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<mods:title id="ED73B23E2804F73082D33B2311853239">Revision of Chaetacanthus Seidler, 1922 (Annelida, Phyllodocida, Polynoidae)</mods:title>
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<paragraph id="8BFAF60FFFD8E54C66D0FF01FD4567D7" blockId="20.[151,664,151,213]" box="[151,664,151,178]" pageId="20" pageNumber="415">
<heading id="D0B24163FFD8E54C66D0FF01FD4567D7" bold="true" box="[151,664,151,178]" fontSize="11" level="1" pageId="20" pageNumber="415" reason="1">
<taxonomicName id="4C458D8CFFD8E54C66D0FF01FD4567D7" ID-CoL="5XMLB" authority="(Kinberg, 1856)" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[151,664,151,178]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="415" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD8E54C66D0FF01FD4567D7" bold="true" box="[151,664,151,178]" pageId="20" pageNumber="415">
<emphasis id="B9312A1DFFD8E54C66D0FF01FE1167D7" bold="true" box="[151,460,151,178]" italics="true" pageId="20" pageNumber="415">Chaetacanthus pomareae</emphasis>
(
<bibRefCitation id="EFD48BFEFFD8E54C679CFF01FD4D67D7" author="Kinberg, J. G. H." box="[475,656,151,177]" pageId="20" pageNumber="415" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856</bibRefCitation>
)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="C35FA584FFD8E54C66D0FF2DFEE567B3" box="[151,312,187,213]" pageId="20" pageNumber="415" type="description">
<paragraph id="8BFAF60FFFD8E54C66D0FF2DFEE567B3" blockId="20.[151,664,151,213]" box="[151,312,187,213]" pageId="20" pageNumber="415">
<figureCitation id="137EEA8AFFD8E54C66D0FF2DFECD67B3" box="[151,272,187,213]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="20" pageNumber="415">Figures 17</figureCitation>
,
<figureCitation id="137EEA8AFFD8E54C675BFF2DFEE567B3" box="[284,312,187,213]" captionStart="FIGURE 18" captionStartId="21.[151,250,1442,1466]" captionTargetBox="[170,1418,598,1418]" captionTargetId="figure-211@21.[170,1418,597,1419]" captionTargetPageId="21" captionText="FIGURE 18. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. First elytron, dorsal view. B. Same, globular macrotubercles. C. Elytron form middle segments. D. Same, surface with conical microtubercles and macrotubercles. E. Same, marginal conical macrotubercles. F. Spiny microtubercles from first elytra. G, H. Close-up of conical macrotubercles. Scale bars: A, C: 0.1 mm; B, D, E: 0.1 mm; F, G: 0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4296809" httpUri="https://zenodo.org/record/4296809/files/figure.png" pageId="20" pageNumber="415">18</figureCitation>
</paragraph>
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<subSubSection id="C35FA584FFD8E54C66D0FE96FC9A6650" pageId="20" pageNumber="415" type="reference_group">
<paragraph id="8BFAF60FFFD8E54C66D0FE96FC9A6650" blockId="20.[151,1389,255,310]" pageId="20" pageNumber="415">
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<treatmentCitation id="0AE4D01EFFD8E54C66D0FE96FD93667E" author="Kinberg, J. G. H." box="[151,590,255,280]" page="383" pageId="20" pageNumber="415" year="1856">
<taxonomicName id="4C458D8CFFD8E54C66D0FE96FD93667E" ID-CoL="3TCD2" authority="Kinberg, 1856: 383" authorityName="Kinberg" authorityPageNumber="383" authorityYear="1856" box="[151,590,255,280]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="415" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD8E54C66D0FE96FEA26671" box="[151,383,256,280]" italics="true" pageId="20" pageNumber="415">Lepidonotus pomareae</emphasis>
<bibRefCitation id="EFD48BFEFFD8E54C67C2FE96FD93667E" author="Kinberg, J. G. H." box="[389,590,255,280]" pageId="20" pageNumber="415" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856: 383</bibRefCitation>
</taxonomicName>
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;
<treatmentCitation id="0AE4D01EFFD8E54C641FFE96FCD1667E" author="Kinberg, J. G. H." box="[600,780,255,280]" page="10" pageId="20" pageNumber="415" year="1858">
<bibRefCitation id="EFD48BFEFFD8E54C641FFE96FCD1667E" author="Kinberg, J. G. H." box="[600,780,255,280]" pageId="20" pageNumber="415" refId="ref14372" refString="Kinberg, J. G. H. (1858) Kongliga Svenska Fregatten Eugenies Resa omkring jorden under befal af C. A. Virgin aren 1851 - 1853. Zoologi, 1. Annulater. Almquist and Wicksells, Uppsala &amp; Stockholm, 32 pp., 8 pls." type="book" year="1858">Kinberg 1858: 10</bibRefCitation>
</treatmentCitation>
, Pl. 3,
<figureCitation id="137EEA8AFFD8E54C6515FE96FC52667E" box="[850,911,256,280]" captionStart="FIGURE 9" captionStartId="11.[151,250,1454,1478]" captionTargetBox="[158,1429,181,1430]" captionTargetId="figure-21@11.[158,1429,181,1430]" captionTargetPageId="11" captionText="FIGURE 9. Chaetacanthus harrisae n. sp., holotype (LACM 245). A. First elytron, dorsal view. B. Same, spiny microtubercles. CE. Same, close up of spiny microtubercles. F. Elytron of middle segment, dorsal view. G. Globular central macrotubercles from first elytron. Scale bars: A, F: 1.1 mm; B, G: 0.1 mm; C: 0.3 mm; D, E: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296791" httpUri="https://zenodo.org/record/4296791/files/figure.png" pageId="20" pageNumber="415">Fig. 9</figureCitation>
, Pl. 10, Fig. 47;
<treatmentCitation id="0AE4D01EFFD8E54C6270FE96FB03667E" author="Seidler, H. J." box="[1079,1246,256,280]" page="20" pageId="20" pageNumber="415" year="1924">
<bibRefCitation id="EFD48BFEFFD8E54C6270FE96FB03667E" author="Seidler, H. J." box="[1079,1246,256,280]" pageId="20" pageNumber="415" pagination="1 - 217" refId="ref15811" refString="Seidler, H. J. (1924 (1923 )) Beitrage zur Kenntnis der Polynoiden, 1. Archiv f ̡ r Natugeschichte Berlin, 89 (11), 1 - 217. https: // doi. org / 10.1002 / mmnz. 4830110105" type="journal article" year="1924">Seidler 1924: 20</bibRefCitation>
</treatmentCitation>
(key), 3233.
<treatmentCitation id="0AE4D01EFFD8E54C66D0FE88FD546650" author="Hartman, O." box="[151,649,286,310]" page="29" pageId="20" pageNumber="415" year="1948">
<taxonomicName id="4C458D8CFFD8E54C66D0FE88FD546650" ID-CoL="5XMLB" authority=": Hartman 1948: 27 - 29" authorityName="Hartman" authorityPageNumber="27 - 29" authorityYear="1948" box="[151,649,286,310]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="415" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD8E54C66D0FE88FE7F6650" box="[151,418,286,310]" italics="true" pageId="20" pageNumber="415">Chaetacanthus pomareae:</emphasis>
<bibRefCitation id="EFD48BFEFFD8E54C67EEFE89FD546650" author="Hartman, O." box="[425,649,286,310]" pageId="20" pageNumber="415" pagination="1 - 137" refId="ref14008" refString="Hartman, O. (1948) The marine annelids erected by Kinberg with some notes on some other types in the Swedish State Museum. Arkiv for Zoologi, 42 A (1), 1 - 137." type="journal article" year="1948">Hartman 1948: 2729</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, Pl. 3,
<figureCitation id="137EEA8AFFD8E54C6488FE88FCC16650" box="[719,796,286,310]" captionStart="FIGURE 13" captionStartId="15.[151,250,1914,1938]" captionTargetBox="[174,1414,1177,1887]" captionTargetId="figure-476@15.[170,1418,1173,1890]" captionTargetPageId="15" captionText="FIGURE 13. AC. Chaetacanthus ornatus n. sp., holotype (ECOSUR 233) elytra in dorsal view. DF. C. brasiliensis, nontype specimen (ECOSUR P 3100) elytra; A, D. Elytra of first pair. B, E. Right elytra of middle segment. C, F. Right elytron of posterior segment. Scale bars: A, B: 0.6 mm; C, D: 0.7 mm; E: 1.4 mm; F: 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4296799" httpUri="https://zenodo.org/record/4296799/files/figure.png" pageId="20" pageNumber="415">Figs 13</figureCitation>
,
<figureCitation id="137EEA8AFFD8E54C6560FE88FC9E6650" box="[807,835,286,310]" captionStart="FIGURE 14" captionStartId="17.[151,250,1828,1852]" captionTargetBox="[160,1425,420,1804]" captionTargetId="figure-102@17.[157,1430,417,1805]" captionTargetPageId="17" captionText="FIGURE 14. Chaetacanthus pilosus (Treadwell, 1937), holotype (AMNH 3531). A. Anterior end, dorsal view. B. First elytron, dorsal view. C. Elytron from second pair, dorsal view. D. Elytron from posterior segment, dorsal view. E. Parapodium from median segment (arrow points to branchiae), anterior view. F. Parapodium from another median segment (arrow points to branchiae). Scale bars: A: 2.4 mm; BD: 1.1 mm; E, F: 0.4 mm." figureDoi="http://doi.org/10.5281/zenodo.4296801" httpUri="https://zenodo.org/record/4296801/files/figure.png" pageId="20" pageNumber="415">14</figureCitation>
.
</treatmentCitationGroup>
</paragraph>
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<paragraph id="8BFAF60FFFD8E54C66D0FEF3FCA266C5" blockId="20.[151,1436,356,563]" pageId="20" pageNumber="415">
<emphasis id="B9312A1DFFD8E54C66D0FEF3FDBC6618" bold="true" box="[151,609,356,383]" pageId="20" pageNumber="415">Type material. Central South Pacific,</emphasis>
<materialsCitation id="3B2DFC52FFD8E54C642BFEF3FCA266C5" ID-GBIF-Occurrence="2989844306" collectionCode="SMNH" country="French Polynesia" latitude="-20.0" location="French Polynesia" longLatPrecision="38946" longitude="-150.0" pageId="20" pageNumber="415" specimenCode="SMNH 395" specimenCount="1" typeStatus="holotype">
<emphasis id="B9312A1DFFD8E54C642BFEF3FCE06618" bold="true" box="[620,829,356,383]" pageId="20" pageNumber="415">
<collectingCountry id="F352B69FFFD8E54C642BFEF3FCE06618" box="[620,829,356,383]" name="French Polynesia" pageId="20" pageNumber="415">French Polynesia</collectingCountry>
</emphasis>
.
<typeStatus id="54FE48ADFFD8E54C6509FEF2FC6B6618" box="[846,950,356,382]" pageId="20" pageNumber="415" type="holotype">Holotype</typeStatus>
of
<taxonomicName id="4C458D8CFFD8E54C65A5FEF2FA456619" authority="Kinberg, 1856" authorityName="Kinberg" authorityYear="1856" box="[994,1432,356,383]" class="Polychaeta" family="Polynoidae" genus="Lepidonotus" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="415" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD8E54C65A5FEF2FB3B6618" box="[994,1254,356,382]" italics="true" pageId="20" pageNumber="415">Lepidonotus pomareae</emphasis>
<bibRefCitation id="EFD48BFEFFD8E54C62B6FEF2FA456619" author="Kinberg, J. G. H." box="[1265,1432,356,383]" pageId="20" pageNumber="415" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856</bibRefCitation>
</taxonomicName>
,
<specimenCode id="DBE35E74FFD8E54C66D0FE1FFEC166C5" box="[151,284,392,419]" collectionCode="SMNH" country="Sweden" httpUri="http://grbio.org/cool/8z6v-f8is" name="Department of Paleozoology, Swedish Museum of Natural History" pageId="20" pageNumber="415">SMNH 395</specimenCode>
, Tahiti,
<geoCoordinate id="EE7190C8FFD8E54C6731FE1FFE7366C5" box="[374,430,393,419]" degrees="20" direction="south" orientation="latitude" pageId="20" pageNumber="415" precision="55555" value="-20.0">20°S</geoCoordinate>
,
<geoCoordinate id="EE7190C8FFD8E54C67FDFE1EFDCC66C5" box="[442,529,392,419]" degrees="150" direction="west" orientation="longitude" pageId="20" pageNumber="415" precision="55555" value="-150.0">150° W</geoCoordinate>
, Freg. Eugenie Exped. 185153.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35FA584FFD8E54C6680FE3AFD006555" pageId="20" pageNumber="415" type="diagnosis">
<paragraph id="8BFAF60FFFD8E54C6680FE3AFD006555" blockId="20.[151,1436,356,563]" pageId="20" pageNumber="415">
<emphasis id="B9312A1DFFD8E54C6680FE3AFE9F66A0" bold="true" box="[199,322,428,454]" pageId="20" pageNumber="415">Diagnosis.</emphasis>
<taxonomicName id="4C458D8CFFD8E54C670EFE3AFE2C66A0" authorityName="Seidler" authorityYear="1922" box="[329,497,428,454]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="415" phylum="Annelida" rank="genus">
<emphasis id="B9312A1DFFD8E54C670EFE3AFE2C66A0" box="[329,497,428,454]" italics="true" pageId="20" pageNumber="415">Chaetacanthus</emphasis>
</taxonomicName>
with branchial filaments single, thick lobes; elytra with globular micropapillae and microtubercles; round macrotubercles larger on first pair of elytra; conical macrotubercles in following elytra, macrotubercles up to five times longer than wide, basally spinous, tips long, smooth; elytra without large, amber-colour macrotubercles arranged in honeycomb like groups.
</paragraph>
</subSubSection>
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<paragraph id="8BFAF60FFFD8E54C66D0F952FF246021" blockId="20.[151,1437,1732,1864]" pageId="20" pageNumber="415">
<emphasis id="B9312A1DFFD8E54C66D0F952FEC661BA" bold="true" box="[151,283,1732,1756]" pageId="20" pageNumber="415">FIGURE 17</emphasis>
.
<taxonomicName id="4C458D8CFFD8E54C676DF952FD0361BA" authority="(Kinberg, 1856)" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[298,734,1732,1756]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="20" pageNumber="415" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD8E54C676DF952FDF361BA" box="[298,558,1732,1756]" italics="true" pageId="20" pageNumber="415">Chaetacanthus pomareae</emphasis>
(
<bibRefCitation id="EFD48BFEFFD8E54C6479F952FD0A61BA" author="Kinberg, J. G. H." box="[574,727,1732,1756]" pageId="20" pageNumber="415" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856</bibRefCitation>
)
</taxonomicName>
, holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm.
</paragraph>
</caption>
<subSubSection id="C35FA584FFD8E54D6680F8E0FE766021" lastPageId="21" lastPageNumber="416" pageId="20" pageNumber="415" type="description">
<paragraph id="8BFAF60FFFD8E54C6680F8E0FC0B60D2" blockId="20.[151,1436,1910,2009]" pageId="20" pageNumber="415">
<emphasis id="B9312A1DFFD8E54C6680F8E0FE8460F6" bold="true" box="[199,345,1910,1936]" pageId="20" pageNumber="415">Description.</emphasis>
<typeStatus id="54FE48ADFFD8E54C6727F8E0FE1560F6" box="[352,456,1910,1936]" pageId="20" pageNumber="415" type="holotype">Holotype</typeStatus>
in poor condition, without posterior end, 1.2 cm long, 0.3 cm wide, 23 segments. Data from description and illustrations by
<bibRefCitation id="EFD48BFEFFD8E54C6471F80CFD3F60D3" author="Kinberg, J. G. H." box="[566,738,1946,1973]" pageId="20" pageNumber="415" refId="ref14372" refString="Kinberg, J. G. H. (1858) Kongliga Svenska Fregatten Eugenies Resa omkring jorden under befal af C. A. Virgin aren 1851 - 1853. Zoologi, 1. Annulater. Almquist and Wicksells, Uppsala &amp; Stockholm, 32 pp., 8 pls." type="book" year="1858">Kinberg (1858)</bibRefCitation>
given in parentheses.
</paragraph>
<paragraph id="8BFAF60FFFD8E54D6680F828FBDE6624" blockId="20.[151,1436,1910,2009]" lastBlockId="21.[151,1437,151,573]" lastPageId="21" lastPageNumber="416" pageId="20" pageNumber="415">
Prostomium bilobed, hexagonal, as long as wide; two pairs of eyes, without pigmentation; anterior pair on middle part, posterior pair near to posterior margin (
<figureCitation id="137EEA8AFFD9E54D6499FF01FC9A67D7" box="[734,839,151,177]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Fig. 17A</figureCitation>
). Facial tubercle prominent, pointed. Median antenna with ceratophore long, inserted frontally, style long, expanded subdistally, tip filiform (slightly longer than laterals). Lateral antennae with ceratophores inserted terminally, styles thin, shorter than median antenna, similar in shape to median antenna (
<figureCitation id="137EEA8AFFD9E54D671FFE95FE1C667B" box="[344,449,259,285]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Fig. 17A</figureCitation>
). Palps with stout papillae, tapered, as long as median antenna. Pharynx missing (previously removed; 12 pairs of marginal papillae; jaws with smooth cutting edge).
</paragraph>
<paragraph id="8BFAF60FFFD9E54D6680FEDDFAFE66EF" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="416">
Tentacular segment not visible dorsally; tentaculophores long, thin, with chaetae; tentacular cirri similar in shape to antennae. Segment two dorsally projected on prostomium as a nuchal rounded lobe (
<figureCitation id="137EEA8AFFD9E54D62E9FEF9FACA66EF" box="[1198,1303,367,393]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Fig. 17A</figureCitation>
).
</paragraph>
<paragraph id="8BFAF60FFFD9E54D6680FE05FAA4655B" blockId="21.[151,1437,151,573]" pageId="21" pageNumber="416">
Elytra with fringe of short papillae; surface granular with numerous microtubercles, and digitiform papillae (
<figureCitation id="137EEA8AFFD9E54D66E7FE21FED766B7" box="[160,266,439,465]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Fig. 17G</figureCitation>
). First pair of elytra with microtubercles and macrotubercles spherical or hemispherical, spinous (
<figureCitation id="137EEA8AFFD9E54D632DFE21FF316693" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Figs 17B, C</figureCitation>
;
<figureCitation id="137EEA8AFFD9E54D66BDFE4DFE916693" box="[250,332,475,501]" captionStart="FIGURE 18" captionStartId="21.[151,250,1442,1466]" captionTargetBox="[170,1418,598,1418]" captionTargetId="figure-211@21.[170,1418,597,1419]" captionTargetPageId="21" captionText="FIGURE 18. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. First elytron, dorsal view. B. Same, globular macrotubercles. C. Elytron form middle segments. D. Same, surface with conical microtubercles and macrotubercles. E. Same, marginal conical macrotubercles. F. Spiny microtubercles from first elytra. G, H. Close-up of conical macrotubercles. Scale bars: A, C: 0.1 mm; B, D, E: 0.1 mm; F, G: 0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4296809" httpUri="https://zenodo.org/record/4296809/files/figure.png" pageId="21" pageNumber="416">18A, F</figureCitation>
), larger in elytrophoral plug area (
<figureCitation id="137EEA8AFFD9E54D649EFE4DFC9C6693" box="[729,833,475,501]" captionStart="FIGURE 18" captionStartId="21.[151,250,1442,1466]" captionTargetBox="[170,1418,598,1418]" captionTargetId="figure-211@21.[170,1418,597,1419]" captionTargetPageId="21" captionText="FIGURE 18. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. First elytron, dorsal view. B. Same, globular macrotubercles. C. Elytron form middle segments. D. Same, surface with conical microtubercles and macrotubercles. E. Same, marginal conical macrotubercles. F. Spiny microtubercles from first elytra. G, H. Close-up of conical macrotubercles. Scale bars: A, C: 0.1 mm; B, D, E: 0.1 mm; F, G: 0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4296809" httpUri="https://zenodo.org/record/4296809/files/figure.png" pageId="21" pageNumber="416">Fig. 18B</figureCitation>
). Elytra of median and posterior segments oval with fringe of slender papillae (
<figureCitation id="137EEA8AFFD9E54D67FEFE69FDC2657F" box="[441,543,511,537]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Figs 17F</figureCitation>
;
<figureCitation id="137EEA8AFFD9E54D646EFE69FDA4657F" box="[553,633,511,537]" captionStart="FIGURE 18" captionStartId="21.[151,250,1442,1466]" captionTargetBox="[170,1418,598,1418]" captionTargetId="figure-211@21.[170,1418,597,1419]" captionTargetPageId="21" captionText="FIGURE 18. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. First elytron, dorsal view. B. Same, globular macrotubercles. C. Elytron form middle segments. D. Same, surface with conical microtubercles and macrotubercles. E. Same, marginal conical macrotubercles. F. Spiny microtubercles from first elytra. G, H. Close-up of conical macrotubercles. Scale bars: A, C: 0.1 mm; B, D, E: 0.1 mm; F, G: 0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4296809" httpUri="https://zenodo.org/record/4296809/files/figure.png" pageId="21" pageNumber="416">18C, E</figureCitation>
), surface with numerous conical macrotubercles, amber incolour, sclerotized, basally spinous, tips long (
<figureCitation id="137EEA8AFFD9E54D6441FDB5FD4C655B" box="[518,657,547,573]" captionStart="FIGURE 17" captionStartId="20.[151,250,1732,1756]" captionTargetBox="[161,1409,604,1701]" captionTargetId="figure-144@20.[151,1436,586,1708]" captionTargetPageId="20" captionText="FIGURE 17. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. Anterior end, dorsal view. B, C. Microtubercles from first elytron. D, E. Macrotubercles, conical, from middle elytra. F. Marginal papillae or fringe. G. Digitiform micropapillae, distally globular. H. Neurochaetal tips form middle parapodium. Scale bars: A: 2 mm; BE, H: 0.03 mm; F, G: 0.01 mm." figureDoi="http://doi.org/10.5281/zenodo.4296807" httpUri="https://zenodo.org/record/4296807/files/figure.png" pageId="21" pageNumber="416">Figs 17D, E</figureCitation>
;
<figureCitation id="137EEA8AFFD9E54D64DBFDB5FCEE655B" box="[668,819,547,573]" captionStart="FIGURE 18" captionStartId="21.[151,250,1442,1466]" captionTargetBox="[170,1418,598,1418]" captionTargetId="figure-211@21.[170,1418,597,1419]" captionTargetPageId="21" captionText="FIGURE 18. Chaetacanthus pomareae (Kinberg, 1856), holotype (SMNH 395). A. First elytron, dorsal view. B. Same, globular macrotubercles. C. Elytron form middle segments. D. Same, surface with conical microtubercles and macrotubercles. E. Same, marginal conical macrotubercles. F. Spiny microtubercles from first elytra. G, H. Close-up of conical macrotubercles. Scale bars: A, C: 0.1 mm; B, D, E: 0.1 mm; F, G: 0.03 mm." figureDoi="http://doi.org/10.5281/zenodo.4296809" httpUri="https://zenodo.org/record/4296809/files/figure.png" pageId="21" pageNumber="416">18D, E, G, H</figureCitation>
), larger macrotubercles over elytrophoral plug area.
</paragraph>
<caption id="DF3AA687FFD9E54D66D0FA34FD2D6140" ID-DOI="http://doi.org/10.5281/zenodo.4296809" ID-Zenodo-Dep="4296809" httpUri="https://zenodo.org/record/4296809/files/figure.png" pageId="21" pageNumber="416" startId="21.[151,250,1442,1466]" targetBox="[170,1418,598,1418]" targetPageId="21">
<paragraph id="8BFAF60FFFD9E54D66D0FA34FD2D6140" blockId="21.[151,1437,1442,1574]" pageId="21" pageNumber="416">
<emphasis id="B9312A1DFFD9E54D66D0FA34FEC462DC" bold="true" box="[151,281,1442,1466]" pageId="21" pageNumber="416">FIGURE 18</emphasis>
.
<taxonomicName id="4C458D8CFFD9E54D6761FA34FD0962DC" authority="(Kinberg, 1856)" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[294,724,1442,1466]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="416" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD9E54D6761FA34FDF562DC" box="[294,552,1442,1466]" italics="true" pageId="21" pageNumber="416">Chaetacanthus pomareae</emphasis>
(
<bibRefCitation id="EFD48BFEFFD9E54D6471FA34FD1062DC" author="Kinberg, J. G. H." box="[566,717,1442,1466]" pageId="21" pageNumber="416" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856</bibRefCitation>
)
</taxonomicName>
, holotype (SMNH 395). A. First elytron, dorsal view. B. Same, globular macrotubercles. C. Elytron form middle segments. D. Same, surface with conical microtubercles and macrotubercles. E. Same, marginal conical macrotubercles. F. Spiny microtubercles from first elytra. G, H. Close-up of conical macrotubercles. Scale bars: A, C: 0.1 mm; B, D, E: 0.1 mm; F, G: 0.03 mm.
</paragraph>
</caption>
<paragraph id="8BFAF60FFFD9E54D6680F9C2FB7261BD" blockId="21.[151,1437,1620,2043]" pageId="21" pageNumber="416">Parapodia biramous, with branchial filaments laterally to eytrophores, cirrophores and between neuropodia. Notopodia shorter than neuropodia. Neuropodia with postchaetal rounded lobe, prechaetal lobe with a small acicular lobe, acicular tips slightly projected. Dorsal cirri short, thin, smooth, with subdistal swelling, tips filiform. Cirrophores short. Ventral cirri thin, short, tips filiform. Nephridial papillae from segment seven.</paragraph>
<paragraph id="8BFAF60FFFD9E54D6680F972FE766021" blockId="21.[151,1437,1620,2043]" pageId="21" pageNumber="416">Notochaetae abundant, long, slender, with minute rows of spines. Neurochaetae thicker than notochaetae; upper bundle with shorter chaetae, tips with subdistal spines. Pygidium with anal cirri long, similar to dorsal cirri, anus terminal in last segment.</paragraph>
</subSubSection>
<subSubSection id="C35FA584FFD9E54E6680F8C7FA51679F" lastPageId="22" lastPageNumber="417" pageId="21" pageNumber="416" type="discussion">
<paragraph id="8BFAF60FFFD9E54D6680F8C7FF3B60D4" blockId="21.[151,1437,1620,2043]" pageId="21" pageNumber="416">
<emphasis id="B9312A1DFFD9E54D6680F8C7FEE6600D" bold="true" box="[199,315,1873,1899]" pageId="21" pageNumber="416">Remarks.</emphasis>
The specific epithet given by Kinberg most probably refers to Queen Animata Pômare, who reigned in Tahiti from 1827 to 1877, an account of whom is given by
<bibRefCitation id="EFD48BFEFFD9E54D6505F8E2FC2F60E9" author="O'Brien, P." box="[834,1010,1908,1935]" pageId="21" pageNumber="416" pagination="108 - 129" refId="ref15009" refString="O'Brien, P. (2006) ' Think of me as a woman': Queen Pomare of Tahiti and Anglo-French imperial contest in the 1840 s Pacific. Gender &amp; History, 18, 108 - 129. https: // doi. org / 10.1111 / j. 1468 - 0424.2006.00417. x" type="journal article" year="2006">OBrien (2006)</bibRefCitation>
. The
<typeStatus id="54FE48ADFFD9E54D6269F8E3FB8360E9" box="[1070,1118,1909,1935]" pageId="21" pageNumber="416">type</typeStatus>
specimen is the only known record.
</paragraph>
<paragraph id="8BFAF60FFFD9E54E6680F82AFA51679F" blockId="21.[151,1437,1620,2043]" lastBlockId="22.[151,1436,151,321]" lastPageId="22" lastPageNumber="417" pageId="21" pageNumber="416">
The
<typeStatus id="54FE48ADFFD9E54D66BBF82AFE8360B0" box="[252,350,1980,2006]" pageId="21" pageNumber="416" type="holotype">holotype</typeStatus>
of
<taxonomicName id="4C458D8CFFD9E54D67C0F82BFD0A60B1" authority="(Kinberg, 1856)" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[391,727,1980,2007]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="21" pageNumber="416" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFD9E54D67C0F82BFDC560B0" box="[391,536,1981,2007]" italics="true" pageId="21" pageNumber="416">C. pomareae</emphasis>
(
<bibRefCitation id="EFD48BFEFFD9E54D646EF82AFD1260B1" author="Kinberg, J. G. H." box="[553,719,1980,2007]" pageId="21" pageNumber="416" pagination="381 - 388" refId="ref14336" refString="Kinberg, J. G. H. (1856) Nya slagten och arter af Annelider. Ofversigt af Kongliga Vetenskaps-Akademiens Forhhandlingar Stockholm, 12 (9 - 10), 381 - 388." type="journal article" year="1856">Kinberg, 1856</bibRefCitation>
)
</taxonomicName>
is about half as long as the critical size indicated by
<bibRefCitation id="EFD48BFEFFD9E54D637BF82BFF39609D" author="Augener, H." pageId="21" pageNumber="416" pagination="39 - 82" refId="ref13181" refString="Augener, H. (1927) Resultaten eener Reis van Dr. C. J. van der Horst in 1920: Polychaeten von Curacao. Bijdragen tot de Kennis der Fauna van Curacao. Bijdragen tot de Dierkunde, 25, 39 - 82. https: // doi. org / 10.1163 / 26660644 - 02501003" type="journal article" year="1927">Augener (1927)</bibRefCitation>
, and herein confirmed; however, instead of having many globular macrotubercles densely packed, there are just a few, and they have very long, conical macrotubercles in the elytrophoral plug area. Consequently, we are confident about using the type, size, and density of globular macrotubercles as diagnostic features, and that
<taxonomicName id="4C458D8CFFDAE54E6349FF2DFA4167B2" baseAuthorityName="Kinberg" baseAuthorityYear="1856" box="[1294,1436,187,213]" class="Polychaeta" family="Polynoidae" genus="Chaetacanthus" kingdom="Animalia" order="Phyllodocida" pageId="22" pageNumber="417" phylum="Annelida" rank="species" species="pomareae">
<emphasis id="B9312A1DFFDAE54E6349FF2DFA4167B2" box="[1294,1436,187,213]" italics="true" pageId="22" pageNumber="417">C. pomareae</emphasis>
</taxonomicName>
differs from its congeners by having a low density of globular macrotubercles, and many conical macrotubercles.
</paragraph>
</subSubSection>
<subSubSection id="C35FA584FFDAE54E6680FE95FCE4667B" box="[199,825,258,285]" pageId="22" pageNumber="417" type="materials_examined">
<paragraph id="8BFAF60FFFDAE54E6680FE95FCE4667B" blockId="22.[151,1436,151,321]" box="[199,825,258,285]" pageId="22" pageNumber="417">
<emphasis id="B9312A1DFFDAE54E6680FE95FEBB667A" bold="true" box="[199,358,258,285]" pageId="22" pageNumber="417">Type locality.</emphasis>
<materialsCitation id="3B2DFC52FFDAE54E672BFE95FCE9667B" ID-GBIF-Occurrence="2989844312" box="[364,820,258,285]" country="French Polynesia" latitude="-20.0" location="Tahiti" longLatPrecision="38946" longitude="-150.0" pageId="22" pageNumber="417" specimenCount="1">
<location id="8E9AA0D4FFDAE54E672BFE95FE6D667B" LSID="urn:lsid:plazi:treatment:03EC4719FFD8E54E66D0FF01FD266627:8E9AA0D4FFDAE54E672BFE95FE6D667B" box="[364,432,259,285]" country="French Polynesia" latitude="-20.0" longLatPrecision="38946" longitude="-150.0" name="Tahiti" pageId="22" pageNumber="417">Tahiti</location>
,
<collectingCountry id="F352B69FFFDAE54E67FCFE95FDA0667B" box="[443,637,259,285]" name="French Polynesia" pageId="22" pageNumber="417">French Polynesia</collectingCountry>
(
<geoCoordinate id="EE7190C8FFDAE54E64CAFE95FD1B667B" box="[653,710,259,285]" degrees="20" direction="south" orientation="latitude" pageId="22" pageNumber="417" precision="55555" value="-20.0">20°S</geoCoordinate>
,
<geoCoordinate id="EE7190C8FFDAE54E6495FE94FCF0667B" box="[722,813,258,285]" degrees="150" direction="west" orientation="longitude" pageId="22" pageNumber="417" precision="55555" value="-150.0">150° W</geoCoordinate>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35FA584FFDAE54E6680FEB0FD266627" box="[199,763,294,321]" pageId="22" pageNumber="417" type="distribution">
<paragraph id="8BFAF60FFFDAE54E6680FEB0FD266627" blockId="22.[151,1436,151,321]" box="[199,763,294,321]" pageId="22" pageNumber="417">
<emphasis id="B9312A1DFFDAE54E6680FEB0FEBC6626" bold="true" box="[199,353,294,320]" pageId="22" pageNumber="417">Distribution.</emphasis>
Only known from the locus typicus.
</paragraph>
</subSubSection>
</treatment>
</document>