403 lines
53 KiB
XML
403 lines
53 KiB
XML
<document id="38494D0BC31E676B58CD5C658F4550E6" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza & Reguero, Marcelo A." docDate="2018" docId="03EC879BCF50831DC1922EBB8F42FA4A" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Protypotherium antiquum Ameghino 1885" docType="treatment" docVersion="7" lastPageNumber="480" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="476" updateTime="1698526258101" updateUser="ExternalLinkService">
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<mods:mods id="31D9248E2FB5D7928308914F7BAD9D5F" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="D1865414BBCFBEF6B8ABA960EFA6A7C8">
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<mods:title id="E616B3BC63BB17E18B41124A37440422">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
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</mods:titleInfo>
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<mods:name id="D1A8B5C674C09BE4AD28358A6B14EF83" type="personal">
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<mods:roleTerm id="261639B4B6FE400EB009020183B72073">Author</mods:roleTerm>
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<mods:namePart id="25E2608BA944C910A328F1C22BE845AC">Fernández, Mercedes</mods:namePart>
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</mods:name>
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<mods:name id="A9C736EBA11D53239BA6D534CB618DA2" type="personal">
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<mods:roleTerm id="C52F08255B0EB3E29E16217870AC69C6">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FBDC9F2E93A29935150E9C433FFE3F5E">Fernicola, Juan Carlos</mods:namePart>
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</mods:name>
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<mods:name id="1B183BEBA1FD41CC949401C7916BC908" type="personal">
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<mods:roleTerm id="6776418B5954C86944171D9C4F73D882">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="8DBC7B0843AB49D22685F861FBE9C7AE">Cerdeño, Esperanza</mods:namePart>
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</mods:name>
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<mods:name id="F52B94048DC47505241884F9E4B84492" type="personal">
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<mods:roleTerm id="4FC2A3C797657FF8B092D59233B306E4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F2B2F18701B5DCCAF342B0DBB3F1525F">Reguero, Marcelo A.</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="161A006247F1F26E78363A1291311545">text</mods:typeOfResource>
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<mods:title id="4E8C3F0705D5F116D0EA2D9F5A1E2652">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="00F07B3D5698390DB3DB652F7A3F205F">
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<mods:date id="4BED3CF939F1B0472CB7B027D62031D8">2018</mods:date>
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<mods:number id="C9E444B4E51860775FDB35C44510E454">2018-02-28</mods:number>
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<mods:detail id="F116B7FFFCD923478EECD384A2DEEA69" type="volume">
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<mods:number id="3C5CF191718FB43526543929006AF2F7">4387</mods:number>
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<mods:detail id="F36B687E59247204E6BF0098CE0284C4" type="issue">
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<mods:number id="E749876D23B99E24C8F8BA976BD8DEF6">3</mods:number>
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<mods:identifier id="7EA102527570A2371542F8F6CF1C1F37" type="DOI">10.11646/zootaxa.4387.3.4</mods:identifier>
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<treatment id="03EC879BCF50831DC1922EBB8F42FA4A" ID-DOI="http://doi.org/10.5281/zenodo.5986677" ID-GBIF-Taxon="141264036" ID-Zenodo-Dep="5986677" LSID="urn:lsid:plazi:treatment:03EC879BCF50831DC1922EBB8F42FA4A" httpUri="http://treatment.plazi.org/id/03EC879BCF50831DC1922EBB8F42FA4A" lastPageId="7" lastPageNumber="480" pageId="3" pageNumber="476">
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<subSubSection id="C35F6506CF508319C1922EBB8E9EF8A3" pageId="3" pageNumber="476" type="nomenclature">
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<paragraph id="8BFA368DCF508319C1922EBB8CE3F8D8" blockId="3.[151,675,1778,1805]" box="[151,675,1778,1805]" pageId="3" pageNumber="476">
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<heading id="D0B281E1CF508319C1922EBB8CE3F8D8" bold="true" box="[151,675,1778,1805]" fontSize="11" level="1" pageId="3" pageNumber="476" reason="1">
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<taxonomicName id="4C454D0ECF508319C1922EBB8CE3F8D8" authority="Ameghino, 1885" authorityName="Ameghino" authorityYear="1885" box="[151,675,1778,1805]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="3" pageNumber="476" phylum="Chordata" rank="species" species="antiquum">
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<emphasis id="B931EA9FCF508319C1922EBB8CE3F8D8" bold="true" box="[151,675,1778,1805]" pageId="3" pageNumber="476">
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<emphasis id="B931EA9FCF508319C1922EBB8F92F8D9" bold="true" box="[151,466,1778,1804]" italics="true" pageId="3" pageNumber="476">Protypotherium antiquum</emphasis>
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<bibRefCitation id="EFD44B7CCF508319C0DC2EBA8CE3F8D8" box="[473,675,1779,1805]" pageId="3" pageNumber="476" refId="ref16851" refString="Ameghino, F. (1885) Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba, 8, 5 - 207." type="journal article">Ameghino, 1885</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8BFA368DCF508319C1922F708E9EF8A3" blockId="3.[151,1436,1849,1982]" pageId="3" pageNumber="476">
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<emphasis id="B931EA9FCF508319C1922F708FB5F887" bold="true" box="[151,501,1849,1874]" pageId="3" pageNumber="476">
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<typeStatus id="54FE882FCF508319C1922F708F4DF887" box="[151,269,1849,1874]" pageId="3" pageNumber="476" type="lectotype">Lectotype</typeStatus>
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(this contribution).
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</emphasis>
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ZMK 21/1877 (ex ZMK 208;
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<figureCitation id="137E2A08CF508319C24F2F708D95F887" box="[842,981,1849,1874]" captionStart="FIGURE 1" captionStartId="4.[151,249,1847,1869]" captionTargetBox="[226,1357,202,1822]" captionTargetId="figure@4.[220,1366,193,1837]" captionTargetPageId="4" captionText="FIGURE 1. Protypotherium antiquum, lectotype ZMK 21/1877, right mandibular fragment with p4-m3, in labial (A) and occlusal (B) views; reproductions of figures 1 (C) and 1a (D) from Ameghino (1889, plate 15); and MACN-A 1288, cast of lectotype, inlabial (E) and occlusal(F)views. Protypotherium australe, MACN-A 1338, cast of theholotype, in occlusalview (G). Protypotherium praerutilum, holotype MACN-A 1081, incomplete skull, in ventral (H) and dorsal (I) views; holotype MACN-A 1082, incomplete mandible, in lateral (J) and occlusal (K) views; reproductions of figures 6 (L) and 6a (M) from Ameghino (1889, plate 14); reproductions of figures 7 (N) and 7a (O) from Ameghino (1889, plate 14). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187699/files/figure.png" pageId="3" pageNumber="476">Figs. 1 A–B</figureCitation>
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), right mandibular fragment with p4–m3 series.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF508319C1C22FC88EAFF86B" pageId="3" pageNumber="476" type="description">
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<paragraph id="8BFA368DCF508319C1C22FC88EAFF86B" blockId="3.[151,1436,1849,1982]" pageId="3" pageNumber="476">
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<emphasis id="B931EA9FCF508319C1C22FC88C1CF84F" bold="true" box="[199,604,1921,1946]" pageId="3" pageNumber="476">
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<typeStatus id="54FE882FCF508319C1C22FC88F2CF84F" box="[199,364,1921,1946]" pageId="3" pageNumber="476" type="paralectotype">Paralectotype</typeStatus>
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(this contribution).
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</emphasis>
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A toothless fragmented mandible with its symphysis that has not been located.
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</paragraph>
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</subSubSection>
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<caption id="DF3A6605CF57831EC1922F7E8B3DF833" httpUri="https://zenodo.org/record/1187699/files/figure.png" pageId="4" pageNumber="477" startId="4.[151,249,1847,1869]" targetBox="[226,1357,202,1822]" targetPageId="4">
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<paragraph id="8BFA368DCF57831EC1922F7E8B3DF833" blockId="4.[151,1436,1846,2022]" pageId="4" pageNumber="477">
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<emphasis id="B931EA9FCF57831EC1922F7E8C71F899" bold="true" box="[151,561,1846,1869]" pageId="4" pageNumber="477">
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FIGURE 1.
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<taxonomicName id="4C454D0ECF57831EC01B2F7F8C71F899" authorityName="Ameghino" authorityYear="1885" box="[286,561,1846,1868]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="antiquum">
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<emphasis id="B931EA9FCF57831EC01B2F7F8C71F899" bold="true" box="[286,561,1846,1868]" italics="true" pageId="4" pageNumber="477">Protypotherium antiquum</emphasis>
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</taxonomicName>
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</emphasis>
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, lectotype ZMK 21/1877, right mandibular fragment with p4-m3, in labial
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<emphasis id="B931EA9FCF57831EC44E2F7E8B2EF898" bold="true" box="[1355,1390,1847,1869]" pageId="4" pageNumber="477">(A)</emphasis>
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and occlusal
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<emphasis id="B931EA9FCF57831EC1F72F1C8F54F8BE" bold="true" box="[242,276,1877,1899]" pageId="4" pageNumber="477">(B)</emphasis>
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views; reproductions of figures 1
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<emphasis id="B931EA9FCF57831EC37C2F1C8CDCF8BE" bold="true" box="[633,668,1877,1899]" pageId="4" pageNumber="477">(C)</emphasis>
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and 1a
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<emphasis id="B931EA9FCF57831EC3F42F1C8D54F8BE" bold="true" box="[753,788,1877,1899]" pageId="4" pageNumber="477">(D)</emphasis>
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from Ameghino (1889, plate 15); and MACN-A 1288, cast of lectotype, in labial
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<emphasis id="B931EA9FCF57831EC0522F3D8F39F85F" bold="true" box="[343,377,1908,1930]" pageId="4" pageNumber="477">(E)</emphasis>
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and occlusal
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<emphasis id="B931EA9FCF57831EC3072F3D8C63F85F" bold="true" box="[514,547,1908,1930]" pageId="4" pageNumber="477">(F)</emphasis>
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views.
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<taxonomicName id="4C454D0ECF57831EC3752F3D8D2EF85F" baseAuthorityName="Ameghino" baseAuthorityYear="1887" box="[624,878,1908,1930]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="australe">
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<emphasis id="B931EA9FCF57831EC3752F3D8D2EF85F" bold="true" box="[624,878,1908,1930]" italics="true" pageId="4" pageNumber="477">Protypotherium australe</emphasis>
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</taxonomicName>
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, MACN-A 1338, cast of the holotype, in occlusal view
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<emphasis id="B931EA9FCF57831EC1922FDA8EFCF87C" bold="true" box="[151,188,1939,1961]" pageId="4" pageNumber="477">(G)</emphasis>
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.
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<taxonomicName id="4C454D0ECF57831EC1CF2FDA8FB4F87C" authorityName="Ameghino" authorityYear="1887" box="[202,500,1939,1961]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="4" pageNumber="477" phylum="Chordata" rank="species" species="praerutilum">
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<emphasis id="B931EA9FCF57831EC1CF2FDA8FB4F87C" bold="true" box="[202,500,1939,1961]" italics="true" pageId="4" pageNumber="477">Protypotherium praerutilum</emphasis>
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</taxonomicName>
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, holotype MACN-A 1081, incomplete skull, in ventral
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<emphasis id="B931EA9FCF57831EC5332FDA8A1BF87C" bold="true" box="[1078,1115,1939,1961]" pageId="4" pageNumber="477">(H)</emphasis>
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and dorsal
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<emphasis id="B931EA9FCF57831EC5D32FDA8AB1F87C" bold="true" box="[1238,1265,1939,1961]" pageId="4" pageNumber="477">(I)</emphasis>
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views; holotype MACN-A 1082, incomplete mandible, in lateral
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<emphasis id="B931EA9FCF57831EC3882FFB8CEBF81D" bold="true" box="[653,683,1970,1992]" pageId="4" pageNumber="477">(J)</emphasis>
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and occlusal
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<emphasis id="B931EA9FCF57831EC23E2FFB8D20F81D" bold="true" box="[827,864,1970,1992]" pageId="4" pageNumber="477">(K)</emphasis>
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views; reproductions of figures 6
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<emphasis id="B931EA9FCF57831EC5C12FFB8AA6F81D" bold="true" box="[1220,1254,1970,1992]" pageId="4" pageNumber="477">(L)</emphasis>
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and 6a
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<emphasis id="B931EA9FCF57831EC43F2FFB8B22F81D" bold="true" box="[1338,1378,1970,1992]" pageId="4" pageNumber="477">(M)</emphasis>
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from Ameghino (1889, plate 14); reproductions of figures 7
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<emphasis id="B931EA9FCF57831EC3C62F998CA5F833" bold="true" box="[707,741,2000,2022]" pageId="4" pageNumber="477">(N)</emphasis>
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and 7a
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<emphasis id="B931EA9FCF57831EC2302F998D1AF833" bold="true" box="[821,858,2000,2022]" pageId="4" pageNumber="477">(O)</emphasis>
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from Ameghino (1889, plate 14). Scale bar = 10 mm.
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</paragraph>
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</caption>
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<caption id="DF3A6605CF56831FC1922E298DD8F833" httpUri="https://zenodo.org/record/1187701/files/figure.png" pageId="5" pageNumber="478" startId="5.[151,249,1632,1654]" targetBox="[308,1273,213,1595]" targetPageId="5">
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<paragraph id="8BFA368DCF56831FC1922E298DD8F833" blockId="5.[151,1436,1632,2022]" pageId="5" pageNumber="478">
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<emphasis id="B931EA9FCF56831FC1922E298C00F9A3" bold="true" box="[151,576,1632,1654]" pageId="5" pageNumber="478">
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FIGURE 2.
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<taxonomicName id="4C454D0ECF56831FC0192E298C00F9A3" authorityName="Ameghino" authorityYear="1887" box="[284,576,1632,1654]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="attenuatum">
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<emphasis id="B931EA9FCF56831FC0192E298C00F9A3" bold="true" box="[284,576,1632,1654]" italics="true" pageId="5" pageNumber="478">Protypotherium attenuatum</emphasis>
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</taxonomicName>
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</emphasis>
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, syntype MACN-A 524, left maxilla with P2–M3 but lacking P1, in occlusal view
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<emphasis id="B931EA9FCF56831FC1922E368EFAF940" bold="true" box="[151,186,1663,1685]" pageId="5" pageNumber="478">(A)</emphasis>
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; syntype MACN-A 628, right fragmented mandible with alveoli of i1–3 and c (broken)–m1 but lacking m2–3, in labial
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<emphasis id="B931EA9FCF56831FC47F2E368BDCF940" bold="true" box="[1402,1436,1663,1685]" pageId="5" pageNumber="478">(B)</emphasis>
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and occlusal
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<emphasis id="B931EA9FCF56831FC0192ED78F7FF961" bold="true" box="[284,319,1694,1716]" pageId="5" pageNumber="478">(C)</emphasis>
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views; reproductions of figures 21
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<emphasis id="B931EA9FCF56831FC3AC2ED78C8CF961" bold="true" box="[681,716,1694,1716]" pageId="5" pageNumber="478">(D)</emphasis>
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, 20
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<emphasis id="B931EA9FCF56831FC3FC2ED78D5BF961" bold="true" box="[761,795,1694,1716]" pageId="5" pageNumber="478">(E)</emphasis>
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and 20a
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<emphasis id="B931EA9FCF56831FC27D2ED78DD9F961" bold="true" box="[888,921,1694,1716]" pageId="5" pageNumber="478">(F)</emphasis>
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from Ameghino (1889, plate 14).
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<taxonomicName id="4C454D0ECF56831FC5F22ED48F4DF907" authorityName="Ameghino" authorityYear="1888" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="obstructum">
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<emphasis id="B931EA9FCF56831FC5F22ED48F4DF907" bold="true" italics="true" pageId="5" pageNumber="478">Protypotherium obstructum</emphasis>
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</taxonomicName>
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, holotype MACN-A 1677, lower premolar in labial, oclusal and lingual views
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<emphasis id="B931EA9FCF56831FC53F2EF58A1FF907" bold="true" box="[1082,1119,1724,1746]" pageId="5" pageNumber="478">(G)</emphasis>
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; reproduction of figure 19
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<emphasis id="B931EA9FCF56831FC4722EF58BDCF907" bold="true" box="[1399,1436,1724,1746]" pageId="5" pageNumber="478">(H)</emphasis>
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from Ameghino (1889, plate 15).
|
||
<taxonomicName id="4C454D0ECF56831FC0EA2E928CB5F924" authorityName="Ameghino" authorityYear="1889" box="[495,757,1755,1777]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="claudum">
|
||
<emphasis id="B931EA9FCF56831FC0EA2E928CB5F924" bold="true" box="[495,757,1755,1777]" italics="true" pageId="5" pageNumber="478">Protypotherium claudum</emphasis>
|
||
</taxonomicName>
|
||
, holotype MACN-A 551, right mandibular fragment with alveolus of p2 and p3–m2, in labial
|
||
<emphasis id="B931EA9FCF56831FC0AD2EB38F83F8C5" bold="true" box="[424,451,1786,1808]" pageId="5" pageNumber="478">(I)</emphasis>
|
||
and occlusal
|
||
<emphasis id="B931EA9FCF56831FC3482EB38C2BF8C5" bold="true" box="[589,619,1786,1808]" pageId="5" pageNumber="478">(J)</emphasis>
|
||
views; reproductions of figures 22
|
||
<emphasis id="B931EA9FCF56831FC2D72EB38DB7F8C5" bold="true" box="[978,1015,1786,1808]" pageId="5" pageNumber="478">(K)</emphasis>
|
||
and 22a
|
||
<emphasis id="B931EA9FCF56831FC5572EB38A34F8C5" bold="true" box="[1106,1140,1786,1808]" pageId="5" pageNumber="478">(L)</emphasis>
|
||
from Ameghino (1889, plate 14).
|
||
<taxonomicName id="4C454D0ECF56831FC1CE2F518FA1F8FB" authorityName="Ameghino" authorityYear="1891" box="[203,481,1816,1838]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="globosum">
|
||
<emphasis id="B931EA9FCF56831FC1CE2F518FA1F8FB" bold="true" box="[203,481,1816,1838]" italics="true" pageId="5" pageNumber="478">Protypotherium globosum</emphasis>
|
||
</taxonomicName>
|
||
, holotype MACN-A 4049, skull, in ventral
|
||
<emphasis id="B931EA9FCF56831FC2CC2F518DB2F8FB" bold="true" box="[969,1010,1816,1838]" pageId="5" pageNumber="478">(M)</emphasis>
|
||
and dorsal
|
||
<emphasis id="B931EA9FCF56831FC57F2F518ADCF8FB" bold="true" box="[1146,1180,1816,1838]" pageId="5" pageNumber="478">(N)</emphasis>
|
||
views.
|
||
<taxonomicName id="4C454D0ECF56831FC5F22F518F57F898" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="convexidens">
|
||
<emphasis id="B931EA9FCF56831FC5F22F518F57F898" bold="true" italics="true" pageId="5" pageNumber="478">Protypotherium convexidens</emphasis>
|
||
</taxonomicName>
|
||
, holotype MACN-A 4054, right maxillary fragment with P3–M2 (broken), in occlusal view
|
||
<emphasis id="B931EA9FCF56831FC5C32F7E8AABF898" bold="true" box="[1222,1259,1847,1869]" pageId="5" pageNumber="478">(O)</emphasis>
|
||
.
|
||
<taxonomicName id="4C454D0ECF56831FC5F22F7E8F4CF8BE" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="diversidens">
|
||
<emphasis id="B931EA9FCF56831FC5F22F7E8F4CF8BE" bold="true" italics="true" pageId="5" pageNumber="478">Protypotherium diversidens</emphasis>
|
||
</taxonomicName>
|
||
, holotype MACN-A 4052, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view
|
||
<emphasis id="B931EA9FCF56831FC4702F1F8BD6F8B9" bold="true" box="[1397,1430,1878,1900]" pageId="5" pageNumber="478">(P)</emphasis>
|
||
.
|
||
<taxonomicName id="4C454D0ECF56831FC1922F3D8F99F85F" authorityName="Ameghino" authorityYear="1891" box="[151,473,1908,1930]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="compressidens">
|
||
<emphasis id="B931EA9FCF56831FC1922F3D8F99F85F" bold="true" box="[151,473,1908,1930]" italics="true" pageId="5" pageNumber="478">Protypotherium compressidens</emphasis>
|
||
</taxonomicName>
|
||
, holotype MACN-A 4029, left maxillary fragment with P3–M3, in occlusal view
|
||
<emphasis id="B931EA9FCF56831FC4172F3D8B77F85F" bold="true" box="[1298,1335,1908,1930]" pageId="5" pageNumber="478">(Q)</emphasis>
|
||
; holotype MACN-A 4030, left mandibular fragment with p4 (talonid)–m3, in occlusal view
|
||
<emphasis id="B931EA9FCF56831FC5002FDA8A68F87C" bold="true" box="[1029,1064,1939,1961]" pageId="5" pageNumber="478">(R)</emphasis>
|
||
.
|
||
<taxonomicName id="4C454D0ECF56831FC53F2FDA8B72F87C" authorityName="Ameghino" authorityYear="1894" box="[1082,1330,1939,1961]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="5" pageNumber="478" phylum="Chordata" rank="species" species="lineare">
|
||
<emphasis id="B931EA9FCF56831FC53F2FDA8B72F87C" bold="true" box="[1082,1330,1939,1961]" italics="true" pageId="5" pageNumber="478">Protypotherium lineare</emphasis>
|
||
</taxonomicName>
|
||
, holotype MACN-A 4039, right mandibular fragment with alveolus of p1 and p2–m1, in occlusal view
|
||
<emphasis id="B931EA9FCF56831FC5412FFB8A22F81D" bold="true" box="[1092,1122,1970,1992]" pageId="5" pageNumber="478">(S)</emphasis>
|
||
; holotype MACN-A 4038, left mandibular fragment with p3–m2, in occlusal view
|
||
<emphasis id="B931EA9FCF56831FC3A62F998C85F833" bold="true" box="[675,709,2000,2022]" pageId="5" pageNumber="478">(T)</emphasis>
|
||
. Scale bar = 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C35F6506CF55831CC1C228DE8DF2F935" pageId="6" pageNumber="479" type="diagnosis">
|
||
<paragraph id="8BFA368DCF55831CC1C228DE8AE8FC39" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="479">
|
||
<emphasis id="B931EA9FCF55831CC1C228DE8F7AFF65" bold="true" box="[199,314,151,176]" pageId="6" pageNumber="479">Diagnosis</emphasis>
|
||
(
|
||
<bibRefCitation id="EFD44B7CCF55831CC04C28DE8C20FF65" box="[329,608,151,176]" pageId="6" pageNumber="479" refId="ref16851" refString="Ameghino, F. (1885) Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba, 8, 5 - 207." type="journal article">Ameghino, 1885: 81–83</bibRefCitation>
|
||
). “[…]
|
||
<emphasis id="B931EA9FCF55831CC3B328DE8AFEFF01" italics="true" pageId="6" pageNumber="479">El fragmento de sínfisis de la mandíbula inferior que me sirvió de base para la fundación del género, es ancho y aplastado en sentido vertical, muy parecido a</emphasis>
|
||
<taxonomicName id="4C454D0ECF55831CC5C328F58B16FF00" box="[1222,1366,188,213]" class="Mammalia" family="Mesotheriidae" genus="Typotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">Typotherium</taxonomicName>
|
||
<emphasis id="B931EA9FCF55831CC46528F48A0EFDCD" italics="true" pageId="6" pageNumber="479">en su parte cóncava interna, pero más aplastado en su parte inferior y con una depresión semilunar en la parte posterior de la sínfisis. El foramen mentale, situado en la parte sinfisaria, es un agujero elíptico, de unos 6 milímetros de diámetro mayor. Las dos ramas de la mandíbula están tan íntimamente unidas formando un solo hueso que en ninguna parte se ve ni el más mínimo rastro de sutura. La parte anterior está rota, pero todavía se distinguen en ella seis alvéolos horizontales en los que sin duda se implantaban seis incisivos. Al nivel del agujero mental, tiene la sínfisis 20 milímetros de ancho y 14 milímetros de alto. La pieza del señor Roth es un pedazo del lado derecho de la mandíbula inferior, con las cuatro últimas muelas y parte del alvéolo de otro molar anterior, de modo que el número de muelas era, por lo menos, de cinco en cada lado de la mandíbula inferior. Estas muelas eran sin raíces separadas, abiertas en la base y construidas sobre el mismo tipo general que las de</emphasis>
|
||
<taxonomicName id="4C454D0ECF55831CC55029B68AF8FDCD" authorityName="Owen" authorityYear="1837" box="[1109,1208,511,536]" class="Mammalia" family="Toxodontidae" genus="Toxodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">Toxodon</taxonomicName>
|
||
<emphasis id="B931EA9FCF55831CC5C52A488A8CFDCD" box="[1216,1228,513,536]" italics="true" pageId="6" pageNumber="479">y</emphasis>
|
||
<taxonomicName id="4C454D0ECF55831CC5D129B68B24FDCD" box="[1236,1380,511,536]" class="Mammalia" family="Mesotheriidae" genus="Typotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">Typotherium</taxonomicName>
|
||
,
|
||
<emphasis id="B931EA9FCF55831CC4712A488C43FD8A" italics="true" pageId="6" pageNumber="479">con una capa de esmalte, que parece haber sido continua en algunas muelas, o a lo menos con no tantas interrupciones como en las muelas del género</emphasis>
|
||
<taxonomicName id="4C454D0ECF55831CC30B2A0E8C2FFDB5" authorityName="Owen" authorityYear="1837" box="[526,623,583,608]" class="Mammalia" family="Toxodontidae" genus="Toxodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">Toxodon</taxonomicName>
|
||
<emphasis id="B931EA9FCF55831CC3792A008A3AFC12" italics="true" pageId="6" pageNumber="479">y todas en serie continua muy apretadas entre sí. La primera muela existente, que corresponde al cuarto y último premolar, si la dentición fuera completa y normal, es más pequeña que las otras, dividida en dos partes desiguales, una anterior más grande y otra posterior bastante más pequeña; esta división es naturalmente producida por dos surcos perpendiculares opuestos, uno colocado sobre el lado interno y el otro sobre la esquina posteroexterna. Las dos muelas que siguen, primero y segundo verdadero molar, están divididas en dos partes o lóbulos más iguales, por dos surcos opuestos, uno interno, poco marcado y acompañado de una pequeña columna formada por la parte posterointerna del primer lóbulo, y el otro externo, más profundo y que forma en la corona un pliegue entrante parecido al que presentan las muelas de los caballos y varios otros mamíferos de órdenes distintos. La última muela, algo más grande que las demás, está dividida en tres partes o lóbulos formados por tres columnas en cada lado (externo e interno), separados por dos surcos. La corona de las muelas está bastante gastada y excavada en el centro, como las del género</emphasis>
|
||
<taxonomicName id="4C454D0ECF55831CC5852BE68B54FC1D" box="[1152,1300,943,968]" class="Mammalia" family="Mesotheriidae" genus="Typotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">Typotherium</taxonomicName>
|
||
.
|
||
<emphasis id="B931EA9FCF55831CC41B2BF88A21FC38" italics="true" pageId="6" pageNumber="479">Parece que la rama ascendente empezaba a levantarse inmediatamente detrás de la última muela.</emphasis>
|
||
[…]”.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF55831CC1C22BBE8DF2F935" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="479">
|
||
English translation. The fragment of mandibular symphysis used to found the genus is wide and vertically compressed, much similar to
|
||
<taxonomicName id="4C454D0ECF55831CC0E02C558C32FBE0" box="[485,626,1052,1077]" class="Mammalia" family="Mesotheriidae" genus="Typotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EA9FCF55831CC0E02C558C32FBE0" box="[485,626,1052,1077]" italics="true" pageId="6" pageNumber="479">Typotherium</emphasis>
|
||
</taxonomicName>
|
||
in its internal concave face, but more compressed at its lower part and with a semilunate depression on its posterior end. The mental foramen is an elliptical opening of about
|
||
<quantity id="4CBD9B68CF55831CC43C2C768B3CFB82" box="[1337,1404,1087,1112]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="6" pageNumber="479" unit="mm" value="6.0">6 mm</quantity>
|
||
of diameter located in the symphyseal region. Both rami of the jaw are intimately fused into a single bone, without any trace of suture. The anterior part is broken, but there are six horizontal alveoli undoubtedly corresponding to six incisors. At the level of the mental foramen, the symphysis is
|
||
<quantity id="4CBD9B68CF55831CC2772CE58D81FB11" box="[882,961,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="6" pageNumber="479" unit="mm" value="20.0">20 mm</quantity>
|
||
wide and
|
||
<quantity id="4CBD9B68CF55831CC5312CE58AC3FB11" box="[1076,1155,1196,1220]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="6" pageNumber="479" unit="mm" value="14.0">14 mm</quantity>
|
||
high. Roth’s specimen is a right mandibular ramus, with the last four cheek teeth and the broken alveolus of another anterior one, so that the number of cheek teeth was, at least, five on each side of the mandible. These teeth did not have separated roots, were open at the base and constructed on the same general way as those of
|
||
<taxonomicName id="4C454D0ECF55831CC2E02D5E8A02FAE5" authorityName="Owen" authorityYear="1837" box="[997,1090,1303,1328]" class="Mammalia" family="Toxodontidae" genus="Toxodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EA9FCF55831CC2E02D5E8A02FAE5" box="[997,1090,1303,1328]" italics="true" pageId="6" pageNumber="479">Toxodon</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C454D0ECF55831CC5792D5E8B49FAE5" box="[1148,1289,1303,1328]" class="Mammalia" family="Mesotheriidae" genus="Typotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EA9FCF55831CC5792D5E8B49FAE5" box="[1148,1289,1303,1328]" italics="true" pageId="6" pageNumber="479">Typotherium</emphasis>
|
||
</taxonomicName>
|
||
, with a layer of enamel that seems to have been continuous in some cheek teeth, or at least with less interruptions than in those of the genus
|
||
<taxonomicName id="4C454D0ECF55831CC0312D168FD1FAAD" authorityName="Owen" authorityYear="1837" box="[308,401,1375,1400]" class="Mammalia" family="Toxodontidae" genus="Toxodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EA9FCF55831CC0312D168FD1FAAD" box="[308,401,1375,1400]" italics="true" pageId="6" pageNumber="479">Toxodon</emphasis>
|
||
</taxonomicName>
|
||
, and all implanted in a very tight, continuous series. The first cheek tooth, assuming a complete and normal dentition, would correspond to the fourth and last premolar; it is smaller than the molars, divided into two unequal parts, the anterior being rather larger than the posterior; this division is naturally produced by two opposite perpendicular grooves, one placed on the inner side and the other on the posteroexternal region. The first and second true molars are divided into two more equal parts or lobes by two opposite grooves, the internal is little marked and accompanied by a small column formed by the posterointernal part of the first lobe, and the external groove is deeper and forms an infolding in the crown similar to that of the cheek teeth of horses and several other mammals of different orders. The last molar, somewhat larger than the others, is divided into three parts or lobes formed by three columns at each side (external and internal), separated by two grooves. The crown of the molars is rather worn and excavated in the centre, such as those of the genus
|
||
<taxonomicName id="4C454D0ECF55831CC5A72EED8B6FF968" box="[1186,1327,1700,1725]" class="Mammalia" family="Mesotheriidae" genus="Typotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EA9FCF55831CC5A72EED8B6FF968" box="[1186,1327,1700,1725]" italics="true" pageId="6" pageNumber="479">Typotherium</emphasis>
|
||
</taxonomicName>
|
||
. It seems that the ascending ramus was rising immediately behind the last molar.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35F6506CF55831DC1C22EA58F42FA4A" lastPageId="7" lastPageNumber="480" pageId="6" pageNumber="479" type="discussion">
|
||
<paragraph id="8BFA368DCF55831DC1C22EA58C28FF2D" blockId="6.[151,1437,151,2013]" lastBlockId="7.[151,1437,151,1439]" lastPageId="7" lastPageNumber="480" pageId="6" pageNumber="479">
|
||
<emphasis id="B931EA9FCF55831CC1C22EA58F0FF8D0" bold="true" box="[199,335,1772,1797]" pageId="6" pageNumber="479">Comments.</emphasis>
|
||
<bibRefCitation id="EFD44B7CCF55831CC05C2EA58C3BF8D1" box="[345,635,1772,1797]" pageId="6" pageNumber="479" refId="ref16851" refString="Ameghino, F. (1885) Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba, 8, 5 - 207." type="journal article">Ameghino (1885: 81–83)</bibRefCitation>
|
||
described
|
||
<taxonomicName id="4C454D0ECF55831CC3FF2EA58A5EF8D1" authorityName="Ameghino" authorityYear="1885" box="[762,1054,1772,1797]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="6" pageNumber="479" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF55831CC3FF2EA58A5EF8D1" box="[762,1054,1772,1797]" italics="true" pageId="6" pageNumber="479">Protypotherium antiquum</emphasis>
|
||
</taxonomicName>
|
||
based on two partial mandibular fragments. One of them is a rolled and shattered, toothless fragmented mandible with its symphysis, which
|
||
<bibRefCitation id="EFD44B7CCF55831CC1922F7D8FCCF899" box="[151,396,1844,1869]" pageId="6" pageNumber="479" refId="ref16851" refString="Ameghino, F. (1885) Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba, 8, 5 - 207." type="journal article">Ameghino (1885: 83)</bibRefCitation>
|
||
recognised to have been used to name the genus and species in a faunal list of a collection catalogue in 1882. However, as already mentioned (see Historical Background), those names are
|
||
<emphasis id="B931EA9FCF55831CC5ED2F108B3CF8A5" box="[1256,1404,1879,1904]" italics="true" pageId="6" pageNumber="479">nomina nuda</emphasis>
|
||
as acknowledged by him years later (e.g.,
|
||
<bibRefCitation id="EFD44B7CCF55831CC34B2F358D5AF841" box="[590,794,1916,1941]" pageId="6" pageNumber="479" refId="ref16898" refString="Ameghino, F. (1887 a) Observaciones generales sobre el orden de mamiferos extinguidos sub-americanos llamados Toxodontes (Toxodontia) y sinopsis de los generos y especies hasta ahora conocidos. Anales del Museo de La Plata, 1 (1), 1 - 66." type="journal article">Ameghino, 1887a</bibRefCitation>
|
||
). The other specimen was collected in the cliffs of Paraná River (
|
||
<collectingRegion id="4981F86FCF55831CC1ED2FE98F97F86C" box="[232,471,1952,1977]" country="Argentina" name="Entre Rios" pageId="6" pageNumber="479">Entre Ríos Province</collectingRegion>
|
||
) by
|
||
<collectingRegion id="4981F86FCF55831CC3172FE98C34F86C" box="[530,628,1952,1977]" country="Philippines" name="Santiago" pageId="6" pageNumber="479">Santiago</collectingRegion>
|
||
Roth, probably during an excursion in 1881, and later borrowed by Ameghino for its study and the description of all the dental characteristics of the species (
|
||
<bibRefCitation id="EFD44B7CCF55831CC57D2F8D8B72F808" box="[1144,1330,1988,2013]" pageId="6" pageNumber="479" refId="ref16851" refString="Ameghino, F. (1885) Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba, 8, 5 - 207." type="journal article">Ameghino 1885</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD44B7CCF55831CC4392F8D8B38F809" box="[1340,1400,1988,2012]" pageId="6" pageNumber="479" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">1889</bibRefCitation>
|
||
). It involves a right mandibular ramus with broken alveolus of p3 and complete p4–m3. As the nomenclatural act that made available the name
|
||
<taxonomicName id="4C454D0ECF54831DC0B028F48C7BFF01" authorityName="Ameghino" authorityYear="1885" box="[437,571,189,212]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC0B028F48C7BFF01" box="[437,571,189,212]" italics="true" pageId="7" pageNumber="480">P. antiquum</emphasis>
|
||
</taxonomicName>
|
||
took place in 1885 when Ameghino described both specimens, they are here recognised as the
|
||
<typeStatus id="54FE882FCF54831DC05A28A98F81FF2D" box="[351,449,224,248]" pageId="7" pageNumber="480" type="syntype">syntypes</typeStatus>
|
||
of the species.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF54831DC1C2294D8DE7FE20" blockId="7.[151,1437,151,1439]" pageId="7" pageNumber="480">
|
||
There are no comments on the
|
||
<typeStatus id="54FE882FCF54831DC335294C8C20FEC8" box="[560,608,261,285]" pageId="7" pageNumber="480">type</typeStatus>
|
||
material of
|
||
<taxonomicName id="4C454D0ECF54831DC3F5294C8D38FEC9" authorityName="Ameghino" authorityYear="1885" box="[752,888,261,284]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC3F5294C8D38FEC9" box="[752,888,261,284]" italics="true" pageId="7" pageNumber="480">P. antiquum</emphasis>
|
||
</taxonomicName>
|
||
in Ameghino’s catalogue and neither in
|
||
<bibRefCitation id="EFD44B7CCF54831DC455294C8EA4FE95" pageId="7" pageNumber="480" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
|
||
. Regarding the toothless symphysis, it should be housed at the MLP, because this institution bought Ameghino’s collection in 1886, unless Ameghino had deviated the specimen to his posterior personal collection; in such a case, it should be in MACN-A. However, this
|
||
<typeStatus id="54FE882FCF54831DC3EA29398D06FE5D" box="[751,838,368,392]" pageId="7" pageNumber="480" type="syntype">syntype</typeStatus>
|
||
could not be located in any of these institutions or in ZMK and, unfortunately, it has not been identified in any of the available photographs taken from AMNH, FM and YPM. Nevertheless, we cannot reject the hypothesis that it could be located in another foreigner institution (see
|
||
<bibRefCitation id="EFD44B7CCF54831DC19229958F3CFE21" box="[151,380,476,501]" pageId="7" pageNumber="480" refId="ref18790" refString="Vizcaino, S. F., Bargo, M. S. & Fernicola, J. C. (2013) Expediciones paleontologicas durante los Siglos XIX y XX a la Formacion Santa Cruz (Mioceno Inferior, Patagonia) y destino de los fosiles. Actas del 3 er Congreso Argentino de Historia de la Geologia, Salta, 2013, 1 - 15." type="journal article">
|
||
Vizcaíno
|
||
<emphasis id="B931EA9FCF54831DC00429948F71FE20" box="[257,305,476,501]" italics="true" pageId="7" pageNumber="480">et al</emphasis>
|
||
. 2013
|
||
</bibRefCitation>
|
||
); further searching will corroborate or refute this.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF54831DC1C22A498D87FC57" blockId="7.[151,1437,151,1439]" pageId="7" pageNumber="480">
|
||
In the case of Roth’s specimen, it was further described by
|
||
<bibRefCitation id="EFD44B7CCF54831DC25929B68A69FDCD" box="[860,1065,511,536]" pageId="7" pageNumber="480" refId="ref16898" refString="Ameghino, F. (1887 a) Observaciones generales sobre el orden de mamiferos extinguidos sub-americanos llamados Toxodontes (Toxodontia) y sinopsis de los generos y especies hasta ahora conocidos. Anales del Museo de La Plata, 1 (1), 1 - 66." type="journal article">Ameghino (1887a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="EFD44B7CCF54831DC5302A498A2EFDCD" box="[1077,1134,512,536]" pageId="7" pageNumber="480" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">1889</bibRefCitation>
|
||
) and even illustrated in his Atlas (
|
||
<bibRefCitation id="EFD44B7CCF54831DC1E12A6D8C15FDE8" box="[228,597,548,573]" pageId="7" pageNumber="480" refId="ref17032" refString="Ameghino, F. (1889) Contribucion al conocimiento de los mamiferos fosiles de la Republica Argentina. Actas de la Academia Nacional de Ciencias de Cordoba, 6, i - xxxii, 1 - 1027, atlas of 98 pls." type="journal article">Ameghino 1889, plate 15: fig.1</bibRefCitation>
|
||
); this illustration is herein reproduced in
|
||
<figureCitation id="137E2A08CF54831DC53E2A6D8AAAFDE9" box="[1083,1258,548,573]" captionStart="FIGURE 1" captionStartId="4.[151,249,1847,1869]" captionTargetBox="[226,1357,202,1822]" captionTargetId="figure@4.[220,1366,193,1837]" captionTargetPageId="4" captionText="FIGURE 1. Protypotherium antiquum, lectotype ZMK 21/1877, right mandibular fragment with p4-m3, in labial (A) and occlusal (B) views; reproductions of figures 1 (C) and 1a (D) from Ameghino (1889, plate 15); and MACN-A 1288, cast of lectotype, inlabial (E) and occlusal(F)views. Protypotherium australe, MACN-A 1338, cast of theholotype, in occlusalview (G). Protypotherium praerutilum, holotype MACN-A 1081, incomplete skull, in ventral (H) and dorsal (I) views; holotype MACN-A 1082, incomplete mandible, in lateral (J) and occlusal (K) views; reproductions of figures 6 (L) and 6a (M) from Ameghino (1889, plate 14); reproductions of figures 7 (N) and 7a (O) from Ameghino (1889, plate 14). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187699/files/figure.png" pageId="7" pageNumber="480">Figures 1 C–D</figureCitation>
|
||
. Roth kept this specimen until he sold it to Dr. V. Lausen, a wealthy Danish citizen settled in Buenos Aires, who sent it to the Zoological Museum, Quaternary Zoology Collections-V, Lausen Collection, University of Copenhagen in Denmark, being housed there since the 25th of November of 1887 (B. E. Kramer, pers. comm. to M. Fernández; M. Reguero’s pers. obs.). This specimen has a label handwritten by Roth with the following information: “
|
||
<taxonomicName id="4C454D0ECF54831DC1A62A938C8FFD26" authority="Amegh. Delta-Egnen" authorityName="Amegh. Delta-Egnen" box="[163,719,730,755]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC1A62A938F8BFD26" box="[163,459,730,755]" italics="true" pageId="7" pageNumber="480">Protypotherium antiquum</emphasis>
|
||
Amegh. Delta-Egnen
|
||
</taxonomicName>
|
||
, Entre Ríos. Original Roth (37.), Laussen.
|
||
<date id="FFFB104DCF54831DC5DE2A938B00FD26" box="[1243,1344,730,755]" pageId="7" pageNumber="480" value="87-11-25">25.11.87</date>
|
||
. 208*”. Original Roth (37.) refers to the number in Roth’s catalogue, and 208 refers to an older catalogue number of Museum of Zoology in Copenhagen (ZMK 208). However, nowadays this material is catalogued as ZMK 21/1887 (
|
||
<figureCitation id="137E2A08CF54831DC19A2B0F8F75FC8A" box="[159,309,838,863]" captionStart="FIGURE 1" captionStartId="4.[151,249,1847,1869]" captionTargetBox="[226,1357,202,1822]" captionTargetId="figure@4.[220,1366,193,1837]" captionTargetPageId="4" captionText="FIGURE 1. Protypotherium antiquum, lectotype ZMK 21/1877, right mandibular fragment with p4-m3, in labial (A) and occlusal (B) views; reproductions of figures 1 (C) and 1a (D) from Ameghino (1889, plate 15); and MACN-A 1288, cast of lectotype, inlabial (E) and occlusal(F)views. Protypotherium australe, MACN-A 1338, cast of theholotype, in occlusalview (G). Protypotherium praerutilum, holotype MACN-A 1081, incomplete skull, in ventral (H) and dorsal (I) views; holotype MACN-A 1082, incomplete mandible, in lateral (J) and occlusal (K) views; reproductions of figures 6 (L) and 6a (M) from Ameghino (1889, plate 14); reproductions of figures 7 (N) and 7a (O) from Ameghino (1889, plate 14). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187699/files/figure.png" pageId="7" pageNumber="480">Figs. 1 A–B</figureCitation>
|
||
). Two casts of ZMK 21/1877 were fortunately kept in MLP and MACN-A: MLP
|
||
<date id="FFFB104DCF54831DC5FB2B0E8BC0FC8A" box="[1278,1408,839,863]" pageId="7" pageNumber="480" value="1997-11-20" valueMax="1997-11-01" valueMin="1997-11-20">97-XI-20-1</date>
|
||
, a complete cast, and MACN-A 1288 (
|
||
<figureCitation id="137E2A08CF54831DC3282B238CF5FC56" box="[557,693,874,899]" captionStart="FIGURE 1" captionStartId="4.[151,249,1847,1869]" captionTargetBox="[226,1357,202,1822]" captionTargetId="figure@4.[220,1366,193,1837]" captionTargetPageId="4" captionText="FIGURE 1. Protypotherium antiquum, lectotype ZMK 21/1877, right mandibular fragment with p4-m3, in labial (A) and occlusal (B) views; reproductions of figures 1 (C) and 1a (D) from Ameghino (1889, plate 15); and MACN-A 1288, cast of lectotype, inlabial (E) and occlusal(F)views. Protypotherium australe, MACN-A 1338, cast of theholotype, in occlusalview (G). Protypotherium praerutilum, holotype MACN-A 1081, incomplete skull, in ventral (H) and dorsal (I) views; holotype MACN-A 1082, incomplete mandible, in lateral (J) and occlusal (K) views; reproductions of figures 6 (L) and 6a (M) from Ameghino (1889, plate 14); reproductions of figures 7 (N) and 7a (O) from Ameghino (1889, plate 14). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187699/files/figure.png" pageId="7" pageNumber="480">Figs. 1 E–F</figureCitation>
|
||
), a cast of the tooth row.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF54831DC1C22BC78EAEFA82" blockId="7.[151,1437,151,1439]" pageId="7" pageNumber="480">
|
||
Before this contribution, there were two mentions regarding the
|
||
<typeStatus id="54FE882FCF54831DC2A72BD98D92FC7D" box="[930,978,912,936]" pageId="7" pageNumber="480">type</typeStatus>
|
||
specimen of
|
||
<taxonomicName id="4C454D0ECF54831DC5692BD98AB3FC72" box="[1132,1267,912,935]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC5692BD98AB3FC72" box="[1132,1267,912,935]" italics="true" pageId="7" pageNumber="480">P. antiquum</emphasis>
|
||
</taxonomicName>
|
||
, with different interpretations:
|
||
<bibRefCitation id="EFD44B7CCF54831DC0492BFB8FBFFC1E" box="[332,511,946,971]" pageId="7" pageNumber="480" refId="ref18435" refString="Schmidt, G. I. (2013) Litopterna y Notoungulata (Mammalia) de la Formacion Ituzaingo (Mioceno Tardio-Plioceno) de la Provincia de Entre Rios: sistematica, bioestratigrafia y paleobiogeografia. Unpublished Ph. D. dissertation, Universidad Nacional de La Plata, La Plata, 336 pp." type="book">Schmidt (2013)</bibRefCitation>
|
||
considered the missing toothless symphysis to be the
|
||
<typeStatus id="54FE882FCF54831DC5822BFB8AACFC1E" box="[1159,1260,946,971]" pageId="7" pageNumber="480" type="holotype">holotype</typeStatus>
|
||
, whereas
|
||
<bibRefCitation id="EFD44B7CCF54831DC4672BFA8EA1FC3A" pageId="7" pageNumber="480" refId="ref18162" refString="Luna, D. A. (2015) Miocene ungulates from Laguna del Laja, Chile, and an assessment of the Laguna del Laja fauna. Unpublished Ph. D. dissertation, University of California, Santa Barbara, 287 pp." type="book">Luna (2015)</bibRefCitation>
|
||
stated the partial mandible ZMK 21/1887 as such. Although
|
||
<bibRefCitation id="EFD44B7CCF54831DC29D2B9F8A20FC3A" box="[920,1120,982,1007]" pageId="7" pageNumber="480" refId="ref16851" refString="Ameghino, F. (1885) Nuevos restos de mamiferos fosiles oligocenos, recogidos por el profesor Pedro Scalabrini y pertenecientes al Museo Provincial de la Ciudad del Parana. Boletin de la Academia Nacional de Ciencias de Cordoba, 8, 5 - 207." type="journal article">Ameghino (1885)</bibRefCitation>
|
||
recognised that the missing toothless symphysis was the object that he used to name
|
||
<taxonomicName id="4C454D0ECF54831DC22C2BB58DF0FBC6" box="[809,944,1020,1043]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC22C2BB58DF0FBC6" box="[809,944,1020,1043]" italics="true" pageId="7" pageNumber="480">P. antiquum</emphasis>
|
||
</taxonomicName>
|
||
in 1882 (
|
||
<emphasis id="B931EA9FCF54831DC5262BB58A83FBC6" box="[1059,1219,1018,1043]" italics="true" pageId="7" pageNumber="480">nomen nudum</emphasis>
|
||
), it is true that the taxonomic conception of
|
||
<taxonomicName id="4C454D0ECF54831DC0B02C698C7AFBE2" box="[437,570,1056,1079]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC0B02C698C7AFBE2" box="[437,570,1056,1079]" italics="true" pageId="7" pageNumber="480">P. antiquum</emphasis>
|
||
</taxonomicName>
|
||
as well as all of its diagnostic dental features (e.g., p4–m3 rootless, with two continuous and opposite, labial and lingual sulci that separates well-delimited trigonid and talonid; tooth row in a very tight continuous series; p4 smaller than m1–3, with the trigonid longer and wider than the talonid; m1–2 larger than premolars, with talonid longer than trigonid, and labial sulcus deeper than the lingual one; m3 larger than the other cheek teeth, with its talonid with two, labial and lingual, distal grooves that delimit a third lobe) were established based on ZMK 21/1887. At the same time, these characteristics were used later by Ameghino to include and contrast the remaining species within the Genus
|
||
<taxonomicName id="4C454D0ECF54831DC3E62CBF8DD3FADA" authorityName="Ameghino" authorityYear="1882" box="[739,915,1270,1295]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="genus">
|
||
<emphasis id="B931EA9FCF54831DC3E62CBF8DD3FADA" box="[739,915,1270,1295]" italics="true" pageId="7" pageNumber="480">Protypotherium</emphasis>
|
||
</taxonomicName>
|
||
. Therefore, we designate ZMK 21/1887 as the
|
||
<typeStatus id="54FE882FCF54831DC1922D538EBEFAE6" box="[151,254,1306,1331]" pageId="7" pageNumber="480" type="lectotype">lectotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C454D0ECF54831DC02D2D558FEFFAE6" box="[296,431,1308,1331]" class="Mammalia" family="Interatheriidae" genus="Protypotherium" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="7" pageNumber="480" phylum="Chordata" rank="species" species="antiquum">
|
||
<emphasis id="B931EA9FCF54831DC02D2D558FEFFAE6" box="[296,431,1308,1331]" italics="true" pageId="7" pageNumber="480">P. antiquum</emphasis>
|
||
</taxonomicName>
|
||
and, consequently, the remaining
|
||
<typeStatus id="54FE882FCF54831DC23E2D528DD2FAE6" box="[827,914,1307,1331]" pageId="7" pageNumber="480" type="syntype">syntype</typeStatus>
|
||
becomes the
|
||
<typeStatus id="54FE882FCF54831DC5372D538A89FAE6" box="[1074,1225,1306,1331]" pageId="7" pageNumber="480" type="paralectotype">paralectotype</typeStatus>
|
||
(ICZN 2000: Art. 73.2.2).
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF54831DC1C22D2B8F42FA4A" blockId="7.[151,1437,151,1439]" pageId="7" pageNumber="480">
|
||
<materialsCitation id="3B2D3CD0CF54831DC1C22D2B8F42FA4A" ID-GBIF-Occurrence="1828873086" collectionCode="SALMA" location="Entre Rios Province" pageId="7" pageNumber="480" specimenCount="1" stateProvince="Entre Rios Province" typeStatus="holotype">
|
||
<emphasis id="B931EA9FCF54831DC1C22D2B8A41FAAE" bold="true" box="[199,1025,1378,1403]" pageId="7" pageNumber="480">
|
||
Chronological and geographical distribution of the
|
||
<typeStatus id="54FE882FCF54831DC2492D2B8DC0FAAE" box="[844,896,1378,1403]" pageId="7" pageNumber="480">type</typeStatus>
|
||
specimen.
|
||
</emphasis>
|
||
Huayquerian
|
||
<collectionCode id="ED54AE48CF54831DC5AF2D2A8B4FFAAE" box="[1194,1295,1379,1403]" pageId="7" pageNumber="480">SALMA</collectionCode>
|
||
,
|
||
<collectingRegion id="4981F86FCF54831DC41A2D2A8EBFFA4A" country="Argentina" name="Entre Rios" pageId="7" pageNumber="480">
|
||
Entre
|
||
<collectingRegion id="4981F86FCF54831DC46C2D2A8EBFFA4A" inferredBySuffix="true" pageId="7" pageNumber="480">Ríos Province</collectingRegion>
|
||
</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |