboonus (female); 8—C. janmillerae sp.n. (paratype, female); 9—C. niger (female) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323022" httpUri="https://zenodo.org/record/5323022/files/figure.png" pageId="23" pageNumber="24">Figs 5</figureCitation>
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,
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<figureCitation id="137E2A36FFFD5816B5F1FF47FEB14AC5" box="[248,280,186,212]" captionStart="FIGURES 28–29" captionStartId="16.[151,269,2004,2028]" captionTargetBox="[288,1303,197,1975]" captionTargetId="figure-23@16.[288,1303,197,1975]" captionTargetPageId="16" captionText="FIGURES 28–29. Distribution of the genus Carabhydrus in Australia." figureDoi="http://doi.org/10.5281/zenodo.5323030" httpUri="https://zenodo.org/record/5323030/files/figure.png" pageId="23" pageNumber="24">28</figureCitation>
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,
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<figureCitation id="137E2A36FFFD5816B42AFF47FEC74AC5" box="[291,366,186,212]" captionStart="FIGURES 36–39" captionStartId="26.[151,270,1273,1297]" captionTargetBox="[151,1433,194,1249]" captionTargetId="figure-158@26.[151,1436,194,1249]" captionTargetPageId="26" captionText="FIGURES 36–39. Type localities of 36, 37—C. turaki sp.n., head waters of Pappinbarra River, Werrikimbe National Park in New South Wales (Photos: Natacha Waddell, April and October 1995); 38, 39—C. storeyi sp.n., Mary Creek at Maryfarms NE Queensland (Photos: C. Watts, December 2008)." figureDoi="http://doi.org/10.5281/zenodo.5323036" httpUri="https://zenodo.org/record/5323036/files/figure.png" pageId="23" pageNumber="24">36, 37</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6538FFFD5816B59EFEFAFC174BAD" pageId="23" pageNumber="24" type="materials_examined">
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<paragraph id="8BFA36B3FFFD5816B59EFEFAFB5E4B33" blockId="23.[151,1436,263,2030]" box="[151,1271,263,290]" pageId="23" pageNumber="24">
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<materialsCitation id="3B2D3CEEFFFD5816B59EFEFAFB5A4B33" box="[151,1267,263,290]" county="Werrikimbe National Park" location="Head" municipality="Werrikimbe Trail" pageId="23" pageNumber="24" specimenCount="1" typeStatus="holotype">
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<emphasis id="B931EAA1FFFD5816B59EFEFAFE9E4B30" bold="true" box="[151,311,263,289]" pageId="23" pageNumber="24">
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<typeStatus id="54FE8811FFFD5816B59EFEFAFF7D4B30" box="[151,212,263,289]" pageId="23" pageNumber="24">Type</typeStatus>
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locality
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</emphasis>
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:
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<location id="8E9A6068FFFD5816B44FFEF5FE2D4B33" LSID="urn:lsid:plazi:treatment:03EC87A5FFFD5819B59EFF6AFF474C7E:8E9A6068FFFD5816B44FFEF5FE2D4B33" box="[326,388,264,290]" county="Werrikimbe National Park" municipality="Werrikimbe Trail" name="Head" pageId="23" pageNumber="24">Head</location>
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waters of
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<location id="8E9A6068FFFD5816B4F7FEF5FD7F4B33" LSID="urn:lsid:plazi:treatment:03EC87A5FFFD5819B59EFF6AFF474C7E:8E9A6068FFFD5816B4F7FEF5FD7F4B33" box="[510,726,264,290]" county="Werrikimbe National Park" municipality="Werrikimbe Trail" name="Pappinbarra River" pageId="23" pageNumber="24">Pappinbarra River</location>
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,
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<collectingMunicipality id="6B9EACC9FFFD5816B7EBFEF5FC064B33" box="[738,943,264,290]" pageId="23" pageNumber="24">Werrikimbe Trail</collectingMunicipality>
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,
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<collectingCounty id="629B4E3FFFFD5816B6B2FEF5FB5A4B33" box="[955,1267,264,290]" pageId="23" pageNumber="24">Werrikimbe National Park</collectingCounty>
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</materialsCitation>
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.
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||
</paragraph>
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||
<paragraph id="8BFA36B3FFFD5816B5CCFED3FC174BAD" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">
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||
<emphasis id="B931EAA1FFFD5816B5CCFED3FED34B59" bold="true" box="[197,378,302,328]" pageId="23" pageNumber="24">Type material</emphasis>
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||
:
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<materialsCitation id="3B2D3CEEFFFD5816B485FED3FC174BAD" accessionNumber="EPA 10834" collectionCode="AM" collectorName="Edge Eren Turak" elevation="220" location="Female" pageId="23" pageNumber="24" specimenCount="2" typeStatus="holotype">
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<emphasis id="B931EAA1FFFD5816B485FED3FDAB4B59" bold="true" box="[396,514,302,328]" pageId="23" pageNumber="24">
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||
<typeStatus id="54FE8811FFFD5816B485FED3FDAB4B59" box="[396,514,302,328]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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||
</emphasis>
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||
:
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||
<location id="8E9A6068FFFD5816B71AFED3FDD84B59" LSID="urn:lsid:plazi:treatment:03EC87A5FFFD5819B59EFF6AFF474C7E:8E9A6068FFFD5816B71AFED3FDD84B59" box="[531,625,302,328]" name="Female" pageId="23" pageNumber="24">Female</location>
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: “NSW EPA Survey MRHI HAST 09 Pappinbarra River u/s Field Study Centre 5/10/95 152.4182 31.2503 [Head waters of Pappinbarra River, Pappinbarra East Arm Road, Werrikimbe National Park,
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<quantity id="4CBD9B56FFFD5816B4EDFE81FD9A4B87" box="[484,563,380,406]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="220.0">
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||
<elevation id="0068D180FFFD5816B4EDFE81FD9A4B87" box="[484,563,380,406]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="23" pageNumber="24" unit="m" value="220.0">220 m</elevation>
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</quantity>
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, NE NSW]
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<collectorName id="26B05365FFFD5816B7CCFE81FC384B87" box="[709,913,380,406]" pageId="23" pageNumber="24">Edge Eren Turak</collectorName>
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leg. EPA 10834”, “
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<typeStatus id="54FE8811FFFD5816B176FE81FB5A4B87" box="[1151,1267,380,406]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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<taxonomicName id="4C454D30FFFD5816B1F5FE81FF774BAA" authorityName="Hendrich & Watts" authorityYear="2009" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="turaki" status="sp. nov.">
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<emphasis id="B931EAA1FFFD5816B1F5FE81FF774BAA" italics="true" pageId="23" pageNumber="24">Carabhydrus turaki</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A20257DAFFFD5816B5ECFE5FFE8C4BAD" box="[229,293,418,444]" pageId="23" pageNumber="24" rank="species">sp. n.</taxonomicNameLabel>
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Hendrich & Watts des. 2008” [red printed label] (
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<collectionCode id="ED54AE76FFFD5816B675FE5FFC184BAD" box="[892,945,418,444]" country="Australia" name="Australian Museum" pageId="23" pageNumber="24" type="Museum">AM</collectionCode>
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||
).
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35F6538FFFD5819B5CCFE35FE264952" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24" type="description">
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<paragraph id="8BFA36B3FFFD5816B5CCFE35FB4A4BF2" blockId="23.[151,1436,263,2030]" box="[197,1251,456,483]" pageId="23" pageNumber="24">
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<emphasis id="B931EAA1FFFD5816B5CCFE35FEFF4BF3" bold="true" box="[197,342,456,482]" pageId="23" pageNumber="24">Description</emphasis>
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:
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<emphasis id="B931EAA1FFFD5816B46CFE35FDB54BF3" bold="true" box="[357,540,456,482]" pageId="23" pageNumber="24">Measurements</emphasis>
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:
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<typeStatus id="54FE8811FFFD5816B725FE34FD344BF2" box="[556,669,457,483]" pageId="23" pageNumber="24" type="holotype">Holotype</typeStatus>
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||
: TL =
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<quantity id="4CBD9B56FFFD5816B7E2FE34FCFF4BF2" box="[747,854,457,483]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" pageId="23" pageNumber="24" unit="mm" value="2.35">2.35 mm</quantity>
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, TL-H =
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||
<quantity id="4CBD9B56FFFD5816B6CDFE34FB884BF2" box="[964,1057,457,483]" metricMagnitude="-3" metricUnit="m" metricValue="2.1" pageId="23" pageNumber="24" unit="mm" value="2.1">2.1 mm</quantity>
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||
, MW =
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||
<quantity id="4CBD9B56FFFD5816B188FE34FB764BF2" box="[1153,1247,457,483]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" pageId="23" pageNumber="24" unit="mm" value="1.1">1.1 mm</quantity>
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||
.
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||
</paragraph>
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||
<paragraph id="8BFA36B3FFFD5816B5CCFE12FC03486F" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">
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||
<emphasis id="B931EAA1FFFD5816B5CCFE12FEA44818" bold="true" box="[197,269,495,521]" pageId="23" pageNumber="24">Color</emphasis>
|
||
(
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<figureCitation id="137E2A36FFFD5816B416FE0DFEC0481B" box="[287,361,496,522]" captionStart="FIGURES 5–9" captionStartId="5.[151,275,1938,1962]" captionTargetBox="[201,1382,187,1918]" captionTargetId="figure-53@5.[195,1391,172,1923]" captionTargetPageId="5" captionText="FIGURES 5–9. Habitus of Carabhydrus. 5—Carabhydrus turaki sp.n. (holotype, female); 6—C. stephanieae (paratype); 7—C. mubboonus (female); 8—C. janmillerae sp.n. (paratype, female); 9—C. niger (female) (scale bar 1 mm) (Photos: L. Hendrich)." figureDoi="http://doi.org/10.5281/zenodo.5323022" httpUri="https://zenodo.org/record/5323022/files/figure.png" pageId="23" pageNumber="24">Fig. 5</figureCitation>
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): Almost dark reddish brown to black, the front of clypeus, antennae, palpi and legs pale reddish brown. Pronotum black with narrow lateral reddish brown margin. Two large basal elytral marks extending to sides but not to suture, and a vague apical band of elytra reddish brown. Ventral surface black except epipleura and last three abdominal segments reddish brown.
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</paragraph>
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||
<paragraph id="8BFA36B3FFFD5816B5CCFD77FD974909" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">
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||
<emphasis id="B931EAA1FFFD5816B5CCFD77FEEB48B5" bold="true" box="[197,322,650,676]" pageId="23" pageNumber="24">Sculpture</emphasis>
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||
: Head anterior to dorsal impression with deep punctures separated by a distance about 1 ½ equal to their own diameters, and a few sparse, minute punctures; interspaces smooth and shiny, microreticulate along hind edge on a strip along small eyes, and in two shallow depressions medially from indistinct swellings above antennae.
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</paragraph>
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<paragraph id="8BFA36B3FFFD5816B5CCFCD8FADB499D" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">Pronotum shining sparsely punctuate medially in front, denser on rest of surface but difficult to see because of many small wrinkles connecting punctures, microreticulation absent. Wrinkles transverse in basal half. Strongest between striae, fine and directed anteriorly or obliquely to outside near front ends of striae.</paragraph>
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<paragraph id="8BFA36B3FFFD5816B5CCFC64FD864E36" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">Elytron with dual irregular punctation similar to that of pronotum but more dense, surface somewhat uneven especially near middle where punctures are larger, near tip they are fine. Lateral part and apical half of elytra with fine microreticulation; epipleura simple, distally narrowed, disappears in distal fifth with dense, more or less contiguous punctures.</paragraph>
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<paragraph id="8BFA36B3FFFD5816B5CCFBC9FEAB4EF9" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">Ventral surface with punctures similar in size than those of dorsal surface, denser and more irregular with surfaces of sclerites irregularly rugose; punctures of metacoxal plate and sterna 1 and 2 not evidently larger than those elsewhere on ventral surface; metasternum medially and metacoxa between metacoxal lines sparsely and finely punctate. Legs with femora, tibiae, and dorsal surface of metatarsomeres 1–3 strongly punctate.</paragraph>
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<paragraph id="8BFA36B3FFFD5816B5CCFB08FE2E4F92" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">
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<emphasis id="B931EAA1FFFD5816B5CCFB08FEEB4F1E" bold="true" box="[197,322,1269,1295]" pageId="23" pageNumber="24">Structure</emphasis>
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: Head very broad, with cervical stria between posterior edge of eyes. Lateral margins of clypeus somewhat produced laterally above base of antenna; frontal fovea distinct, ovate; dorsal surface with a strong transverse impression between posterior margins of eyes; maxillary and labial palpi each with apical palpomere fusiform.
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</paragraph>
|
||
<paragraph id="8BFA36B3FFFD5816B5CCFA6DFD994CCE" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">Pronotum slightly transverse, less sinuate and cordiform, at base same width as head. Basal half flat, anterior part curved down, especially near projecting anterior angles where there is a slight depression, parallel to rear margin of head. Lateral margin in dorsal aspect more or less evenly rounded; lateral bead sharply defined, narrow; basal margin slightly lobed medially; disc with a pair of broad and deep longitudinal impressions, each extending from apical third of disc to hind margin and strongly limited on outer side by a more or less carinate ridge, one large deep depression just behind centre of pronotum and of same size and form as longitudinal impressions, mesal margin less distinct; several additional shorter and more irregular rugae or impressions as well as elongate punctures along basal margin adjacent to the major impressions. Scutellum visible, short and broad.</paragraph>
|
||
<paragraph id="8BFA36B3FFFD5816B5CCF911FD7E4DB1" blockId="23.[151,1436,263,2030]" pageId="23" pageNumber="24">Elytra together elongate-oval with point of maximum width near middle, sides converging basally; disc of each elytron with a medial and a sublateral, broad, deep and sharply defined longitudinal groove each bearing a row of serial punctures, and a less evidently defined lateral groove adjacent to lateral margin; medial and sublateral groove beginning approximately opposite basolateral impression of pronotum, sharply defined on basal three-fourths, becoming shallower apically.</paragraph>
|
||
<paragraph id="8BFA36B3FFFD5819B5CCF850FC034B62" blockId="23.[151,1436,263,2030]" lastBlockId="24.[151,1436,152,1647]" lastPageId="24" lastPageNumber="25" pageId="23" pageNumber="24">Ventral surface, lateral portions of thorax and abdomen narrowed ventrally, medially forming a more or less flattened area comprising prosternal process, medial area of metasternum, intralinear space of metacoxae, and abdominal sterna medially. Prosternum with anterolateral pore well removed from anterior margin. Prosternal process hardly deflexed, grooved in middle, sides raised, oval. Metasternum largely in a vertical plane, prosternal process touches metasternum between mesocoxae. Metasternum medially flat, faintly raised anteriorly joined lines extend from between mesocoxae posteriorly, where they become indistinct. Metacoxal lines raised, first directed a little outward as usual but their front ends curved slightly medially. Metacoxal process flat, distally hardly sinuous, indistinctly notched in middle.</paragraph>
|
||
<paragraph id="8BFA36B3FFF25819B5CCFE7DFBCA490D" blockId="24.[151,1436,152,1647]" pageId="24" pageNumber="25">Legs long. Coxae and trochanters normal. Femora simple, metafemur shorter than others and much shorter than tibia. All femora largely visible from above. Profemur with anteroventral margin slightly emarginate subapically and fringed with a row of short, stout setae. Protibia with medially enlarged apical half, a regular row of very fine and short setae on inside below. Three basal tarsal segments enlarged, symmetrical, with adhesive hairs below and long fine hairs above. Protibia with medially enlarged apical half, a regular row of very fine short setae on inside below. Apex oblique on outside, with several spines. Normal spurs present. Three basal tarsal segments enlarged, symmetrical, with adhesive hairs below and long fine hairs above. Tarsal segment 4 minute, fifth slender, long, with equal slender claws. Mesotibia straight, inner face with long hairs. Pro- and mesotarsomeres 1–3 ovate; protarsomere 4 small, situated within emargination of tarsomere 3 but visible in dorsal aspect. Hind legs slender; metatibia straight, evenly broadened from base to apex, natatorial setae absent from ventral face in female, metatarsal claws equal.</paragraph>
|
||
<paragraph id="8BFA36B3FFF25819B5CCFCD5FE264952" blockId="24.[151,1436,152,1647]" box="[197,399,808,835]" pageId="24" pageNumber="25">
|
||
<emphasis id="B931EAA1FFF25819B5CCFCD5FEAD4953" bold="true" box="[197,260,808,834]" pageId="24" pageNumber="25">Male</emphasis>
|
||
: Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6538FFF25819B5CCFCB2FF5A49A6" pageId="24" pageNumber="25" type="etymology">
|
||
<paragraph id="8BFA36B3FFF25819B5CCFCB2FF5A49A6" blockId="24.[151,1436,152,1647]" pageId="24" pageNumber="25">
|
||
<emphasis id="B931EAA1FFF25819B5CCFCB2FEE74978" bold="true" box="[197,334,847,873]" pageId="24" pageNumber="25">Etymology</emphasis>
|
||
: The species is dedicated to the Australian limnologist and ecologist Dr Eren Turak (NSW Department of Environment and Climate Change, Sydney) who collected the only known specimen of the species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6538FFF25819B5CCFC3EFD1A4E69" pageId="24" pageNumber="25" type="diagnosis">
|
||
<paragraph id="8BFA36B3FFF25819B5CCFC3EFD1A4E69" blockId="24.[151,1436,152,1647]" pageId="24" pageNumber="25">
|
||
<emphasis id="B931EAA1FFF25819B5CCFC3EFE7B49CC" bold="true" box="[197,466,963,989]" pageId="24" pageNumber="25">Differential diagnosis</emphasis>
|
||
: In colour and size
|
||
<taxonomicName id="4C454D30FFF25819B7C9FC39FC8349CC" authorityName="Hendrich & Watts" authorityYear="2009" box="[704,810,964,989]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="turaki" status="sp. nov.">
|
||
<emphasis id="B931EAA1FFF25819B7C9FC39FC8349CC" box="[704,810,964,989]" italics="true" pageId="24" pageNumber="25">C. turaki</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B931EAA1FFF25819B63BFC3EFCC749CC" bold="true" box="[818,878,963,989]" pageId="24" pageNumber="25">
|
||
<taxonomicNameLabel id="A20257DAFFF25819B63BFC3EFCC749CC" box="[818,878,963,989]" pageId="24" pageNumber="25" rank="species">sp.n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C454D30FFF25819B6FDFC39FBDC49CC" authorityName="Zwick" authorityYear="1981" box="[1012,1141,964,989]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="andreas">
|
||
<emphasis id="B931EAA1FFF25819B6FDFC39FBDC49CC" box="[1012,1141,964,989]" italics="true" pageId="24" pageNumber="25">C. andreas</emphasis>
|
||
</taxonomicName>
|
||
but has a more flattened body, much better defined and very deep elytral grooves reminisent of those in
|
||
<emphasis id="B931EAA1FFF25819B175FC17FA9F4E12" box="[1148,1334,1002,1027]" italics="true" pageId="24" pageNumber="25">
|
||
<taxonomicName id="4C454D30FFF25819B175FC17FA984E12" authorityName="Larson & Storey" authorityYear="1994" box="[1148,1329,1002,1027]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="mubboonus">C. mubboonus</taxonomicName>
|
||
,
|
||
</emphasis>
|
||
a wider appearance than
|
||
<taxonomicName id="4C454D30FFF25819B457FBECFE764E3B" authorityName="Zwick" authorityYear="1981" box="[350,479,1041,1066]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="andreas">
|
||
<emphasis id="B931EAA1FFF25819B457FBECFE764E3B" box="[350,479,1041,1066]" italics="true" pageId="24" pageNumber="25">C. andreas</emphasis>
|
||
</taxonomicName>
|
||
due to the pronotum being less strongly cordiform and sinuate laterally, the pale elytral spots are less defined than in
|
||
<taxonomicName id="4C454D30FFF25819B776FBC5FCA44E40" authorityName="Zwick" authorityYear="1981" box="[639,781,1080,1105]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="andreas">
|
||
<emphasis id="B931EAA1FFF25819B776FBC5FCA44E40" box="[639,781,1080,1105]" italics="true" pageId="24" pageNumber="25">C. andreas</emphasis>
|
||
</taxonomicName>
|
||
and almost half of the elytra is covered by a fine microretulation which is absent in
|
||
<taxonomicName id="4C454D30FFF25819B724FBA2FD054E69" authorityName="Zwick" authorityYear="1981" box="[557,684,1119,1144]" class="Insecta" family="Dytiscidae" genus="Carabhydrus" kingdom="Animalia" order="Coleoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="andreas">
|
||
<emphasis id="B931EAA1FFF25819B724FBA2FD054E69" box="[557,684,1119,1144]" italics="true" pageId="24" pageNumber="25">C. andreas</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6538FFF25819B5CCFB78FF474C7E" pageId="24" pageNumber="25" type="distribution">
|
||
<paragraph id="8BFA36B3FFF25819B5CCFB78FE584EFC" blockId="24.[151,1436,152,1647]" pageId="24" pageNumber="25">
|
||
<emphasis id="B931EAA1FFF25819B5CCFB78FEF74E8E" bold="true" box="[197,350,1157,1183]" pageId="24" pageNumber="25">Distribution</emphasis>
|
||
: North-eastern
|
||
<collectingRegion id="4981F851FFF25819B71CFB78FD4F4E8E" box="[533,742,1157,1183]" country="Australia" name="New South Wales" pageId="24" pageNumber="25">New South Wales</collectingRegion>
|
||
(
|
||
<figureCitation id="137E2A36FFF25819B7FCFB78FCE44E8E" box="[757,845,1157,1183]" captionStart="FIGURES 28–29" captionStartId="16.[151,269,2004,2028]" captionTargetBox="[288,1303,197,1975]" captionTargetId="figure-23@16.[288,1303,197,1975]" captionTargetPageId="16" captionText="FIGURES 28–29. Distribution of the genus Carabhydrus in Australia." figureDoi="http://doi.org/10.5281/zenodo.5323030" httpUri="https://zenodo.org/record/5323030/files/figure.png" pageId="24" pageNumber="25">Fig. 28</figureCitation>
|
||
). Only known from the
|
||
<typeStatus id="54FE8811FFF25819B16CFB78FB3E4E8E" box="[1125,1175,1157,1183]" pageId="24" pageNumber="25">type</typeStatus>
|
||
locality. According to the geographical coordinates on the locality label most probably a head water stream of the Pappinbarra River in Werrikimbe National Park.
|
||
</paragraph>
|
||
<paragraph id="8BFA36B3FFF25819B5CCFB05FF474C7E" blockId="24.[151,1436,152,1647]" pageId="24" pageNumber="25">
|
||
<emphasis id="B931EAA1FFF25819B5CCFB05FE8E4F03" bold="true" box="[197,295,1272,1298]" pageId="24" pageNumber="25">Habitat</emphasis>
|
||
: A rheophilic species collected in a river with good riffle habitats with cobble (40%), boulder (20%) and pebble (15%) substratum and fringed by old growth forest (
|
||
<figureCitation id="137E2A36FFF25819B6C6FADDFBFE4F2B" box="[975,1111,1312,1338]" captionStart="FIGURES 36–39" captionStartId="26.[151,270,1273,1297]" captionTargetBox="[151,1433,194,1249]" captionTargetId="figure-158@26.[151,1436,194,1249]" captionTargetPageId="26" captionText="FIGURES 36–39. Type localities of 36, 37—C. turaki sp.n., head waters of Pappinbarra River, Werrikimbe National Park in New South Wales (Photos: Natacha Waddell, April and October 1995); 38, 39—C. storeyi sp.n., Mary Creek at Maryfarms NE Queensland (Photos: C. Watts, December 2008)." figureDoi="http://doi.org/10.5281/zenodo.5323036" httpUri="https://zenodo.org/record/5323036/files/figure.png" pageId="24" pageNumber="25">Figs 36, 37</figureCitation>
|
||
). The width varied between 4 and
|
||
<quantity id="4CBD9B56FFF25819B5E8FABBFEB44F71" box="[225,285,1350,1376]" metricMagnitude="1" metricUnit="m" metricValue="1.2" pageId="24" pageNumber="25" unit="m" value="12.0">12 m</quantity>
|
||
and the oxygen level was high (dissolved oxygen
|
||
<quantity id="4CBD9B56FFF25819B671FABBFC744F71" box="[888,989,1350,1376]" metricMagnitude="-5" metricUnit="kg" metricValue="1.04" pageId="24" pageNumber="25" unit="mg" value="10.4">10.4 mg</quantity>
|
||
/l). No true macrophytes were present but a small amount of filamentous algae and a tiny amount of aquatic moss were observed in
|
||
<date id="FFFB1073FFF25819B1D6FA90FACA4F96" box="[1247,1379,1389,1415]" pageId="24" pageNumber="25" value="1996-04">April 1996</date>
|
||
. The riparian vegetation was fairly dense with three distinct layers. The bottom layer was sedges, grasses and ferns dominated by
|
||
<taxonomicName id="4C454D30FFF25819B44DFA46FD974FC5" authorityName="Labillardiere" authorityYear="1805" box="[324,574,1467,1492]" class="Liliopsida" family="Asparagaceae" genus="Lomandra" kingdom="Plantae" order="Asparagales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="longifolia">
|
||
<emphasis id="B931EAA1FFF25819B44DFA46FD974FC5" box="[324,574,1467,1492]" italics="true" pageId="24" pageNumber="25">Lomandra longifolia</emphasis>
|
||
</taxonomicName>
|
||
and bracken (
|
||
<emphasis id="B931EAA1FFF25819B7E5FA46FC5B4FC5" box="[748,1010,1467,1492]" italics="true" pageId="24" pageNumber="25">Pretidium esculentum</emphasis>
|
||
). The middle layer was shrubs and small trees (e.g
|
||
<taxonomicName id="4C454D30FFF25819B45EFA1CFE7E4FEA" box="[343,471,1505,1531]" class="Magnoliopsida" family="Fabaceae" genus="Acacia" kingdom="Plantae" order="Fabales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="undetermined">
|
||
<emphasis id="B931EAA1FFF25819B45EFA1CFE034FEB" box="[343,426,1505,1530]" italics="true" pageId="24" pageNumber="25">Acacia</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D30FFF25819B4ECFA1CFD584FEB" box="[485,753,1505,1530]" class="Magnoliopsida" family="Cunoniaceae" genus="Callicoma" kingdom="Plantae" order="Oxalidales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="serratifolia">
|
||
<emphasis id="B931EAA1FFF25819B4ECFA1CFD584FEB" box="[485,753,1505,1530]" italics="true" pageId="24" pageNumber="25">Callicoma serratifolia</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D30FFF25819B624FA1CFC4B4FEB" box="[813,994,1505,1530]" class="Magnoliopsida" family="Moraceae" genus="Ficus" kingdom="Plantae" order="Rosales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="coronota">
|
||
<emphasis id="B931EAA1FFF25819B624FA1CFC4B4FEB" box="[813,994,1505,1530]" italics="true" pageId="24" pageNumber="25">Ficus coronota</emphasis>
|
||
</taxonomicName>
|
||
). The upper layer consisted of taller trees such as large acacias as well as some rainforest tree species such as
|
||
<taxonomicName id="4C454D30FFF25819B102F9F5FA9A4C30" box="[1035,1331,1544,1569]" class="Magnoliopsida" family="Cunoniaceae" genus="Ceratopetalum" kingdom="Plantae" order="Oxalidales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="apetalum">
|
||
<emphasis id="B931EAA1FFF25819B102F9F5FA9A4C30" box="[1035,1331,1544,1569]" italics="true" pageId="24" pageNumber="25">Ceratopetalum apetalum</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C454D30FFF25819B04DF9F5FEBA4C59" class="Magnoliopsida" family="Ehretiaceae" genus="Ehretia" kingdom="Plantae" order="Boraginales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="acuminata">
|
||
<emphasis id="B931EAA1FFF25819B04DF9F5FEBA4C59" italics="true" pageId="24" pageNumber="25">Ehretia acuminata</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D30FFF25819B445F9D2FD8B4C59" box="[332,546,1583,1608]" class="Magnoliopsida" family="Myrtaceae" genus="Syzygium" kingdom="Plantae" order="Myrtales" pageId="24" pageNumber="25" phylum="Tracheophyta" rank="species" species="australe">
|
||
<emphasis id="B931EAA1FFF25819B445F9D2FD8B4C59" box="[332,546,1583,1608]" italics="true" pageId="24" pageNumber="25">Syzygium australe</emphasis>
|
||
</taxonomicName>
|
||
). There were some weeds such as
|
||
<emphasis id="B931EAA1FFF25819B6B3F9D2FBDF4C59" box="[954,1142,1583,1608]" italics="true" pageId="24" pageNumber="25">Camphor laurel</emphasis>
|
||
and wild tobacco (Turak in litt.).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |