200 lines
21 KiB
XML
200 lines
21 KiB
XML
<document id="BA98ED162ADFAE03AC84412CF3872BFB" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza & Reguero, Marcelo A." docDate="2018" docId="03EC879BCF47830EC1922A278F42F911" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patriarchus altus Ameghino 1891" docType="treatment" docVersion="7" lastPageNumber="493" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="493" updateTime="1698526258101" updateUser="ExternalLinkService">
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<mods:title id="9B93984F07AC5B98F281E7B28CDF4CEC">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
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<mods:namePart id="C35C952850BBEFDF3B8F927B5BEE2677">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="76FB50E4965FE0368C984EABDAC2A5F7">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="7EC97D85D081FC8D135FAFCA94267C9B">Cerdeño, Esperanza</mods:namePart>
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<mods:namePart id="1C64190AE340743979FFC9C02C213768">Reguero, Marcelo A.</mods:namePart>
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<mods:title id="AF19AA88326B3CE527FBDE167116E347">Zootaxa</mods:title>
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<mods:date id="731BB9D13C0F31C2D3AFC3459F03362E">2018</mods:date>
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<mods:number id="D54DB85579F9DA81D046D23801D9221A">2018-02-28</mods:number>
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<mods:number id="3472D01C2C65A03292F9943AFBD75E2D">4387</mods:number>
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<treatment id="03EC879BCF47830EC1922A278F42F911" ID-DOI="http://doi.org/10.5281/zenodo.5986713" ID-GBIF-Taxon="141264040" ID-Zenodo-Dep="5986713" LSID="urn:lsid:plazi:treatment:03EC879BCF47830EC1922A278F42F911" httpUri="http://treatment.plazi.org/id/03EC879BCF47830EC1922A278F42F911" lastPageNumber="493" pageId="20" pageNumber="493">
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<subSubSection id="C35F6506CF47830EC1922A278C7AFD5D" box="[151,570,622,648]" pageId="20" pageNumber="493" type="nomenclature">
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<paragraph id="8BFA368DCF47830EC1922A278C7AFD5D" blockId="20.[151,570,622,648]" box="[151,570,622,648]" pageId="20" pageNumber="493">
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<heading id="D0B281E1CF47830EC1922A278C7AFD5D" bold="true" box="[151,570,622,648]" fontSize="11" level="1" pageId="20" pageNumber="493" reason="1">
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<taxonomicName id="4C454D0ECF47830EC1922A278C7AFD5D" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[151,570,622,648]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="altus">
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<emphasis id="B931EA9FCF47830EC1922A278C7AFD5D" bold="true" box="[151,570,622,648]" pageId="20" pageNumber="493">
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<emphasis id="B931EA9FCF47830EC1922A278F29FD5D" bold="true" box="[151,361,622,648]" italics="true" pageId="20" pageNumber="493">Patriarchus altus</emphasis>
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<bibRefCitation id="EFD44B7CCF47830EC06A2A278C7AFD5D" box="[367,570,622,648]" pageId="20" pageNumber="493" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino, 1891</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF47830EC1922AFC8F50FCC3" pageId="20" pageNumber="493" type="materials_examined">
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<paragraph id="8BFA368DCF47830EC1922AFC8D2FFD1B" blockId="20.[151,1437,693,1732]" box="[151,879,693,718]" pageId="20" pageNumber="493">
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<emphasis id="B931EA9FCF47830EC1922AFC8F4BFD1B" bold="true" box="[151,267,693,718]" pageId="20" pageNumber="493">Holotype.</emphasis>
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MACN-A 4001 (
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<figureCitation id="137E2A08CF47830EC0D72AFC8C1EFD1B" box="[466,606,693,718]" captionStart="FIGURE 4" captionStartId="16.[151,249,1284,1306]" captionTargetBox="[162,1415,221,1217]" captionTargetId="figure@16.[155,1432,193,1262]" captionTargetPageId="16" captionText="FIGURE 4. Patriarchus leptocephalus, holotype MACN-A 3989, incomplete skull, in ventral (A) and dorsal (B) views. Patriarchus altus, holotype MACN-A 4001, incomplete skull, in ventral (C) and dorsal (D) views. Patriarchus icochiloides, holotype MACN-A 4021, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (E); holotype MACN-A 4022, right mandibular fragment with alveoli of p2–3 and p4–m3, in occlusal view (F). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187705/files/figure.png" pageId="20" pageNumber="493">Figs. 4 C–D</figureCitation>
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), almost complete skull.
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</paragraph>
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<paragraph id="8BFA368DCF47830EC1C22A908F50FCC3" blockId="20.[151,1437,693,1732]" pageId="20" pageNumber="493">
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<emphasis id="B931EA9FCF47830EC1C22A908F7BFD27" bold="true" box="[199,315,729,754]" pageId="20" pageNumber="493">
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<typeStatus id="54FE882FCF47830EC1C22A908F76FD27" box="[199,310,729,754]" pageId="20" pageNumber="493" type="paratype">Paratype</typeStatus>
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.
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</emphasis>
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<collectionCode id="ED54AE48CF47830EC0402A938FF7FD27" box="[325,439,730,754]" pageId="20" pageNumber="493">MACN-A</collectionCode>
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3999, incomplete skull, and
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<collectionCode id="ED54AE48CF47830EC20A2A938DC1FD27" box="[783,897,730,754]" pageId="20" pageNumber="493">MACN-A</collectionCode>
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4000, incomplete mandible, both of the same individual.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF47830EC1C22B688D3CFB8F" pageId="20" pageNumber="493" type="diagnosis">
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<paragraph id="8BFA368DCF47830EC1C22B688EBEFC1F" blockId="20.[151,1437,693,1732]" pageId="20" pageNumber="493">
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<emphasis id="B931EA9FCF47830EC1C22B688F7AFCEF" bold="true" box="[199,314,801,826]" pageId="20" pageNumber="493">Diagnosis</emphasis>
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(
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<bibRefCitation id="EFD44B7CCF47830EC0552B688C13FCEF" box="[336,595,801,826]" pageId="20" pageNumber="493" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891: 293</bibRefCitation>
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). “
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<emphasis id="B931EA9FCF47830EC37C2B688B46FCEF" box="[633,1286,801,826]" italics="true" pageId="20" pageNumber="493">Igual conformación general que la especie precedente</emphasis>
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[
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<taxonomicName id="4C454D0ECF47830EC41E2B688F74FC8B" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="leptocephalus">Patriarchus leptocephalus</taxonomicName>
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],
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<emphasis id="B931EA9FCF47830EC04B2B0F8EABFC1F" italics="true" pageId="20" pageNumber="493">pero de tamaño mucho mayor; rostro más elevado y paladar un poco más ancho adelante. Longitud del cráneo, 102 milímetros; longitud de la serie dentaria superior, 59 milímetros; ancho interorbital de los frontales, 28 milímetros; longitud de los frontales, 30 milímetros. Los frontales no penetran adelante entre los nasales</emphasis>
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||
”.
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||
</paragraph>
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<paragraph id="8BFA368DCF47830EC1C22B9C8D3CFB8F" blockId="20.[151,1437,693,1732]" pageId="20" pageNumber="493">
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English translation. Same general configuration as the preceding species [
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<taxonomicName id="4C454D0ECF47830EC52A2B9C8B1CFC3B" authorityName="Ameghino" authorityYear="1891" box="[1071,1372,981,1006]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="leptocephalus">
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<emphasis id="B931EA9FCF47830EC52A2B9C8B1CFC3B" box="[1071,1372,981,1006]" italics="true" pageId="20" pageNumber="493">Patriarchus leptocephalus</emphasis>
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</taxonomicName>
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], but much larger; higher rostrum and palate a little wider anteriorly. Length of the skull, 102 millimetres; length of upper tooth row, 59 millimetres; interorbital width of the frontals, 28 millimetres; length of the frontals, 30 millimetres. Frontals do not penetrate forward between the nasals.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C35F6506CF47830EC1C22C2C8F42F911" pageId="20" pageNumber="493" type="discussion">
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<paragraph id="8BFA368DCF47830EC1C22C2C8C52F9E1" blockId="20.[151,1437,693,1732]" pageId="20" pageNumber="493">
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<emphasis id="B931EA9FCF47830EC1C22C2C8F0FFBAB" bold="true" box="[199,335,1125,1150]" pageId="20" pageNumber="493">Comments.</emphasis>
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<bibRefCitation id="EFD44B7CCF47830EC05B2C2C8C6EFBAB" box="[350,558,1125,1150]" pageId="20" pageNumber="493" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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described this species, but did not add any figure of it. According to Ameghino’s catalogue, the skull MACN-A 4001 is the
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<typeStatus id="54FE882FCF47830EC3FB2CC38D6EFB77" box="[766,814,1162,1186]" pageId="20" pageNumber="493">type</typeStatus>
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of this species, whereas the cranial fragment MACN-A 3999 and its mandibular portion MACN-A 4000 are indicated each as “
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<emphasis id="B931EA9FCF47830EC2CF2CF88A58FB1D" box="[970,1048,1200,1224]" italics="true" pageId="20" pageNumber="493">tipo 2°</emphasis>
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” (2nd
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<typeStatus id="54FE882FCF47830EC5592CF98AD0FB1D" box="[1116,1168,1200,1224]" pageId="20" pageNumber="493">type</typeStatus>
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), meaning that they are
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<typeStatus id="54FE882FCF47830EC1922C9C8F46FB38" box="[151,262,1237,1261]" pageId="20" pageNumber="493" type="paratype">paratypes</typeStatus>
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.
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<bibRefCitation id="EFD44B7CCF47830EC0142C9C8FEFFB38" box="[273,431,1236,1261]" pageId="20" pageNumber="493" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
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also indicated MACN-A 3999 to MACN-A 4001 as the
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<typeStatus id="54FE882FCF47830EC5372C9C8A22FB38" box="[1074,1122,1237,1261]" pageId="20" pageNumber="493">type</typeStatus>
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specimens of
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<taxonomicName id="4C454D0ECF47830EC4002C9C8B29FB38" authorityName="Ameghino" authorityYear="1891" box="[1285,1385,1236,1261]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="altus">
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<emphasis id="B931EA9FCF47830EC4002C9C8B29FB38" box="[1285,1385,1236,1261]" italics="true" pageId="20" pageNumber="493">Pa. altus</emphasis>
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</taxonomicName>
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, but did not distinguish between
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<typeStatus id="54FE882FCF47830EC0DA2CB58C4FFAC1" box="[479,527,1276,1300]" pageId="20" pageNumber="493">type</typeStatus>
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and 2nd
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<typeStatus id="54FE882FCF47830EC3762CB58CE7FAC1" box="[627,679,1276,1300]" pageId="20" pageNumber="493">type</typeStatus>
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. The skull MACN-A 4001 matches the original diagnosis as it is larger (length of C–M3 =
|
||
<quantity id="4CBD9B68CF47830EC0C12D568C7AFAE2" box="[452,570,1311,1336]" metricMagnitude="-2" metricUnit="m" metricValue="4.505" pageId="20" pageNumber="493" unit="mm" value="45.05">45.05 mm</quantity>
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||
) than the
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<typeStatus id="54FE882FCF47830EC3B52D568D52FAED" box="[688,786,1311,1336]" pageId="20" pageNumber="493" type="holotype">holotype</typeStatus>
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||
of
|
||
<taxonomicName id="4C454D0ECF47830EC2392D698A48FAED" authorityName="Ameghino" authorityYear="1891" box="[828,1032,1311,1336]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="leptocephalus">
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||
<emphasis id="B931EA9FCF47830EC2392D698A48FAED" box="[828,1032,1311,1336]" italics="true" pageId="20" pageNumber="493">Pa. leptocephalus</emphasis>
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||
</taxonomicName>
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(length of C–M3 =
|
||
<quantity id="4CBD9B68CF47830EC5F12D568B2CFAE2" box="[1268,1388,1311,1336]" metricMagnitude="-2" metricUnit="m" metricValue="3.735" pageId="20" pageNumber="493" unit="mm" value="37.35">37.35 mm</quantity>
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||
), its rostrum is more elevated and the anterior region of the palate is wider. All the measurements (skull length =
|
||
<quantity id="4CBD9B68CF47830EC44A2D0A8E86FAAA" metricMagnitude="-1" metricUnit="m" metricValue="1.0325" pageId="20" pageNumber="493" unit="mm" value="103.25">103.25 mm</quantity>
|
||
; length of the upper tooth row =
|
||
<quantity id="4CBD9B68CF47830EC3362D2E8CE7FAAA" box="[563,679,1383,1408]" metricMagnitude="-2" metricUnit="m" metricValue="5.8549999999999995" pageId="20" pageNumber="493" unit="mm" value="58.55">58.55 mm</quantity>
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||
; interorbital width of the frontals =
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||
<quantity id="4CBD9B68CF47830EC5312D2E8AE8FAAA" box="[1076,1192,1383,1408]" metricMagnitude="-2" metricUnit="m" metricValue="2.926" pageId="20" pageNumber="493" unit="mm" value="29.26">29.26 mm</quantity>
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||
; length of the frontals =
|
||
<quantity id="4CBD9B68CF47830EC1A82DC28EBDFA76" box="[173,253,1419,1443]" metricMagnitude="-2" metricUnit="m" metricValue="3.3" pageId="20" pageNumber="493" unit="mm" value="33.0">33 mm</quantity>
|
||
) also match those provided by
|
||
<bibRefCitation id="EFD44B7CCF47830EC35F2DC28D65FA76" box="[602,805,1419,1444]" pageId="20" pageNumber="493" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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||
. As a result, we confirm MACN-A 4001 as the
|
||
<typeStatus id="54FE882FCF47830EC43F2DC28BDCFA71" box="[1338,1436,1419,1444]" pageId="20" pageNumber="493" type="holotype">holotype</typeStatus>
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of
|
||
<taxonomicName id="4C454D0ECF47830EC1B02DF98F58FA1D" authorityName="Ameghino" authorityYear="1891" box="[181,280,1455,1480]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="altus">
|
||
<emphasis id="B931EA9FCF47830EC1B02DF98F58FA1D" box="[181,280,1455,1480]" italics="true" pageId="20" pageNumber="493">Pa. altus</emphasis>
|
||
</taxonomicName>
|
||
. In the case of the individual MACN-A 3999 plus MACN-A 4000, it is also confirmed as the
|
||
<typeStatus id="54FE882FCF47830EC4392DF98BDCFA1D" box="[1340,1436,1456,1480]" pageId="20" pageNumber="493" type="paratype">paratype</typeStatus>
|
||
because, despite the fact that MACN-A 3999 is not complete enough to take any of the given measurements, none of these specimens contradicts the diagnosis and both were recognised by Ameghino as “
|
||
<emphasis id="B931EA9FCF47830EC5C92DB18B5EF9DA" box="[1228,1310,1527,1551]" italics="true" pageId="20" pageNumber="493">tipo 2°</emphasis>
|
||
” when he acknowledged the
|
||
<typeStatus id="54FE882FCF47830EC06C2E558FD9F9E1" box="[361,409,1564,1588]" pageId="20" pageNumber="493">type</typeStatus>
|
||
specimen.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF47830EC1C22E098D52F9A9" blockId="20.[151,1437,693,1732]" pageId="20" pageNumber="493">
|
||
In summary, we confirm MACN-A 4001 as the
|
||
<typeStatus id="54FE882FCF47830EC3F72E768D14F98D" box="[754,852,1599,1624]" pageId="20" pageNumber="493" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C454D0ECF47830EC27B2E098A09F98D" authorityName="Ameghino" authorityYear="1891" box="[894,1097,1599,1624]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="leptocephalus">
|
||
<emphasis id="B931EA9FCF47830EC27B2E098A09F98D" box="[894,1097,1599,1624]" italics="true" pageId="20" pageNumber="493">Pa. leptocephalus</emphasis>
|
||
</taxonomicName>
|
||
and the individual MACN-A 3999 plus MACN-A 4000 as the
|
||
<typeStatus id="54FE882FCF47830EC30E2E2D8C2BF9A9" box="[523,619,1636,1660]" pageId="20" pageNumber="493" type="paratype">paratype</typeStatus>
|
||
of the species.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF47830EC1C22ECE8F42F911" blockId="20.[151,1437,693,1732]" pageId="20" pageNumber="493">
|
||
<materialsCitation id="3B2D3CD0CF47830EC1C22ECE8F42F911" ID-GBIF-Occurrence="1828873088" collectionCode="SALMA" location="Santa Cruz Province" pageId="20" pageNumber="493" specimenCount="1" stateProvince="Santa Cruz Province" typeStatus="holotype">
|
||
<emphasis id="B931EA9FCF47830EC1C22ECE8DBEF975" bold="true" box="[199,1022,1671,1696]" pageId="20" pageNumber="493">
|
||
Chronological and geographical distribution of the
|
||
<typeStatus id="54FE882FCF47830EC24C2ECE8D3DF975" box="[841,893,1671,1696]" pageId="20" pageNumber="493">type</typeStatus>
|
||
specimen.
|
||
</emphasis>
|
||
Santacrucian
|
||
<collectionCode id="ED54AE48CF47830EC5A32ECE8B4BF94A" box="[1190,1291,1671,1695]" pageId="20" pageNumber="493">SALMA</collectionCode>
|
||
,
|
||
<collectingRegion id="4981F86FCF47830EC41E2ECE8EBFF911" country="Argentina" name="Santa Cruz" pageId="20" pageNumber="493">
|
||
Santa
|
||
<collectingRegion id="4981F86FCF47830EC4632ECE8EBFF911" inferredBySuffix="true" pageId="20" pageNumber="493">Cruz Province</collectingRegion>
|
||
</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |