146 lines
19 KiB
XML
146 lines
19 KiB
XML
<document id="97C4E5879A4F88688EF82706310AA062" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza & Reguero, Marcelo A." docDate="2018" docId="03EC879BCF598310C1922B9D8FB0F8BA" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protypotherium obstructum Ameghino 1888" docType="treatment" docVersion="7" lastPageNumber="483" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="483" updateTime="1698526258101" updateUser="ExternalLinkService">
|
||
<mods:mods id="D6451D359112684DCD57F2AB31EBE3BF" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="CD65F3101955A87403A5C271FA9210B0">
|
||
<mods:title id="5B29BA9DF40B0B3CBE71641509F9557A">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="9D488FA8D0243247793F6BC2D8AABDA3" type="personal">
|
||
<mods:role id="659C8F0B67AABD8AFDDFBF62A44AB460">
|
||
<mods:roleTerm id="A3B2C2DD65083530991EB681B7C545A7">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="39A3CDEC2BB471E1938876CF84FA1A8F">Fernández, Mercedes</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8DA63CAF0CBD3B6CAC4D7373E5FB3633" type="personal">
|
||
<mods:role id="2CF9440044F0220B4B90BBB6C533484F">
|
||
<mods:roleTerm id="001AA04F5F962AA9C0A913D83D27033B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E3653A293DF9C2DC282AA8F2F9884308">Fernicola, Juan Carlos</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F29E2F37113D959584A98F4844D1FBE1" type="personal">
|
||
<mods:role id="78F3E116092C459E2CA02072B6B0D5F4">
|
||
<mods:roleTerm id="4278B5AC580BC770EEE3BD33E8ED1575">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="FF3146D870A11003368A6CEEF190ED9E">Cerdeño, Esperanza</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="8330D9A410EB1298063B530AA275A5A9" type="personal">
|
||
<mods:role id="F2511758C13F939BDA3EF8BCB79B0720">
|
||
<mods:roleTerm id="51992D0EECD52B285FCDF8052ECD6518">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="8B4EA8B39DE5EDCD7774AAD1FC812C22">Reguero, Marcelo A.</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="AE42A48234E6968F278C1F9633005307">text</mods:typeOfResource>
|
||
<mods:relatedItem id="13F41479279EB954FB19BADB7A3A34A3" type="host">
|
||
<mods:titleInfo id="2467534DF1FA7932C8909B7867D5536F">
|
||
<mods:title id="A1F05ED0E7F17A45035C0D8BE05F4786">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="C24C8DEA84E84B108A1A16652BAFA048">
|
||
<mods:date id="DDBFEC7B2A474D8D5AEB69F7CFDAD2DB">2018</mods:date>
|
||
<mods:detail id="2812D3DAFC513C65B64EE872974A671F" type="pubDate">
|
||
<mods:number id="BAA5C8C3B15BC5000065192D5FE574F6">2018-02-28</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="320021E0B4F9535C1F9CF60B72442BE0" type="volume">
|
||
<mods:number id="44867C4AF5C418E62F02C0DD29678F44">4387</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="033A8DA511C18BBA19E48A95A47FC9FB" type="issue">
|
||
<mods:number id="E8BF75BC5A99180B1D6C8A3015997BE7">3</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="75F54BB9582B81933C4B5FB701592CB6" unit="page">
|
||
<mods:start id="DEF04564051C7F5467CCE5BF6D099F11">473</mods:start>
|
||
<mods:end id="361880FFFE77700280BE424089AF17D5">498</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="96B92CAC4483A1DD5A117DB1E3E9CD2A">journal article</mods:classification>
|
||
<mods:identifier id="0B54807AF8558CE4963305BA81BF997A" type="CLB-Dataset">30626</mods:identifier>
|
||
<mods:identifier id="4F6AC278B6D38A918F6126F8A0BABEB5" type="DOI">10.11646/zootaxa.4387.3.4</mods:identifier>
|
||
<mods:identifier id="17AC567BCF240A977A97B795F7D8FE50" type="GBIF-Dataset">43de8956-0706-4f8d-b5a9-f51c30c276d8</mods:identifier>
|
||
<mods:identifier id="3EA9AD6A97C8004D9BC5E2B77A438AFB" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="0F7BC23B92CD48BFB174430598D960F7" type="Zenodo-Dep">1187697</mods:identifier>
|
||
<mods:identifier id="EB0BB508E6DF158AB50A40CBE9E3627C" type="ZooBank">B53A6B20-5386-4E5A-A00F-A559EDF640DF</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EC879BCF598310C1922B9D8FB0F8BA" ID-DOI="http://doi.org/10.5281/zenodo.5986685" ID-GBIF-Taxon="141264032" ID-Zenodo-Dep="5986685" LSID="urn:lsid:plazi:treatment:03EC879BCF598310C1922B9D8FB0F8BA" httpUri="http://treatment.plazi.org/id/03EC879BCF598310C1922B9D8FB0F8BA" lastPageNumber="483" pageId="10" pageNumber="483">
|
||
<subSubSection id="C35F6506CF598310C1922B9D8CEFFC3B" box="[151,687,980,1006]" pageId="10" pageNumber="483" type="nomenclature">
|
||
<paragraph id="8BFA368DCF598310C1922B9D8CEFFC3B" blockId="10.[151,687,980,1006]" box="[151,687,980,1006]" pageId="10" pageNumber="483">
|
||
<heading id="D0B281E1CF598310C1922B9D8CEFFC3B" bold="true" box="[151,687,980,1006]" fontSize="11" level="1" pageId="10" pageNumber="483" reason="1">
|
||
<taxonomicName id="4C454D0ECF598310C1922B9D8CEFFC3B" authority="Ameghino 1888" authorityName="Ameghino" authorityYear="1888" box="[151,687,980,1006]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="obstructum">
|
||
<emphasis id="B931EA9FCF598310C1922B9D8CEFFC3B" bold="true" box="[151,687,980,1006]" pageId="10" pageNumber="483">
|
||
<emphasis id="B931EA9FCF598310C1922B9D8FA6FC3B" bold="true" box="[151,486,980,1006]" italics="true" pageId="10" pageNumber="483">Protypotherium obstructum</emphasis>
|
||
<bibRefCitation id="EFD44B7CCF598310C0E92B9D8CEFFC3B" box="[492,687,980,1006]" pageId="10" pageNumber="483" refId="ref16996" refString="Ameghino, F. (1888) Lista de las especies de mamiferos fosiles del Mioceno superior de Monte Hermoso hasta ahora conocidas. Obras Completas y Correspondencia Cientifica, 5, 481 - 496." type="journal article">Ameghino 1888</bibRefCitation>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6506CF598310C1922C538CB9FBE6" box="[151,761,1050,1075]" pageId="10" pageNumber="483" type="materials_examined">
|
||
<paragraph id="8BFA368DCF598310C1922C538CB9FBE6" blockId="10.[151,1437,1050,1903]" box="[151,761,1050,1075]" pageId="10" pageNumber="483">
|
||
<emphasis id="B931EA9FCF598310C1922C538F4BFBE6" bold="true" box="[151,267,1050,1075]" pageId="10" pageNumber="483">Holotype.</emphasis>
|
||
MACN-A 1677 (
|
||
<figureCitation id="137E2A08CF598310C0D72C538C72FBE6" box="[466,562,1050,1075]" captionStart="FIGURE 2" captionStartId="5.[151,249,1632,1654]" captionTargetBox="[308,1273,213,1595]" captionTargetId="figure@5.[305,1281,193,1611]" captionTargetPageId="5" captionText="FIGURE 2. Protypotherium attenuatum, syntype MACN-A 524, left maxilla with P2–M3 but lacking P1, in occlusal view (A); syntype MACN-A 628, right fragmented mandible with alveoli of i1–3 and c (broken)–m1 but lacking m2–3, in labial (B) and occlusal (C) views; reproductions of figures 21 (D), 20 (E) and 20a (F) from Ameghino (1889, plate 14). Protypotherium obstructum, holotype MACN-A 1677, lower premolar in labial, oclusal and lingual views (G); reproduction of figure 19 (H) from Ameghino (1889, plate 15). Protypotherium claudum, holotype MACN-A 551, right mandibular fragment with alveolus of p2 andp3–m2, in labial (I) and occlusal (J)views; reproductions of figures 22 (K) and 22a (L) from Ameghino (1889, plate 14). Protypotherium globosum, holotype MACN-A 4049, skull, in ventral (M) and dorsal (N) views. Protypotherium convexidens, holotype MACN-A 4054, right maxillary fragment with P3–M2 (broken), in occlusal view (O). Protypotherium diversidens, holotype MACN-A 4052, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (P). Protypotheriumcompressidens, holotype MACN-A 4029, left maxillaryfragment withP3–M3, in occlusal view (Q); holotype MACN-A 4030, left mandibular fragment with p4 (talonid)–m3, in occlusal view (R). Protypotherium lineare, holotype MACN-A4039, right mandibular fragment with alveolus of p1 and p2–m1, in occlusal view (S); holotype MACN-A 4038, left mandibular fragment with p3–m2, in occlusal view (T). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187701/files/figure.png" pageId="10" pageNumber="483">Fig. 2 G</figureCitation>
|
||
), lower premolar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6506CF598310C1C22C778EA0FA36" pageId="10" pageNumber="483" type="diagnosis">
|
||
<paragraph id="8BFA368DCF598310C1C22C778A5FFADE" blockId="10.[151,1437,1050,1903]" pageId="10" pageNumber="483">
|
||
<emphasis id="B931EA9FCF598310C1C22C778F7AFB82" bold="true" box="[199,314,1086,1111]" pageId="10" pageNumber="483">Diagnosis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B7CCF598310C04F2C778C72FB82" box="[330,562,1086,1111]" pageId="10" pageNumber="483" refId="ref16996" refString="Ameghino, F. (1888) Lista de las especies de mamiferos fosiles del Mioceno superior de Monte Hermoso hasta ahora conocidas. Obras Completas y Correspondencia Cientifica, 5, 481 - 496." type="journal article">Ameghino 1888: 14</bibRefCitation>
|
||
). “
|
||
<emphasis id="B931EA9FCF598310C3512C768A4BFADF" italics="true" pageId="10" pageNumber="483">Esta especie está representada por un solo premolar inferior, pero de una forma tan distinta de los mismos dientes de las otras especies que indica claramente no puede proceder de ninguna de las ya conocidas. Es el premolar tercero o cuarto de la mandíbula inferior, y difiere de los mismos dientes de las demás especies, por el surco perpendicular externo menos profundo, y por no presentar vestigios del surco interno opuesto. La cara perpendicular interna es así casi plana, con una muy pequeña elevación perpendicular en el medio. Largo de la corona, 0.0045. Es esta la especie más moderna del género</emphasis>
|
||
”.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF598310C1C22D5F8EA0FA36" blockId="10.[151,1437,1050,1903]" pageId="10" pageNumber="483">
|
||
English translation. This species is represented by a single lower premolar so different from the same teeth of the other known species that clearly indicates that it does not belong to any of them. It is the third or fourth lower premolar, and differs from the same teeth of the other species by the shallower external perpendicular groove and the lack of the opposite internal groove. Thus, the internal perpendicular face is almost flat, with a very small perpendicular elevation in the middle. Length of the tooth crown,
|
||
<quantity id="4CBD9B68CF598310C27D2DEF8DA5FA6B" box="[888,997,1446,1471]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="483" unit="m" value="0.0045">0.0045 m</quantity>
|
||
. This is the most modern species of the genus.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6506CF598310C1C22DA78FB0F8BA" pageId="10" pageNumber="483" type="discussion">
|
||
<paragraph id="8BFA368DCF598310C1C22DA78FFEF8F3" blockId="10.[151,1437,1050,1903]" pageId="10" pageNumber="483">
|
||
<emphasis id="B931EA9FCF598310C1C22DA78F0FF9D2" bold="true" box="[199,335,1518,1543]" pageId="10" pageNumber="483">Comments.</emphasis>
|
||
<bibRefCitation id="EFD44B7CCF598310C0532DA68FF0F9D2" box="[342,432,1519,1543]" pageId="10" pageNumber="483" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">In 1889</bibRefCitation>
|
||
, Ameghino expanded the original diagnosis of the species by describing with more detail the lower premolar collected at Monte Hermoso (
|
||
<collectingRegion id="4981F86FCF598310C3ED2E5A8DBAF9FE" box="[744,1018,1554,1579]" country="Argentina" name="Buenos Aires" pageId="10" pageNumber="483">
|
||
Buenos
|
||
<collectingRegion id="4981F86FCF598310C24E2E5B8DBAF9FE" box="[843,1018,1554,1579]" inferredBySuffix="true" pageId="10" pageNumber="483">Aires Province</collectingRegion>
|
||
</collectingRegion>
|
||
) by his brother Carlos, which was indicated by Florentino as the only assignable material. Florentino Ameghino also illustrated this tooth (
|
||
<bibRefCitation id="EFD44B7CCF598310C4212E7F8FE7F9A6" pageId="10" pageNumber="483" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">Ameghino 1889: plate 14, fig. 14</bibRefCitation>
|
||
), which is herein reproduced in
|
||
<figureCitation id="137E2A08CF598310C2362E138DFBF9A7" box="[819,955,1626,1651]" captionStart="FIGURE 2" captionStartId="5.[151,249,1632,1654]" captionTargetBox="[308,1273,213,1595]" captionTargetId="figure@5.[305,1281,193,1611]" captionTargetPageId="5" captionText="FIGURE 2. Protypotherium attenuatum, syntype MACN-A 524, left maxilla with P2–M3 but lacking P1, in occlusal view (A); syntype MACN-A 628, right fragmented mandible with alveoli of i1–3 and c (broken)–m1 but lacking m2–3, in labial (B) and occlusal (C) views; reproductions of figures 21 (D), 20 (E) and 20a (F) from Ameghino (1889, plate 14). Protypotherium obstructum, holotype MACN-A 1677, lower premolar in labial, oclusal and lingual views (G); reproduction of figure 19 (H) from Ameghino (1889, plate 15). Protypotherium claudum, holotype MACN-A 551, right mandibular fragment with alveolus of p2 andp3–m2, in labial (I) and occlusal (J)views; reproductions of figures 22 (K) and 22a (L) from Ameghino (1889, plate 14). Protypotherium globosum, holotype MACN-A 4049, skull, in ventral (M) and dorsal (N) views. Protypotherium convexidens, holotype MACN-A 4054, right maxillary fragment with P3–M2 (broken), in occlusal view (O). Protypotherium diversidens, holotype MACN-A 4052, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (P). Protypotheriumcompressidens, holotype MACN-A 4029, left maxillaryfragment withP3–M3, in occlusal view (Q); holotype MACN-A 4030, left mandibular fragment with p4 (talonid)–m3, in occlusal view (R). Protypotherium lineare, holotype MACN-A4039, right mandibular fragment with alveolus of p1 and p2–m1, in occlusal view (S); holotype MACN-A 4038, left mandibular fragment with p3–m2, in occlusal view (T). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187701/files/figure.png" pageId="10" pageNumber="483">Figure 2 H</figureCitation>
|
||
. No material is indicated as the
|
||
<typeStatus id="54FE882FCF598310C44C2E128B39F9A6" box="[1353,1401,1627,1651]" pageId="10" pageNumber="483">type</typeStatus>
|
||
in Ameghino´s catalogue, but
|
||
<bibRefCitation id="EFD44B7CCF598310C0D52E368C2FF942" box="[464,623,1662,1687]" pageId="10" pageNumber="483" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
|
||
recognised MACN-A 1677 as such. This specimen is a lower premolar that completely matches the original description (e.g., length equals
|
||
<quantity id="4CBD9B68CF598310C29B2EEA8A47F96F" box="[926,1031,1699,1723]" metricMagnitude="-3" metricUnit="m" metricValue="4.88" pageId="10" pageNumber="483" unit="mm" value="4.88">4.88 mm</quantity>
|
||
; its morphology differs from all the remaining species of the genus as the labial sulcus is extremely shallow, the lingual face is completely flat and lacks any sulcus) and posterior illustration (
|
||
<bibRefCitation id="EFD44B7CCF598310C3792EA38D77F8D6" box="[636,823,1770,1795]" pageId="10" pageNumber="483" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">Ameghino 1889</bibRefCitation>
|
||
). Therefore, MACN-A 1677 is herein confirmed as the
|
||
<typeStatus id="54FE882FCF598310C1922F478EB9F8F2" box="[151,249,1806,1831]" pageId="10" pageNumber="483" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C454D0ECF598310C01B2F468FF8F8F2" authorityName="Ameghino" authorityYear="1888" box="[286,440,1806,1831]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="10" pageNumber="483" phylum="Chordata" rank="species" species="obstructum">
|
||
<emphasis id="B931EA9FCF598310C01B2F468FF8F8F2" box="[286,440,1806,1831]" italics="true" pageId="10" pageNumber="483">P. obstructum</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF598310C1C22F7B8FB0F8BA" blockId="10.[151,1437,1050,1903]" pageId="10" pageNumber="483">
|
||
<materialsCitation id="3B2D3CD0CF598310C1C22F7B8FB0F8BA" ID-GBIF-Occurrence="1828873145" collectionCode="SALMA" location="Buenos Aires Province" pageId="10" pageNumber="483" specimenCount="1" stateProvince="Buenos Aires Province" typeStatus="holotype">
|
||
<emphasis id="B931EA9FCF598310C1C22F7B8DA9F89E" bold="true" box="[199,1001,1842,1867]" pageId="10" pageNumber="483">
|
||
Chronological and geographical distribution of the
|
||
<typeStatus id="54FE882FCF598310C2322F7B8D2BF89E" box="[823,875,1842,1867]" pageId="10" pageNumber="483">type</typeStatus>
|
||
specimen.
|
||
</emphasis>
|
||
Montehermosan
|
||
<collectionCode id="ED54AE48CF598310C5B42F7A8B53F89E" box="[1201,1299,1843,1867]" pageId="10" pageNumber="483">SALMA</collectionCode>
|
||
(
|
||
<bibRefCitation id="EFD44B7CCF598310C4212F7B8E93F8BA" pageId="10" pageNumber="483" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">Ameghino 1889</bibRefCitation>
|
||
),
|
||
<collectingRegion id="4981F86FCF598310C1E32F1E8FACF8BA" box="[230,492,1878,1903]" country="Argentina" name="Buenos Aires" pageId="10" pageNumber="483">
|
||
Buenos
|
||
<collectingRegion id="4981F86FCF598310C0442F1F8FACF8BA" box="[321,492,1878,1903]" inferredBySuffix="true" pageId="10" pageNumber="483">Aires Province</collectingRegion>
|
||
</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |