176 lines
20 KiB
XML
176 lines
20 KiB
XML
<document id="B15BB51AD32A588B06E0331C66779897" ID-CLB-Dataset="30626" ID-DOI="10.11646/zootaxa.4387.3.4" ID-GBIF-Dataset="43de8956-0706-4f8d-b5a9-f51c30c276d8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1187697" ID-ZooBank="B53A6B20-5386-4E5A-A00F-A559EDF640DF" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1520059483769" checkinUser="plazi" docAuthor="Fernández, Mercedes, Fernicola, Juan Carlos, Cerdeño, Esperanza & Reguero, Marcelo A." docDate="2018" docId="03EC879BCF40830EC1922C538F42FDCE" docLanguage="en" docName="zootaxa.4387.3.4.pdf" docOrigin="Zootaxa 4387 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Patriarchus leptocephalus Ameghino 1891" docType="treatment" docVersion="7" lastPageNumber="493" masterDocId="FFD5FFE3CF53831AC10528498E40FFD5" masterDocTitle="Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino" masterLastPageNumber="498" masterPageNumber="473" pageNumber="492" updateTime="1698526258101" updateUser="ExternalLinkService">
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<mods:title id="C7665CF2616F5A703A9A5D2374ACDA5D">Identification of type materials of the species of Protypotherium Ameghino, 1885 and Patriarchus Ameghino, 1889 (Notoungulata: Interatheriidae) erected by Florentino Ameghino</mods:title>
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<mods:namePart id="6FE82460AE4422FA1D7F24828411D410">Fernández, Mercedes</mods:namePart>
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<mods:namePart id="E2818661E21C5D79F901545B8246FC3E">Fernicola, Juan Carlos</mods:namePart>
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<mods:namePart id="F77287B5FE98F4817FE78CB32F56CC83">Cerdeño, Esperanza</mods:namePart>
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<mods:namePart id="87C9377CB3E574B40CA0187D23ECC967">Reguero, Marcelo A.</mods:namePart>
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<mods:date id="DD4AB6AE8F87793F1236B1D3387DA172">2018</mods:date>
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<treatment id="03EC879BCF40830EC1922C538F42FDCE" ID-DOI="http://doi.org/10.5281/zenodo.5986711" ID-GBIF-Taxon="141264044" ID-Zenodo-Dep="5986711" LSID="urn:lsid:plazi:treatment:03EC879BCF40830EC1922C538F42FDCE" httpUri="http://treatment.plazi.org/id/03EC879BCF40830EC1922C538F42FDCE" lastPageId="20" lastPageNumber="493" pageId="19" pageNumber="492">
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<subSubSection id="C35F6506CF408309C1922C538CE5FBE0" box="[151,677,1050,1077]" pageId="19" pageNumber="492" type="nomenclature">
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<paragraph id="8BFA368DCF408309C1922C538CE5FBE0" blockId="19.[151,677,1050,1077]" box="[151,677,1050,1077]" pageId="19" pageNumber="492">
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<heading id="D0B281E1CF408309C1922C538CE5FBE0" bold="true" box="[151,677,1050,1077]" fontSize="11" level="1" pageId="19" pageNumber="492" reason="1">
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<taxonomicName id="4C454D0ECF408309C1922C538CE5FBE0" authority="Ameghino, 1891" authorityName="Ameghino" authorityYear="1891" box="[151,677,1050,1077]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="leptocephalus">
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<emphasis id="B931EA9FCF408309C1922C538CE5FBE0" bold="true" box="[151,677,1050,1077]" pageId="19" pageNumber="492">
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<emphasis id="B931EA9FCF408309C1922C538F94FBE1" bold="true" box="[151,468,1050,1076]" italics="true" pageId="19" pageNumber="492">Patriarchus leptocephalus</emphasis>
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<bibRefCitation id="EFD44B7CCF408309C0DE2C528CE5FBE0" box="[475,677,1051,1077]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino, 1891</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C35F6506CF408309C1922C288DE4FB17" pageId="19" pageNumber="492" type="materials_examined">
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<paragraph id="8BFA368DCF408309C1922C288D2FFBAF" blockId="19.[151,1437,1121,2016]" box="[151,879,1121,1146]" pageId="19" pageNumber="492">
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<emphasis id="B931EA9FCF408309C1922C288F4BFBAF" bold="true" box="[151,267,1121,1146]" pageId="19" pageNumber="492">Holotype.</emphasis>
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MACN-A 3989 (
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<figureCitation id="137E2A08CF408309C0D72C288C1EFBAF" box="[466,606,1121,1146]" captionStart="FIGURE 4" captionStartId="16.[151,249,1284,1306]" captionTargetBox="[162,1415,221,1217]" captionTargetId="figure@16.[155,1432,193,1262]" captionTargetPageId="16" captionText="FIGURE 4. Patriarchus leptocephalus, holotype MACN-A 3989, incomplete skull, in ventral (A) and dorsal (B) views. Patriarchus altus, holotype MACN-A 4001, incomplete skull, in ventral (C) and dorsal (D) views. Patriarchus icochiloides, holotype MACN-A 4021, right maxillary fragment with alveolus of P1 and P2–M3, in occlusal view (E); holotype MACN-A 4022, right mandibular fragment with alveoli of p2–3 and p4–m3, in occlusal view (F). Scale bar = 10 mm." httpUri="https://zenodo.org/record/1187705/files/figure.png" pageId="19" pageNumber="492">Figs. 4 A–B</figureCitation>
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), almost complete skull.
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</paragraph>
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<paragraph id="8BFA368DCF408309C1C22CCC8DE4FB17" blockId="19.[151,1437,1121,2016]" pageId="19" pageNumber="492">
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<emphasis id="B931EA9FCF408309C1C22CCC8F7BFB4B" bold="true" box="[199,315,1157,1182]" pageId="19" pageNumber="492">Paratype.</emphasis>
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MACN-A 3990, left cranial fragment with complete
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<date id="FFFB104DCF408309C29A2CCF8DF2FB4B" box="[927,946,1158,1182]" pageId="19" pageNumber="492">I1</date>
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–M3 series, and MACN-A 3991, complete left mandibular ramus with
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<date id="FFFB104DCF408309C0CB2CE08F9FFB17" box="[462,479,1193,1218]" pageId="19" pageNumber="492">i1</date>
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–m3 series, both of the same individual.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF408309C1C22C848AD8F9D3" pageId="19" pageNumber="492" type="diagnosis">
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<paragraph id="8BFA368DCF408309C1C22C848C71FAA3" blockId="19.[151,1437,1121,2016]" pageId="19" pageNumber="492">
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<emphasis id="B931EA9FCF408309C1C22C848F7AFB33" bold="true" box="[199,314,1229,1254]" pageId="19" pageNumber="492">Diagnosis</emphasis>
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(
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<bibRefCitation id="EFD44B7CCF408309C04F2C848C03FB33" box="[330,579,1229,1254]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino 1891: 293</bibRefCitation>
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). “
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<emphasis id="B931EA9FCF408309C3612C878BDCFB33" box="[612,1436,1229,1254]" italics="true" pageId="19" pageNumber="492">Tamaño pequeño, pero poco mayor aun que el de la especie precedente</emphasis>
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[
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<taxonomicName id="4C454D0ECF408309C1982CB88FEDFADF" authorityName="Ameghino" authorityYear="1891" box="[157,429,1265,1290]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="diastematus">Patriarchus diastematus</taxonomicName>
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].
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<emphasis id="B931EA9FCF408309C0C62CB88C5DFAA3" italics="true" pageId="19" pageNumber="492">Frontales muy angostos; hocico angosto y prolongado; paladar muy angosto adelante. Toda la forma del cráneo angosta y prolongada. Longitud del cráneo, 82 milímetros; ancho interorbital de los frontales, 21 milímetros; longitud de los frontales sobre la línea media, 21 milímetros. Adelante, los frontales penetran un poco entre los nasales</emphasis>
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”.
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</paragraph>
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<paragraph id="8BFA368DCF408309C1C22DC88AD8F9D3" blockId="19.[151,1437,1121,2016]" pageId="19" pageNumber="492">
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English translation. Species of small size, but slightly larger than the previous species [
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<taxonomicName id="4C454D0ECF408309C4122DC88F5DFA6B" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="diastematus">
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<emphasis id="B931EA9FCF408309C4122DC88F5DFA6B" italics="true" pageId="19" pageNumber="492">Patriarchus diastematus</emphasis>
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</taxonomicName>
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]. Frontals are very narrow; snout is narrow and long; palate very narrow anteriorly. The whole skull is narrow and long. Length of the skull, 82 millimetres; interorbital width of the frontals, 21 millimetres; length of the frontals on the midline, 21 millimetres. Anteriorly, the frontals slightly penetrate the nasals.
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</paragraph>
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</subSubSection>
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<subSubSection id="C35F6506CF40830EC1C22E588F42FDCE" lastPageId="20" lastPageNumber="493" pageId="19" pageNumber="492" type="discussion">
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<paragraph id="8BFA368DCF40830EC1C22E588DE0FE5E" blockId="19.[151,1437,1121,2016]" lastBlockId="20.[151,1437,151,539]" lastPageId="20" lastPageNumber="493" pageId="19" pageNumber="492">
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<emphasis id="B931EA9FCF408309C1C22E588F0FF9FF" bold="true" box="[199,335,1553,1578]" pageId="19" pageNumber="492">Comments.</emphasis>
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<bibRefCitation id="EFD44B7CCF408309C05F2E588C66F9FF" box="[346,550,1553,1578]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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described this taxon without illustrating any specimens of it. According to Ameghino’s catalogue the skull MACN-A 3989 is the type of this species, whereas both incomplete skull MACN- A 3990 and mandible MACN-A 3991 are indicated each as “
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<emphasis id="B931EA9FCF408309C24B2E148DDBF9A1" box="[846,923,1628,1652]" italics="true" pageId="19" pageNumber="492">tipo 2°</emphasis>
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” (2nd type), an old and informal term used to refer to a paratype (C. Sciosia, pers. comm.).
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<bibRefCitation id="EFD44B7CCF408309C38A2EC88D6CF94C" box="[655,812,1664,1689]" pageId="19" pageNumber="492" refId="ref18248" refString="Mones A. (1986) Palaeovertebrata Sudamericana. Catalogo Sistematico de los Vertebrados Fosiles de America del Sur, Parte I. Preliminary List and Bibliography. Courier Forschungsinstitut Senckenberg, 82, 1 - 625." type="journal article">Mones (1986)</bibRefCitation>
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also indicated MACN-A 3989 to MACN-A 3991 as the type specimens of the species, but did not make a distinction between type and 2nd type. The skull MACN-A 3989 matches the original diagnosis as it is slightly larger (length of P2–M3 =
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<quantity id="4CBD9B68CF408309C5052E828A2AF936" box="[1024,1130,1739,1763]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="19" pageNumber="492" unit="mm" value="30.2">30.2 mm</quantity>
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) than the holotype of
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<taxonomicName id="4C454D0ECF408309C4732E858F5DF8D2" authorityName="Ameghino" authorityYear="1891" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="19" pageNumber="492" phylum="Chordata" rank="species" species="diastematus">
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<emphasis id="B931EA9FCF408309C4732E858F5DF8D2" italics="true" pageId="19" pageNumber="492">Pa. diastematus</emphasis>
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</taxonomicName>
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(length of P2–M3 =
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<quantity id="4CBD9B68CF408309C30A2EA68C39F8D2" box="[527,633,1775,1799]" metricMagnitude="-2" metricUnit="m" metricValue="2.81" pageId="19" pageNumber="492" unit="mm" value="28.1">28.1 mm</quantity>
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), the frontals penetrate the nasals, the rostrum is long and the palate is anteriorly narrow; and almost all the measurements (length of the skull =
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<quantity id="4CBD9B68CF408309C2FB2F5A8A37F8FE" box="[1022,1143,1811,1836]" metricMagnitude="-2" metricUnit="m" metricValue="8.345" pageId="19" pageNumber="492" unit="mm" value="83.45">83.45 mm</quantity>
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; intraorbital width of the frontals =
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<quantity id="4CBD9B68CF408309C0082F7E8F1EF89A" box="[269,350,1847,1871]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="19" pageNumber="492" unit="mm" value="22.0">22 mm</quantity>
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) also match those given by
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<bibRefCitation id="EFD44B7CCF408309C39A2F7F8D2CF89A" box="[671,876,1846,1871]" pageId="19" pageNumber="492" refId="ref17076" refString="Ameghino, F. (1891) Nuevos restos de mamiferos fosiles descubiertos por Carlos Ameghino en el Eoceno inferior de la Patagonia austral. Especies nuevas, adiciones y correcciones. Revista Argentina de Historia Natural, 1 (5), 289 - 328." type="journal article">Ameghino (1891)</bibRefCitation>
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. Just the length of the frontals on the middle line (
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<quantity id="4CBD9B68CF408309C19A2F128EB3F8A6" box="[159,243,1883,1907]" metricMagnitude="-2" metricUnit="m" metricValue="3.1" pageId="19" pageNumber="492" unit="mm" value="31.0">31 mm</quantity>
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) is greater than the length (
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||
<quantity id="4CBD9B68CF408309C3372F128CC5F8A6" box="[562,645,1883,1907]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="19" pageNumber="492" unit="mm" value="21.0">21 mm</quantity>
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) provided by the author. Nevertheless, this was likely a typing error, because Ameghino established that the frontals were ‘very narrow’, which is contradictory with the same measure indicated for both the width and length of the frontals (
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<quantity id="4CBD9B68CF408309C3FE2FEA8D0EF86E" box="[763,846,1955,1979]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="19" pageNumber="492" unit="mm" value="21.0">21 mm</quantity>
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). MACN-A 3990 is not complete enough to take any of the measurements given by Ameghino, but it cannot be discarded as part of the paratype, because it does not contradict the diagnosis and the author recognised this specimen as 2nd type when he acknowledged MACN-A 3989 as type in his catalogue. As MACN-A 3991 is a partial mandible of the same individual as MACN-A 3990, it is also recognised in this contribution as part of the paratype of
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<taxonomicName id="4C454D0ECF47830EC27F28AD8A06FF2E" authorityName="Ameghino" authorityYear="1891" box="[890,1094,226,251]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="leptocephalus">
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<emphasis id="B931EA9FCF47830EC27F28AD8A06FF2E" box="[890,1094,226,251]" italics="true" pageId="20" pageNumber="493">Pa. leptocephalus</emphasis>
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</taxonomicName>
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. Regarding the remaining ten specimens assigned to the species in Ameghino’s catalogue, MACN-A 9653 is a cranial element, but only the length of the skull (
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<quantity id="4CBD9B68CF47830EC07D29628F8BFE96" box="[376,459,299,323]" metricMagnitude="-2" metricUnit="m" metricValue="8.3" pageId="20" pageNumber="493" unit="mm" value="83.0">83 mm</quantity>
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) resembles Ameghino’s (1891) diagnosis, so it is discarded as part of the type series. The other nine catalogued materials are fragmented mandibles and, consequently, discarded as the type specimens; among them, only MACN-A 11191 is currently within the collection.
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</paragraph>
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<paragraph id="8BFA368DCF47830EC1C229D18D52FE06" blockId="20.[151,1437,151,539]" pageId="20" pageNumber="493">
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In summary, MACN-A 3989 is confirmed as the
|
||
<typeStatus id="54FE882FCF47830EC3FD29DF8D1AFE7A" box="[760,858,406,431]" pageId="20" pageNumber="493" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C454D0ECF47830EC28729D18A0DFE7A" authorityName="Ameghino" authorityYear="1891" box="[898,1101,406,431]" class="Mammalia" family="Interatheriidae" genus="Patriarchus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Notoungulata" pageId="20" pageNumber="493" phylum="Chordata" rank="species" species="leptocephalus">
|
||
<emphasis id="B931EA9FCF47830EC28729D18A0DFE7A" box="[898,1101,406,431]" italics="true" pageId="20" pageNumber="493">Pa. leptocephalus</emphasis>
|
||
</taxonomicName>
|
||
and the individual MACN-A 3990 plus MACN-A 3991 as the
|
||
<typeStatus id="54FE882FCF47830EC30E29F28C2BFE06" box="[523,619,443,467]" pageId="20" pageNumber="493" type="paratype">paratype</typeStatus>
|
||
of the species.
|
||
</paragraph>
|
||
<paragraph id="8BFA368DCF47830EC1C229978F42FDCE" blockId="20.[151,1437,151,539]" pageId="20" pageNumber="493">
|
||
<materialsCitation id="3B2D3CD0CF47830EC1C229978F42FDCE" ID-GBIF-Occurrence="1828873083" collectionCode="SALMA" location="Santa Cruz Province" pageId="20" pageNumber="493" specimenCount="1" stateProvince="Santa Cruz Province" typeStatus="holotype">
|
||
<emphasis id="B931EA9FCF47830EC1C229978DBEFE22" bold="true" box="[199,1022,478,503]" pageId="20" pageNumber="493">
|
||
Chronological and geographical distribution of the
|
||
<typeStatus id="54FE882FCF47830EC24C29978D3DFE22" box="[841,893,478,503]" pageId="20" pageNumber="493">type</typeStatus>
|
||
specimen.
|
||
</emphasis>
|
||
Santacrucian
|
||
<collectionCode id="ED54AE48CF47830EC5A329968B4BFE22" box="[1190,1291,479,503]" pageId="20" pageNumber="493">SALMA</collectionCode>
|
||
,
|
||
<collectingRegion id="4981F86FCF47830EC41E29968EBFFDCE" country="Argentina" name="Santa Cruz" pageId="20" pageNumber="493">
|
||
Santa
|
||
<collectingRegion id="4981F86FCF47830EC46329968EBFFDCE" inferredBySuffix="true" pageId="20" pageNumber="493">Cruz Province</collectingRegion>
|
||
</collectingRegion>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |