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<document id="5691F177BD4285E98A40C059462C3B61" ID-DOI="10.11646/zootaxa.5361.3.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="10144864" ID-ZooBank="AFEFD8C9-ABC1-4793-8B83-49FB7B526389" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="juliana" checkinTime="1700163474722" checkinUser="plazi" docAuthor="Koreeda, Reo, Maeda, Ken &amp; Motomura, Hiroyuki" docDate="2023" docId="03EC5C3E1C460C71FF7BFB60FAEDFED0" docLanguage="en" docName="zootaxa.5361.3.5.pdf" docOrigin="Zootaxa 5361 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.3.5/52197" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Luciogobius platycephalus Shiogaki &amp; Dotsu 1976" docType="treatment" docVersion="3" lastPageNumber="391" masterDocId="FFD524461C470C73FFECFFAFFF84FFCD" masterDocTitle="A new subtropical species of goby of the genus Luciogobius (Gobiidae) from southwestern Japan" masterLastPageNumber="408" masterPageNumber="390" pageNumber="391" updateTime="1700496976976" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:title id="AE347DA5E21658528F9689F153D959C6">A new subtropical species of goby of the genus Luciogobius (Gobiidae) from southwestern Japan</mods:title>
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<mods:namePart id="B5A2ECF50026D8C2FF61A247F3715028">Koreeda, Reo</mods:namePart>
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<mods:namePart id="D967AAD9EBBF5AE434CAA2259F9D103B">Maeda, Ken</mods:namePart>
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<paragraph id="8BFAED281C460C72FF7BFB60FC7DFB24" blockId="1.[151,1017,1231,1258]" box="[151,1017,1231,1258]" pageId="1" pageNumber="391">
<heading id="D0B25A441C460C72FF7BFB60FC7DFB24" bold="true" box="[151,1017,1231,1258]" fontSize="11" level="1" pageId="1" pageNumber="391" reason="1">
<emphasis id="B931313A1C460C72FF7BFB60FC7DFB24" bold="true" box="[151,1017,1231,1258]" pageId="1" pageNumber="391">
The
<taxonomicName id="4C4596AB1C460C72FF23FB60FD97FB27" ID-CoL="3WCMP" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[207,531,1231,1258]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FF23FB60FD97FB27" bold="true" box="[207,531,1231,1258]" italics="true" pageId="1" pageNumber="391">Luciogobius platycephalus</emphasis>
</taxonomicName>
complex
<emphasis id="B931313A1C460C72FD66FB60FD4AFB24" bold="true" box="[650,718,1231,1257]" italics="true" pageId="1" pageNumber="391">sensu</emphasis>
<bibRefCitation id="EFD490D91C460C72FD3AFB60FC7DFB24" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[726,1017,1231,1258]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FC85FB60FC19FB27" bold="true" box="[873,925,1231,1258]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
. (2019)
</bibRefCitation>
</emphasis>
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</paragraph>
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<subSubSection id="C35FBEA31C460C72FF7BFAB8FE1EFA28" pageId="1" pageNumber="391" type="description">
<paragraph id="8BFAED281C460C72FF7BFAB8FE1EFA28" blockId="1.[151,1437,1303,2014]" pageId="1" pageNumber="391">
<emphasis id="B931313A1C460C72FF7BFAB8FE57FAFC" bold="true" box="[151,467,1303,1329]" pageId="1" pageNumber="391">Morphological characters.</emphasis>
Species of the
<taxonomicName id="4C4596AB1C460C72FD91FAB8FC2FFAFC" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[637,939,1303,1329]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FD91FAB8FC2FFAFC" box="[637,939,1303,1329]" italics="true" pageId="1" pageNumber="391">Luciogobius platycephalus</emphasis>
</taxonomicName>
complex possess the following combination of characters: second vertebrae with pleural ribs (sometimes one side absent); anus to anal-fin origin (AAA) longer than half body depth at anus to anal-fin origin; dorsal-fin origin just above or behind anterior 1/3 part of anal-fin base; total dorsal-fin rays usually 1012; pectoral fin with 27 free uppermost rays; posterior margin of pectoral-fin membrane between each fin ray relatively concave; anteriormost pterygiophore of anal fin inserted between second to fourth haemal spine.
</paragraph>
</subSubSection>
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<paragraph id="8BFAED281C460C72FF2BFA40FBFBF9E0" blockId="1.[151,1437,1303,2014]" pageId="1" pageNumber="391">
<emphasis id="B931313A1C460C72FF2BFA40FC37F9C4" bold="true" box="[199,947,1519,1546]" pageId="1" pageNumber="391">
Species included in the
<taxonomicName id="4C4596AB1C460C72FE17FA40FCBCF9C7" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[507,824,1519,1546]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FE17FA40FCBCF9C7" bold="true" box="[507,824,1519,1546]" italics="true" pageId="1" pageNumber="391">Luciogobius platycephalus</emphasis>
</taxonomicName>
complex:
</emphasis>
<taxonomicName id="4C4596AB1C460C72FC2DFA40FB2FF9C4" authority="Koreeda &amp; Maeda &amp; Motomura, 2023" authorityName="Koreeda &amp; Maeda &amp; Motomura" authorityYear="2023" box="[961,1195,1519,1545]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="griseus" status="sp. nov.">
<emphasis id="B931313A1C460C72FC2DFA40FB2FF9C4" box="[961,1195,1519,1545]" italics="true" pageId="1" pageNumber="391">Luciogobius griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2028C411C460C72FB55FA5FFB7AF9C7" box="[1209,1278,1520,1546]" pageId="1" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
,
<taxonomicName id="4C4596AB1C460C72FAFEFA40FDC9F9E0" authority="Shiogaki &amp; Dotsu, 1976" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FAFEFA40FEB7F9E0" italics="true" pageId="1" pageNumber="391">Luciogobius platycephalus</emphasis>
<bibRefCitation id="EFD490D91C460C72FED6F9BCFDC9F9E0" author="Shiogaki, M. &amp; Dotsu, Y." box="[314,589,1555,1582]" pageId="1" pageNumber="391" pagination="125 - 129" refId="ref12198" refString="Shiogaki, M. &amp; Dotsu, Y. (1976) Two new species of the genus Luciogobius (family Gobiidae) from Japan. Japanese Journal of Ichthyology, 23 (3), 125 - 129. [https: // www. jstage. jst. go. jp / article / jji 1950 / 23 / 3 / 23 _ 3 _ 125 / _ pdf / - char / ja]" type="journal article" year="1976">Shiogaki &amp; Dotsu, 1976</bibRefCitation>
</taxonomicName>
,
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<emphasis id="B931313A1C460C72FDB4F9BCFD66F9E0" box="[600,738,1555,1581]" italics="true" pageId="1" pageNumber="391">Luciogobius</emphasis>
sp. 7
</taxonomicName>
<emphasis id="B931313A1C460C72FCC9F9BBFCE7F9E0" box="[805,867,1556,1581]" italics="true" pageId="1" pageNumber="391">sensu</emphasis>
<bibRefCitation id="EFD490D91C460C72FC87F9BCFBFFF9E0" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[875,1147,1555,1582]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FC02F9BBFB9BF9E0" box="[1006,1055,1555,1581]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
. (2019)
</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35FBEA31C460C71FF2BF998FAEDFED0" lastPageId="2" lastPageNumber="392" pageId="1" pageNumber="391" type="discussion">
<paragraph id="8BFAED281C460C72FF2BF998FCFFF8BC" blockId="1.[151,1437,1303,2014]" pageId="1" pageNumber="391">
<emphasis id="B931313A1C460C72FF2BF998FEBFF99C" bold="true" box="[199,315,1591,1617]" pageId="1" pageNumber="391">Remarks.</emphasis>
Members of the
<taxonomicName id="4C4596AB1C460C72FDEFF997FD3BF99C" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[515,703,1591,1617]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FDEFF997FD3BF99C" box="[515,703,1591,1617]" italics="true" pageId="1" pageNumber="391">L. platycephalus</emphasis>
</taxonomicName>
complex can be distinguished from other species complexes by the presence of pleural ribs on the second vertebra, except the
<taxonomicName id="4C4596AB1C460C72FCB3F9F3FC50F9B8" authorityName="Arai" authorityYear="1970" box="[863,980,1627,1653]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B931313A1C460C72FCB3F9F3FC50F9B8" box="[863,980,1627,1653]" italics="true" pageId="1" pageNumber="391">L. grandis</emphasis>
</taxonomicName>
complex
<emphasis id="B931313A1C460C72FBA9F9F3FB07F9B8" box="[1093,1155,1628,1653]" italics="true" pageId="1" pageNumber="391">sensu</emphasis>
<bibRefCitation id="EFD490D91C460C72FB67F9F4FA18F9B8" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[1163,1436,1627,1654]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FAFCF9F3FAC7F9B8" box="[1296,1347,1627,1653]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
. (2019)
</bibRefCitation>
(
<bibRefCitation id="EFD490D91C460C72FF73F9D0FE27F954" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[159,419,1663,1690]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FECFF92FFED0F954" box="[291,340,1663,1689]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
., 2019
</bibRefCitation>
; this study). The latter complex differs from the
<taxonomicName id="4C4596AB1C460C72FC50F92FFBF1F954" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[956,1141,1663,1689]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FC50F92FFBF1F954" box="[956,1141,1663,1689]" italics="true" pageId="1" pageNumber="391">L. platycephalus</emphasis>
</taxonomicName>
complex by having 1218 (usually more than 13) dorsal-fin rays, AAA shorter than half the body depth at the anus to anal-fin origin [except for
<taxonomicName id="4C4596AB1C460C72FF53F968FE02F92C" box="[191,390,1735,1762]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="undefined-6">
<emphasis id="B931313A1C460C72FF53F968FECDF92C" box="[191,329,1735,1761]" italics="true" pageId="1" pageNumber="391">Luciogobius</emphasis>
sp. 6
</taxonomicName>
<emphasis id="B931313A1C460C72FE62F967FE48F92C" box="[398,460,1736,1761]" italics="true" pageId="1" pageNumber="391">sensu</emphasis>
<bibRefCitation id="EFD490D91C460C72FE3FF968FD62F92C" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[467,742,1735,1762]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FDBBF967FD0DF92C" box="[599,649,1735,1761]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
. (2019)
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], dorsal-fin origin just vertical through or slightly posterior to anal-fin origin (except for
<taxonomicName id="4C4596AB1C460C72FE51F944FD00F8C8" box="[445,644,1771,1798]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="undefined-6">
<emphasis id="B931313A1C460C72FE51F944FDC3F8C8" box="[445,583,1771,1797]" italics="true" pageId="1" pageNumber="391">Luciogobius</emphasis>
sp. 6
</taxonomicName>
), and the anteriormost anal-fin pterygiophore inserted between the first and second haemal spines [sometimes inserted between the second and third haemal spines in
<taxonomicName id="4C4596AB1C460C72FB3EF8A0FA18F8E4" box="[1234,1436,1807,1834]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="undefined-4">
<emphasis id="B931313A1C460C72FB3EF8A0FAD8F8E4" box="[1234,1372,1807,1833]" italics="true" pageId="1" pageNumber="391">Luciogobius</emphasis>
sp. 4
</taxonomicName>
<emphasis id="B931313A1C460C72FF7BF89BFF51F880" box="[151,213,1844,1869]" italics="true" pageId="1" pageNumber="391">sensu</emphasis>
<bibRefCitation id="EFD490D91C460C72FF31F89CFE39F880" author="Maeda, K. &amp; Yamasaki, N. &amp; Kondo, M. &amp; Tachihara, K." box="[221,445,1843,1870]" pageId="1" pageNumber="391" pagination="162 - 174" refId="ref11457" refString="Maeda, K., Yamasaki, N., Kondo, M. &amp; Tachihara, K. (2008) Occurrence and morphology of larvae and juveniles of six Luciogobius species from Aritsu Beach, Okinawa Island. Ichthyological Research, 55, 162 - 174. https: // doi. org / 10.1007 / s 10228 - 007 - 0018 - x" type="journal article" year="2008">
Maeda
<emphasis id="B931313A1C460C72FEDCF89BFEE6F880" box="[304,354,1843,1869]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
. (2008)
</bibRefCitation>
] (
<bibRefCitation id="EFD490D91C460C72FE3FF89CFD5EF880" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[467,730,1843,1870]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FDB4F89BFD0EF880" box="[600,650,1843,1869]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
., 2019
</bibRefCitation>
;
<bibRefCitation id="EFD490D91C460C72FD0AF89CFBADF880" author="Koreeda, R. &amp; Motomura, H." box="[742,1065,1843,1870]" pageId="1" pageNumber="391" pagination="27 - 52" refId="ref11086" refString="Koreeda, R. &amp; Motomura, H. (2021) Records of seven temperate and subtropical species of Luciogobius from Shimokoshikishima island, Koshiki Islands, East China Sea, Japan, and the first record of the possible hybrid individuals for Luciogobius. Ichthy, Natural History of Fishes of Japan, 11, 27 - 52. [In Japanese] https: // doi. org / 10.34583 / ichthy. 11.0 _ 27" type="journal article" year="2021">Koreeda &amp; Motomura, 2021</bibRefCitation>
). Sexual dimorphism of the head is apparent in the
<taxonomicName id="4C4596AB1C460C72FEB2F8F7FD9CF8BC" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[350,536,1879,1905]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FEB2F8F7FD9CF8BC" box="[350,536,1879,1905]" italics="true" pageId="1" pageNumber="391">L. platycephalus</emphasis>
</taxonomicName>
complex (see
<taxonomicName id="4C4596AB1C460C72FD56F8F7FCACF8BC" authority="Koreeda &amp; Maeda &amp; Motomura, 2023" authorityName="Koreeda &amp; Maeda &amp; Motomura" authorityYear="2023" box="[698,808,1880,1905]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="griseus" status="n. sp.">
<emphasis id="B931313A1C460C72FD56F8F7FCACF8BC" box="[698,808,1880,1905]" italics="true" pageId="1" pageNumber="391">L. griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2028C411C460C72FCC3F8F7FCEAF8BC" box="[815,878,1879,1906]" pageId="1" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
).
</paragraph>
<paragraph id="8BFAED281C460C71FF2BF8D4FAEDFED0" blockId="1.[151,1437,1303,2014]" lastBlockId="2.[151,1437,151,285]" lastPageId="2" lastPageNumber="392" pageId="1" pageNumber="391">
<bibRefCitation id="EFD490D91C460C72FF2BF8D4FE54F858" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[199,464,1915,1942]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FEA5F8D3FEFDF858" box="[329,377,1915,1941]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
. (2019)
</bibRefCitation>
listed differences in fresh or live coloration between the
<taxonomicName id="4C4596AB1C460C72FBACF8D3FB36F858" authorityName="Arai" authorityYear="1970" box="[1088,1202,1915,1941]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="grandis">
<emphasis id="B931313A1C460C72FBACF8D3FB36F858" box="[1088,1202,1915,1941]" italics="true" pageId="1" pageNumber="391">L. grandis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C4596AB1C460C72FB09F8D3FA19F858" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[1253,1437,1915,1941]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C460C72FB09F8D3FA19F858" box="[1253,1437,1915,1941]" italics="true" pageId="1" pageNumber="391">L. platycephalus</emphasis>
</taxonomicName>
complexes: dark green to dark gray in the former vs. bright yellow to orange in the latter. However, the fresh or live coloration of
<taxonomicName id="4C4596AB1C460C72FEC1F86BFE1FF810" authority="Koreeda &amp; Maeda &amp; Motomura, 2023" authorityName="Koreeda &amp; Maeda &amp; Motomura" authorityYear="2023" box="[301,411,1988,2013]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="griseus" status="n. sp.">
<emphasis id="B931313A1C460C72FEC1F86BFE1FF810" box="[301,411,1988,2013]" italics="true" pageId="1" pageNumber="391">L. griseus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2028C411C460C72FE4EF86BFE59F813" box="[418,477,1988,2014]" pageId="1" pageNumber="391" rank="species">n. sp.</taxonomicNameLabel>
and
<taxonomicName id="4C4596AB1C460C72FDFFF86CFD5CF813" box="[531,728,1987,2014]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="391" phylum="Chordata" rank="species" species="undefined-7">
<emphasis id="B931313A1C460C72FDFFF86CFD19F810" box="[531,669,1987,2013]" italics="true" pageId="1" pageNumber="391">Luciogobius</emphasis>
sp. 7
</taxonomicName>
are also dark green or dark yellow (see
<bibRefCitation id="EFD490D91C460C72FB78F86CFA1CF810" author="Shibukawa, K. &amp; Aizawa, M. &amp; Suzuki, T. &amp; Kanagawa, N. &amp; Muto, F." box="[1172,1432,1987,2014]" pageId="1" pageNumber="391" pagination="29 - 96" refId="ref12013" refString="Shibukawa, K., Aizawa, M., Suzuki, T., Kanagawa, N. &amp; Muto, F. (2019) Preliminary review of earthworm gobies of the genus Luciogobius (Gobiiformes, Oxudercidae) from Shizuoka Prefecture, Japan. Bulletin of the Museum of Natural and Environmental History, Shizuoka, 12, 29 - 96. [In Japanese]" type="journal article" year="2019">
Shibukawa
<emphasis id="B931313A1C460C72FAFBF86BFACCF810" box="[1303,1352,1987,2013]" italics="true" pageId="1" pageNumber="391">et al</emphasis>
., 2019
</bibRefCitation>
: figs. 2021; this study). Juveniles and adults of
<taxonomicName id="4C4596AB1C450C71FD40FF38FCE2FF7C" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[684,870,151,177]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="392" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C450C71FD40FF38FCE2FF7C" box="[684,870,151,177]" italics="true" pageId="2" pageNumber="392">L. platycephalus</emphasis>
</taxonomicName>
are usually bright yellow to green in life, although larger adults often have relatively darker coloration (this study). In addition, the population of
<taxonomicName id="4C4596AB1C450C71FB29FF14FAFBFF18" authorityName="Shiogaki &amp; Dotsu" authorityYear="1976" box="[1221,1407,187,213]" family="Gobiidae" genus="Luciogobius" kingdom="Animalia" order="Perciformes" pageId="2" pageNumber="392" phylum="Chordata" rank="species" species="platycephalus">
<emphasis id="B931313A1C450C71FB29FF14FAFBFF18" box="[1221,1407,187,213]" italics="true" pageId="2" pageNumber="392">L. platycephalus</emphasis>
</taxonomicName>
in northwestern
<collectingRegion id="498123CA1C450C71FEDEFF70FE36FF34" box="[306,434,223,249]" country="Japan" name="Kagoshima" pageId="2" pageNumber="392">Kagoshima</collectingRegion>
Bay, Kyushu,
<collectingCountry id="F352ADB81C450C71FDB0FF70FD18FF34" box="[604,668,223,249]" name="Japan" pageId="2" pageNumber="392">Japan</collectingCountry>
is relatively darker than those in other sites (
<bibRefCitation id="EFD490D91C450C71FB4FFF70FA15FF34" author="Koreeda, R. &amp; Yamashita, R. &amp; Furuhashi, R. &amp; Saito, H. &amp; Motomura, H." box="[1187,1425,223,249]" pageId="2" pageNumber="392" pagination="357 - 366" refId="ref11281" refString="Koreeda, R., Yamashita, R., Furuhashi, R., Saito, H. &amp; Motomura, H. (2020) First records of Luciogobius sp. 1 (sensu Shibukawa et al. 2019), L. platycephalus, and L. elongatus from Kagoshima Bay, southern Kyushu, Japan. Nature of Kagoshima, 46, 357 - 366. [In Japanese]" type="journal article" year="2020">
Koreeda
<emphasis id="B931313A1C450C71FAE7FF70FAB9FF34" box="[1291,1341,223,249]" italics="true" pageId="2" pageNumber="392">et al</emphasis>
., 2020
</bibRefCitation>
). Possibly, the darker bodied specimen reported by
<bibRefCitation id="EFD490D91C450C71FD29FEACFC34FED0" author="Koreeda, R. &amp; Yamashita, R. &amp; Furuhashi, R. &amp; Saito, H. &amp; Motomura, H." box="[709,944,259,285]" pageId="2" pageNumber="392" pagination="357 - 366" refId="ref11281" refString="Koreeda, R., Yamashita, R., Furuhashi, R., Saito, H. &amp; Motomura, H. (2020) First records of Luciogobius sp. 1 (sensu Shibukawa et al. 2019), L. platycephalus, and L. elongatus from Kagoshima Bay, southern Kyushu, Japan. Nature of Kagoshima, 46, 357 - 366. [In Japanese]" type="journal article" year="2020">
Koreeda
<emphasis id="B931313A1C450C71FCC7FEACFCD8FED0" box="[811,860,259,285]" italics="true" pageId="2" pageNumber="392">et al</emphasis>
. (2020
</bibRefCitation>
: fig. 3C, F) was an axanthic individual.
</paragraph>
</subSubSection>
</treatment>
</document>