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<document id="AC2769B5CD2B19F223CC953B8E314B32" ID-CLB-Dataset="46232" ID-DOI="10.5281/zenodo.205387" ID-GBIF-Dataset="7760eb64-e5d4-4c83-9934-64a603f461b2" ID-ISSN="1175-5326" ID-Zenodo-Dep="205387" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460396634351" checkinUser="plazi" docAuthor="Naderloo, Reza" docDate="2011" docId="03AACA4C4A456D7355F90873F0A5FBB0" docLanguage="en" docName="zt03048p043.pdf" docOrigin="Zootaxa 3048" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Grapsus albolineatus Latreille" docType="treatment" docVersion="14" lastPageNumber="5" masterDocId="FF93B2344A446D77556E0D7EF408FFBF" masterDocTitle="Grapsoid crabs (Decapoda: Brachyura: Thoracotremata) of the Persian Gulf and the Gulf of Oman" masterLastPageNumber="43" masterPageNumber="1" pageNumber="2" updateTime="1698298771184" updateUser="ExternalLinkService">
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<heading id="D0F4CC364A456D7655F90873F6D2FA98" bold="true" box="[151,730,1293,1319]" fontSize="11" level="1" pageId="1" pageNumber="2" reason="1">
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<taxonomicName id="4C0300D94A456D7655F90873F601FA98" ID-CoL="6L4W5" authority="Latreille" authorityName="Latreille" box="[151,521,1293,1319]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="albolineatus">
<emphasis id="B977A7484A456D7655F90873F59FFA98" bold="true" box="[151,407,1293,1319]" italics="true" pageId="1" pageNumber="2">Grapsus albolineatus</emphasis>
Latreille
</taxonomicName>
in Milbert, 1812
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBC7B5A4A456D7655F90851F569FAF8" blockId="1.[151,730,1293,1351]" box="[151,353,1327,1351]" pageId="1" pageNumber="2">
(
<figureCitation id="133867DF4A456D7655F10851F4E0FAF8" box="[159,232,1327,1351]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="1" pageNumber="2">Figs. 1</figureCitation>
ae, 5a, b)
</paragraph>
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<subSubSection id="C31928D14A456D7655F90807F5E6F9F9" pageId="1" pageNumber="2" type="reference_group">
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<taxonomicName id="4C0300D94A456D7655F90807F5C4FA31" ID-CoL="6L4W5" authority="Latreille" authorityName="Latreille" box="[151,460,1400,1422]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="albolineatus">
<emphasis id="B977A7484A456D7655F90807F578FA31" box="[151,368,1400,1422]" italics="true" pageId="1" pageNumber="2">Grapsus albolineatus</emphasis>
Latreille
</taxonomicName>
in Milbert, 1812: 274. —
<treatmentCitation id="0AA25D4B4A456D7657B60806F7F2FA31" author="Vannini" box="[728,1018,1400,1422]" page="64" pageId="1" pageNumber="2" year="1981">
<bibRefCitation id="EF9206AB4A456D7657B60806F7DDFA32" author="Vannini" box="[728,981,1400,1422]" pageId="1" pageNumber="2" refString="Vannini, M. &amp; Valmori, P. (1981) Researches on the coast of Somalia. The shore and the dune of Sar Uanle. 30. Grapsidae (Decapoda, Brachyura). Monitore Zoologico Italiano, (N. S.), Supplement 14, 57 - 101." type="journal article" year="1981">Vannini &amp; Valmori 1981</bibRefCitation>
: 64
</treatmentCitation>
, figs. 1D, 2D, 3D, 4D. —
<treatmentCitation id="0AA25D4B4A456D76506B0806F4EAFA13" author="Crosnier" page="13" pageId="1" pageNumber="2" year="1965">
<bibRefCitation id="EF9206AB4A456D76506B0806F190FA31" author="Crosnier" box="[1285,1432,1400,1422]" pageId="1" pageNumber="2" refString="Crosnier, A. (1965) Crustaces Decapodes, Grapsidae et Ocypodidae. Faune de Madagascar, 18, 1 - 143." type="journal article" year="1965">Crosnier 1965</bibRefCitation>
: 13
</treatmentCitation>
, figs. 8, 11. —
<treatmentCitation id="0AA25D4B4A456D76541508E8F649FA13" author="Holthuis" box="[379,577,1430,1452]" page="145" pageId="1" pageNumber="2" year="1977">
<bibRefCitation id="EF9206AB4A456D76541508E8F607FA13" author="Holthuis" box="[379,527,1430,1452]" pageId="1" pageNumber="2" refString="Holthuis, L. B. (1977) The Grapsidae, Gecarcinidae and Palicidae (Crustacea: Decapoda: Brachyura) of the Red Sea. Israel Journal of Zoology, 26 (3 - 4), 141 - 192." type="journal article" year="1977">Holthuis 1977</bibRefCitation>
: 145
</treatmentCitation>
. —
<treatmentCitation id="0AA25D4B4A456D76570408E8F711FA13" author="Titgen" box="[618,793,1430,1452]" page="142" pageId="1" pageNumber="2" year="1982">
<bibRefCitation id="EF9206AB4A456D76570408E8F6E0FA13" author="Titgen" box="[618,744,1430,1452]" pageId="1" pageNumber="2" refString="Titgen, R. H. (1982) The Systematics and Ecology of the Decapods of Dubai, and their Zoogeographic Relationships to the Persian Gulf and the Western Indian Ocean. Doctoral Dissertation, Texas A &amp; M University." type="book" year="1982">Titgen 1982</bibRefCitation>
: 142
</treatmentCitation>
, 253 (list). —
<treatmentCitation id="0AA25D4B4A456D7656C408E8F0C8FA13" author="Tirmizi" box="[938,1216,1430,1452]" page="138" pageId="1" pageNumber="2" year="1996">
<bibRefCitation id="EF9206AB4A456D7656C408E8F087FA13" author="Tirmizi" box="[938,1167,1430,1452]" pageId="1" pageNumber="2" refString="Tirmizi, N. M., Ghani, N. (1996) Marine Fauna of Pakistan: 5: Crustacea: Brachyura, Brachyrhyncha, Part 1: Xanthidae, Goneplacidae, Pinnotheridae, Ocypodidae, Grapsidae. Center of Excellence, University of Karachi, 1 - 188." type="journal article" year="1996">Tirmizi &amp; Ghani 1996</bibRefCitation>
: 138
</treatmentCitation>
, fig. 53. —
<treatmentCitation id="0AA25D4B4A456D76505608E9F553FA74" author="Carpenter" page="40" pageId="1" pageNumber="2" year="1997">
<bibRefCitation id="EF9206AB4A456D76505608E9F53DFA74" author="Carpenter" pageId="1" pageNumber="2" refString="Carpenter, K. E., Krupp, F., Jones, D. A. &amp; Zajonz, U. (1997) The Living Marine Resources of Kuwait, Eastern Saudi Arabia, Bahrain, Qatar, and the United Arab Emirates. FAO Species Identification Field Guide for Fishery Purposes, Rome, I - VIII + 1 - 293, poss. - XVII." type="book chapter" year="1997">
Carpenter
<emphasis id="B977A7484A456D7655A908C8F4FAFA74" box="[199,242,1461,1483]" italics="true" pageId="1" pageNumber="2">et al</emphasis>
. 1997
</bibRefCitation>
: 40
</treatmentCitation>
. —
<treatmentCitation id="0AA25D4B4A456D7654EB08CBF62CFA74" author="Apel" box="[389,548,1461,1483]" page="116" pageId="1" pageNumber="2" year="2001">
<bibRefCitation id="EF9206AB4A456D7654EB08CBF5FBFA74" author="Apel" box="[389,499,1461,1483]" pageId="1" pageNumber="2" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes, pp. 1 - 268. Unpublished doctoral dissertation, Johann Wolfgang Goethe-Universitat, Frankfurt am Main, Germany, 260 pp." type="book chapter" year="2001">Apel 2001</bibRefCitation>
: 116
</treatmentCitation>
, 124, 134 (list). — Ng
<emphasis id="B977A7484A456D76566108C8F732FA74" box="[783,826,1461,1483]" italics="true" pageId="1" pageNumber="2">et al</emphasis>
. 2008: 216 (list).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBC7B5A4A456D7655F908AAF501F9B7" blockId="1.[151,1437,1400,1606]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB135C744A456D7655F908AAF501F9B7" pageId="1" pageNumber="2">
<taxonomicName id="4C0300D94A456D7655F908AAF553FA55" ID-CoL="6KSW2" box="[151,347,1492,1514]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B977A7484A456D7655F908AAF553FA55" box="[151,347,1492,1514]" italics="true" pageId="1" pageNumber="2">Grapsus maculatus</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="4C0300D94A456D7654C708ABF631FA55" ID-CoL="6KT84" box="[425,569,1493,1514]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A456D7654C708ABF631FA55" box="[425,569,1493,1514]" italics="true" pageId="1" pageNumber="2">tenuicrustatus</emphasis>
</taxonomicName>
<treatmentCitation id="0AA25D4B4A456D76573108ADF749FA56" author="Stephensen" box="[607,833,1491,1513]" page="194" pageId="1" pageNumber="2" year="1945">
<bibRefCitation id="EF9206AB4A456D76573108ADF718FA56" author="Stephensen" box="[607,784,1491,1513]" pageId="1" pageNumber="2" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen 1945</bibRefCitation>
: 194
</treatmentCitation>
, 210 (list), figs. 59AD [not
<taxonomicName id="4C0300D94A456D76510508AAF4F7F9B7" authority="Herbst, 1783" authorityName="Herbst" authorityYear="1783" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A456D76510508AAF143FA55" box="[1131,1355,1492,1514]" italics="true" pageId="1" pageNumber="2">Cancer tenuicrustatus</emphasis>
<bibRefCitation id="EF9206AB4A456D76503C08ADF4F7F9B7" author="Herbst" pageId="1" pageNumber="2" refString="Herbst, J. F. W. (1783) Versuch einer Naturgeschichte der Krabben und Krebse, nebst einer systematischen Beschreibung ihrer verschiedenen Arten. Zurich, Joh. Casper Fuessly. 1 (2 - 5), 87 - 182, pls II - IX. [For dates of publication see Rathbun 1897: 157]" type="journal article" year="1783">Herbst, 1783</bibRefCitation>
</taxonomicName>
].
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBC7B5A4A456D7655F90B6CF5E6F9F9" blockId="1.[151,1437,1400,1606]" pageId="1" pageNumber="2">
<treatmentCitationGroup id="AB135C744A456D7655F90B6CF5E6F9F9" pageId="1" pageNumber="2">
<taxonomicName id="4C0300D94A456D7655F90B6CF58CF998" ID-CoL="6KT84" box="[151,388,1554,1575]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A456D7655F90B6CF58CF998" box="[151,388,1554,1575]" italics="true" pageId="1" pageNumber="2">Grapsus tenuicrustatus</emphasis>
</taxonomicName>
<treatmentCitation id="0AA25D4B4A456D7654C20B6CF682F998" author="Pretzmann" box="[428,650,1554,1575]" page="479" pageId="1" pageNumber="2" year="1971">
<bibRefCitation id="EF9206AB4A456D7654C20B6CF65DF998" author="Pretzmann" box="[428,597,1554,1575]" pageId="1" pageNumber="2" refString="Pretzmann, G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien. 2. Teil: Marine Brachyura. Annalen des Naturhistorischen Museums Wien, 75, 477 - 487." type="journal article" year="1971">Pretzmann 1971</bibRefCitation>
: 479
</treatmentCitation>
. —
<treatmentCitation id="0AA25D4B4A456D7657DB0B6CF797F998" author="Basson" box="[693,927,1553,1575]" page="47" pageId="1" pageNumber="2" year="1977">
<bibRefCitation id="EF9206AB4A456D7657DB0B6CF770F998" author="Basson" box="[693,888,1553,1575]" pageId="1" pageNumber="2" refString="Basson, P. W., Burchard, J. E., Hardy, J. T. &amp; Price, A. R. G. (1977) Biotopes of the Western Arabian Gulf: Marine Life and Environments of Saudi Arabia. ARAMCO, Dept. of Loss Prevention and environmental affairs, Dhahran, 289 pp." type="book" year="1977">
Basson
<emphasis id="B977A7484A456D7656680B6CF73CF998" box="[774,820,1553,1575]" italics="true" pageId="1" pageNumber="2">et al</emphasis>
. 1977
</bibRefCitation>
: 47
</treatmentCitation>
, 145, 227. —
<treatmentCitation id="0AA25D4B4A456D76515C0B6FF0EAF998" author="Titgen" box="[1074,1250,1553,1575]" page="253" pageId="1" pageNumber="2" year="1982">
<bibRefCitation id="EF9206AB4A456D76515C0B6FF0B9F998" author="Titgen" box="[1074,1201,1553,1575]" pageId="1" pageNumber="2" refString="Titgen, R. H. (1982) The Systematics and Ecology of the Decapods of Dubai, and their Zoogeographic Relationships to the Persian Gulf and the Western Indian Ocean. Doctoral Dissertation, Texas A &amp; M University." type="book" year="1982">Titgen 1982</bibRefCitation>
: 253
</treatmentCitation>
(list) [not
<taxonomicName id="4C0300D94A456D76503F0B6CF5ECF9F9" authority="Herbst, 1783" authorityName="Herbst" authorityYear="1783" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="2" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A456D76503F0B6CF55EF9F9" italics="true" pageId="1" pageNumber="2">Cancer tenuicrustatus</emphasis>
<bibRefCitation id="EF9206AB4A456D7654320B4EF5ECF9F9" author="Herbst" box="[348,484,1584,1606]" pageId="1" pageNumber="2" refString="Herbst, J. F. W. (1783) Versuch einer Naturgeschichte der Krabben und Krebse, nebst einer systematischen Beschreibung ihrer verschiedenen Arten. Zurich, Joh. Casper Fuessly. 1 (2 - 5), 87 - 182, pls II - IX. [For dates of publication see Rathbun 1897: 157]" type="journal article" year="1783">Herbst, 1783</bibRefCitation>
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].
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</paragraph>
</subSubSection>
<subSubSection id="C31928D14A456D7555F90BE9F7C7FDF5" lastPageId="2" lastPageNumber="3" pageId="1" pageNumber="2" type="materials_examined">
<paragraph id="8BBC7B5A4A456D7655F90BE9F5A1F911" blockId="1.[151,1436,1686,2009]" box="[151,425,1686,1711]" pageId="1" pageNumber="2">
<emphasis id="B977A7484A456D7655F90BE9F53DF910" bold="true" box="[151,309,1686,1711]" pageId="1" pageNumber="2">
<typeStatus id="54B8C5F84A456D7655F90BE9F4DAF910" box="[151,210,1687,1711]" pageId="1" pageNumber="2">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3143BCA4A456D7654520BE8F5A1F911" box="[316,425,1686,1710]" name="Mauritius" pageId="1" pageNumber="2">Mauritius</collectingCountry>
</paragraph>
<paragraph id="8BBC7B5A4A456D7655AB0BC5F799F866" blockId="1.[151,1436,1686,2009]" pageId="1" pageNumber="2">
<emphasis id="B977A7484A456D7655AB0BC5F5B8F96B" bold="true" box="[197,432,1723,1748]" pageId="1" pageNumber="2">Material examined.</emphasis>
PERSIAN
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,
<collectingCountry id="F3143BCA4A456D7657E20BC2F6C9F96B" box="[652,705,1724,1748]" name="Iran" pageId="1" pageNumber="2">
<emphasis id="B977A7484A456D7657E20BC2F6C9F96B" bold="true" box="[652,705,1724,1748]" pageId="1" pageNumber="2">Iran</emphasis>
</collectingCountry>
:
<specimenCount id="9D05B0D34A456D7657BC0BC2F716F96B" box="[722,798,1724,1748]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
(
<collectionCode id="ED12E39F4A456D7656410BC2F761F96B" LSID="urn:lsid:biocol.org:col:34838" box="[815,873,1724,1748]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="2">SMF</collectionCode>
38394), Basaeedu,
<collectingCountry id="F3143BCA4A456D7651250BC2F092F96B" box="[1099,1178,1724,1748]" name="Iran" pageId="1" pageNumber="2">Qeshm</collectingCountry>
I.,
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,
<geoCoordinate id="EE371D9D4A456D76505B0BC2F19FF96B" box="[1333,1431,1724,1748]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="55.266666">55º 16'E</geoCoordinate>
, coll. R. Naderloo &amp; A. Kazemi,
<date id="FFBD5D9A4A456D76576C0B9FF68CF946" box="[514,644,1761,1785]" pageId="1" pageNumber="2" value="2008-07-05">07.05.2008</date>
;
<specimenCount id="9D05B0D34A456D7657E00B9FF6EFF946" box="[654,743,1761,1785]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
,
<specimenCount id="9D05B0D34A456D76579C0B9FF75BF946" box="[754,851,1761,1785]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A456D76560C0B9FF794F946" LSID="urn:lsid:biocol.org:col:34838" box="[866,924,1761,1785]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="2">SMF</collectionCode>
38395), Zeytun (olive) park beach,
<collectingCountry id="F3143BCA4A456D7650400B9FF176F946" box="[1326,1406,1761,1785]" name="Iran" pageId="1" pageNumber="2">Qeshm</collectingCountry>
I.,
<geoCoordinate id="EE371D9D4A456D7655F90A78F4F2F8A1" box="[151,250,1798,1822]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="27.183332">27º 11'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A456D7654660A78F562F8A1" box="[264,362,1798,1822]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="56.4">56º 24'E</geoCoordinate>
, coll. A. Sari, M. Türkay &amp; R. Naderloo, 0 8.01.2008.
<collectingCountry id="F3143BCA4A456D7656B20A78F01FF8A0" box="[988,1047,1798,1823]" name="United Arab Emirates" pageId="1" pageNumber="2">
<emphasis id="B977A7484A456D7656B20A78F01FF8A0" bold="true" box="[988,1047,1798,1823]" pageId="1" pageNumber="2">UAE</emphasis>
</collectingCountry>
:
<specimenCount id="9D05B0D34A456D7651490A78F077F8A1" box="[1063,1151,1798,1822]" pageId="1" pageNumber="2" type="male">3 males</specimenCount>
(
<collectionCode id="ED12E39F4A456D7651E10A78F0C1F8A1" LSID="urn:lsid:biocol.org:col:34838" box="[1167,1225,1798,1822]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="2">SMF</collectionCode>
37384) Lullayyah Harbour, Sharjah,
<geoCoordinate id="EE371D9D4A456D7654060A52F5C6F8FB" box="[360,462,1836,1860]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="25.383333">25º 23'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A456D7654B50A52F635F8FB" box="[475,573,1836,1860]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="56.366665">56º 22'E</geoCoordinate>
, coll. M. Apel,
<date id="FFBD5D9A4A456D76579A0A52F77EF8FB" box="[756,886,1836,1860]" pageId="1" pageNumber="2" value="1995-06-27">27.06.1995</date>
;
<specimenCount id="9D05B0D34A456D7656EC0A52F7EDF8FB" box="[898,997,1836,1860]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A456D7656980A52F038F8FB" LSID="urn:lsid:biocol.org:col:34838" box="[1014,1072,1836,1860]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34838" name="Forschungsinstitut und Natur-Museum Senckenberg" pageId="1" pageNumber="2">SMF</collectionCode>
37385), N. coast of Ash Sham, Ras Al-Khaymah,
<geoCoordinate id="EE371D9D4A456D7654070A2FF5C5F8D6" box="[361,461,1873,1897]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="26.033333">26º 02'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A456D7654B70A2FF632F8D6" box="[473,570,1873,1897]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="55.033333">55º 02'E</geoCoordinate>
, coll. M. Apel,
<date id="FFBD5D9A4A456D7657850A2FF765F8D6" box="[747,877,1873,1897]" pageId="1" pageNumber="2" value="1995-10-07">10.07.1995</date>
.
<collectingCountry id="F3143BCA4A456D7656160A2FF010F8D5" box="[888,1048,1873,1898]" name="Saudi Arabia" pageId="1" pageNumber="2">
<emphasis id="B977A7484A456D7656160A2FF010F8D5" bold="true" box="[888,1048,1873,1898]" pageId="1" pageNumber="2">Saudi Arabia</emphasis>
</collectingCountry>
:
<specimenCount id="9D05B0D34A456D7651490A2FF07EF8D6" box="[1063,1142,1873,1897]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
,
<specimenCount id="9D05B0D34A456D7651EC0A2FF0ECF8D6" box="[1154,1252,1873,1897]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A456D76519D0A2FF152F8D6" box="[1267,1370,1873,1897]" httpUri="http://grbio.org/cool/0aes-7fp6" name="Saudi Arabian National Museum of Natural History" pageId="1" pageNumber="2">SNMNH</collectionCode>
156), Ash Sharghiyah, Karan I.,
<geoCoordinate id="EE371D9D4A456D7654A50A08F639F831" box="[459,561,1910,1934]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="27.716667">27º 43'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A456D7657510A08F6A9F831" box="[575,673,1910,1934]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="49.816666">49º 49'E</geoCoordinate>
, coll. M. Apel,
<date id="FFBD5D9A4A456D7656370A08F7D3F831" box="[857,987,1910,1934]" pageId="1" pageNumber="2" value="1992-07-17">17.07.1992</date>
;
<specimenCount id="9D05B0D34A456D7656860A08F031F831" box="[1000,1081,1910,1934]" pageId="1" pageNumber="2" type="male">1 male</specimenCount>
,
<specimenCount id="9D05B0D34A456D7651280A08F0A2F831" box="[1094,1194,1910,1934]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A456D7651D50A08F12AF831" box="[1211,1314,1910,1934]" httpUri="http://grbio.org/cool/0aes-7fp6" name="Saudi Arabian National Museum of Natural History" pageId="1" pageNumber="2">SNMNH</collectionCode>
157), Ash Sharghiyah, Karan I.,
<geoCoordinate id="EE371D9D4A456D7654F20AE2F60DF80B" box="[412,517,1948,1972]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="27.716667">27º 43'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A456D76577B0AE2F672F80B" box="[533,634,1948,1972]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="49.816666">49º 49'E</geoCoordinate>
, coll. M. Apel,
<date id="FFBD5D9A4A456D7656520AE2F7B6F80B" box="[828,958,1948,1972]" pageId="1" pageNumber="2" value="1995-05-20">20.05.1995</date>
;
<specimenCount id="9D05B0D34A456D7656A00AE2F024F80B" box="[974,1068,1948,1972]" pageId="1" pageNumber="2" type="male">2 males</specimenCount>
,
<specimenCount id="9D05B0D34A456D7651520AE2F0AAF80B" box="[1084,1186,1948,1972]" pageId="1" pageNumber="2" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A456D7651D80AE2F115F80B" box="[1206,1309,1948,1972]" httpUri="http://grbio.org/cool/0aes-7fp6" name="Saudi Arabian National Museum of Natural History" pageId="1" pageNumber="2">SNMNH</collectionCode>
158), Ash Sharghiyah, Karan I.,
<geoCoordinate id="EE371D9D4A456D7654E30ABFF5F9F866" box="[397,497,1985,2009]" direction="north" orientation="latitude" pageId="1" pageNumber="2" precision="925" value="27.716667">27º 43'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A456D7654920ABFF654F866" box="[508,604,1985,2009]" direction="east" orientation="longitude" pageId="1" pageNumber="2" precision="925" value="49.816666">49º 49'E</geoCoordinate>
, coll. M. Apel,
<date id="FFBD5D9A4A456D7656650ABFF784F866" box="[779,908,1985,2009]" pageId="1" pageNumber="2" value="1992-07-17">17.07.1992</date>
.
</paragraph>
<paragraph id="8BBC7B5A4A466D7555AB0DE9F7C7FDF5" blockId="2.[151,1436,151,1855]" pageId="2" pageNumber="3">
<collectionCode id="ED12E39F4A466D7555AB0DE9F506FF10" box="[197,270,151,175]" pageId="2" pageNumber="3">GULF</collectionCode>
OF
<collectingCountry id="F3143BCA4A466D75542E0DE9F591FF10" box="[320,409,151,175]" name="Oman" pageId="2" pageNumber="3">OMAN</collectingCountry>
,
<collectingCountry id="F3143BCA4A466D7554CA0DE6F5D1FF0F" box="[420,473,152,176]" name="Iran" pageId="2" pageNumber="3">
<emphasis id="B977A7484A466D7554CA0DE6F5D1FF0F" bold="true" box="[420,473,152,176]" pageId="2" pageNumber="3">Iran</emphasis>
</collectingCountry>
:
<specimenCount id="9D05B0D34A466D7554890DE9F640FF10" box="[487,584,151,175]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A466D7557390DE9F6AAFF10" box="[599,674,151,175]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach 1130), rocky shore, Gwadr, Baluchestan,
<geoCoordinate id="EE371D9D4A466D7551AD0DE9F121FF10" box="[1219,1321,151,175]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.133333">25° 08'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D75505B0DE9F19FFF10" box="[1333,1431,151,175]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="61.45">61° 27'E</geoCoordinate>
, coll. R. Naderloo &amp; A. Sari,
<date id="FFBD5D9A4A466D7554B90DC3F65FFF6A" box="[471,599,189,213]" pageId="2" pageNumber="3" value="2005-11-19">19.11.2005</date>
;
<specimenCount id="9D05B0D34A466D75570C0DC3F6B8FF6A" box="[610,688,189,213]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
,
<specimenCount id="9D05B0D34A466D7557D50DC3F6F1FF6A" box="[699,761,189,213]" pageId="2" pageNumber="3" type="generic">1 juv.</specimenCount>
(
<collectionCode id="ED12E39F4A466D7556680DC3F759FF6A" box="[774,849,189,213]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach 1179), rocky shore, Pasabandar, Baluchestan,
<geoCoordinate id="EE371D9D4A466D7555F90D9CF4F7FF45" box="[151,255,226,250]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.066668">25° 04'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D7554650D9CF578FF45" box="[267,368,226,250]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="61.4">61° 24'E</geoCoordinate>
, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D7557A60D9CF741FF45" box="[712,841,226,250]" pageId="2" pageNumber="3" value="2005-11-18">18.11.2005</date>
;
<specimenCount id="9D05B0D34A466D75563B0D9CF7B0FF45" box="[853,952,226,250]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A466D7556A60D9CF01BFF45" box="[968,1043,226,250]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach1132), rocky shore, Bandar- Ramin, Baluchestan,
<geoCoordinate id="EE371D9D4A466D7554E40C79F5FAFEA0" box="[394,498,263,287]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.266666">25° 16'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D7554910C79F665FEA0" box="[511,621,263,287]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="60.733334">60° 44' E</geoCoordinate>
, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D7556A60C79F040FEA0" box="[968,1096,263,287]" pageId="2" pageNumber="3" value="2005-11-18">18.11.2005</date>
;
<specimenCount id="9D05B0D34A466D75513B0C79F0A5FEA0" box="[1109,1197,263,287]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
(
<collectionCode id="ED12E39F4A466D7551D00C79F101FEA0" box="[1214,1289,263,287]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach1135), rocky shore, Hotel Lipar, Chabahar, Baluchestan,
<geoCoordinate id="EE371D9D4A466D7557A20C53F73CFEFA" box="[716,820,301,325]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.316668">25° 19'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D75562E0C53F7A4FEFA" box="[832,940,301,325]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="60.616665">60° 37' E</geoCoordinate>
, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D75506F0C53F189FEFA" box="[1281,1409,301,325]" pageId="2" pageNumber="3" value="2005-11-17">17.11.2005</date>
;
<specimenCount id="9D05B0D34A466D7550E00C53F4EBFED5" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A466D75559A0C2CF537FED5" box="[244,319,338,362]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Barch1175), rocky/cobble, Portuguese castle, Chabahar, Baluchestan,
<geoCoordinate id="EE371D9D4A466D75510B0C2CF0C3FED5" box="[1125,1227,338,362]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.266666">25º 16'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D7551B60C2CF132FED5" box="[1240,1338,338,362]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="60.616665">60º 37'E</geoCoordinate>
, coll. R. Naderloo &amp; A. Kazemi,
<date id="FFBD5D9A4A466D7554DE0C09F638FE30" box="[432,560,375,399]" pageId="2" pageNumber="3" value="2005-11-17">17.11.2005</date>
;
<specimenCount id="9D05B0D34A466D7557530C09F69DFE30" box="[573,661,375,399]" pageId="2" pageNumber="3" type="male">4 males</specimenCount>
(juv.) (
<collectionCode id="ED12E39F4A466D7557870C09F73CFE30" box="[745,820,375,399]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach1177), rocky shore, Chabahar, Baluchestan,
<geoCoordinate id="EE371D9D4A466D75501B0C09F4C6FE0A" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.266666">25° 16'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D7555B40CE3F534FE0A" box="[218,316,413,437]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="60.666668">60° 40'E</geoCoordinate>
, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D7557E80CE3F70FFE0A" box="[646,775,413,437]" pageId="2" pageNumber="3" value="2005-11-17">17.11.2005</date>
;
<specimenCount id="9D05B0D34A466D75567F0CE3F756FE0A" box="[785,862,413,437]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
,
<specimenCount id="9D05B0D34A466D7556070CE3F7C1FE0A" box="[873,969,413,437]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
(
<collectionCode id="ED12E39F4A466D7556B60CE3F02BFE0A" box="[984,1059,413,437]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach1133), sandy/rocky, shrimp aquaculture station of Gorgij, Bandar-Jask, Hormozgan, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D7551300CBCF0D6FE65" box="[1118,1246,450,474]" pageId="2" pageNumber="3" value="2005-11-16">16.11.2005</date>
;
<specimenCount id="9D05B0D34A466D7551840CBCF149FE65" box="[1258,1345,450,474]" pageId="2" pageNumber="3" type="male">2 males</specimenCount>
(
<collectionCode id="ED12E39F4A466D75503F0CBCF194FE65" box="[1361,1436,450,474]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach1134), sandy/rocky, fishery jetty, Sorgolm village, Bandar-Jask, Hormozgan,
<geoCoordinate id="EE371D9D4A466D7551540C99F095FE40" box="[1082,1181,487,511]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.583334">25º 35'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D7551C70C99F101FE40" box="[1193,1289,487,511]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="58.033333">58º 02'E</geoCoordinate>
, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D7554250F73F5C4FD9A" box="[331,460,525,549]" pageId="2" pageNumber="3" value="2005-11-21">21.11.2005</date>
;
<specimenCount id="9D05B0D34A466D7554B90F73F62DFD9A" box="[471,549,525,549]" pageId="2" pageNumber="3" type="male">1 male</specimenCount>
,
<specimenCount id="9D05B0D34A466D75575E0F73F69CFD9A" box="[560,660,525,549]" pageId="2" pageNumber="3" type="female">1 female</specimenCount>
,
<specimenCount id="9D05B0D34A466D7557CE0F73F6D7FD9A" box="[672,735,525,549]" pageId="2" pageNumber="3" type="generic">1 juv.</specimenCount>
(
<collectionCode id="ED12E39F4A466D7557820F73F73FFD9A" box="[748,823,525,549]" pageId="2" pageNumber="3">ZUTC</collectionCode>
Brach1157) rocky shore, Yekdar village, Bandar-Jask, Hormozgan,
<geoCoordinate id="EE371D9D4A466D7554470F4CF587FDF5" box="[297,399,562,586]" direction="north" orientation="latitude" pageId="2" pageNumber="3" precision="925" value="25.55">25° 33'N</geoCoordinate>
,
<geoCoordinate id="EE371D9D4A466D7554F40F4CF5F4FDF5" box="[410,508,562,586]" direction="east" orientation="longitude" pageId="2" pageNumber="3" precision="925" value="58.25">58° 15'E</geoCoordinate>
, coll. A. Sari &amp; R. Naderloo,
<date id="FFBD5D9A4A466D7556240F4CF7C3FDF5" box="[842,971,562,586]" pageId="2" pageNumber="3" value="2005-11-21">21.11.2005</date>
.
</paragraph>
</subSubSection>
<subSubSection id="C31928D14A466D7355AB0F29F0C2FEDB" lastPageId="4" lastPageNumber="5" pageId="2" pageNumber="3" type="description">
<paragraph id="8BBC7B5A4A466D7555AB0F29F664FC00" blockId="2.[151,1436,151,1855]" pageId="2" pageNumber="3">
<emphasis id="B977A7484A466D7555AB0F29F57BFDCF" bold="true" box="[197,371,599,624]" pageId="2" pageNumber="3">Redescription.</emphasis>
Carapace subcircular (
<figureCitation id="133867DF4A466D7557100F29F6C4FDD0" box="[638,716,599,623]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="2" pageNumber="3">Figs. 1</figureCitation>
a, 5a), slightly broader than long, about 1.1 times as broad as long, maximum breadth across middle part of carapace; carapace weakly convex; regions defined with shallow grooves; lateral regions with numerous curved ridges, getting longer posteriorly; transverse ridge along posterior region, parallel to posterior margin; short ridges on anterior half portion of carapace. Lateral margin of carapace remarkably arched (nearly straight in small specimens); 2 anterolateral teeth (including exorbital tooth), sharply triangular, separated by V-shaped notch; shallow groove extending obliquely from this notch towards cardiac region. Frontal region gently flexed downwards; 4-lobed, median 2 slightly larger than lateral 2; large tubercles on lobes, tubercles smaller posteriorly, large tubercles behind lateral lobes, small tubercles between frontal lobes, frontal edge; frontal ridge slightly wider than posterior margin of carapace, finely crenulated, nearly straight, slightly concave medially, with lateral angle rounded.
</paragraph>
<paragraph id="8BBC7B5A4A466D7555AB0EB3F00FFBC5" blockId="2.[151,1436,151,1855]" pageId="2" pageNumber="3">Orbits deep, U-shaped notch on supraorbital margin, separating inner from lateral part; granular elevation behind notch; outer part slightly granular. Infraorbital margin with inner angle acutely triangular; middle portion finely granular, with relatively large denticles on outer part, separated from outer angle by deep U-shaped notch; outer angle triangle with relatively large tooth on inner surface. Eyestalks short, proximal part wide, with granules on anterior face. Antennae remarkably short, flagellum not reaching to cornea.</paragraph>
<paragraph id="8BBC7B5A4A466D7555AB09F9F69AFB7A" blockId="2.[151,1436,151,1855]" pageId="2" pageNumber="3">
Third maxilliped (
<figureCitation id="133867DF4A466D7554F809F9F5DFFB20" box="[406,471,1159,1183]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
c) with ischium slightly longer than merus, 2 large spines on proximal part of inner margin of ischium, concealed by dense setae.
</paragraph>
<paragraph id="8BBC7B5A4A466D7555AB09ACF03FF985" blockId="2.[151,1436,151,1855]" pageId="2" pageNumber="3">
Chelipeds subequal, in comparison to walking legs relatively small, ischium with 3 or 4 spine-shaped teeth on anterior margin. Merus with spine-shaped teeth on anterior margin, normally larger distally; lower posterior margin granulate, granules dentiform distally; upper posterior margin serrate; outer surface with transverse ridges. Carpus with strong tooth on inner margin, granular on outer surface. Manus (
<figureCitation id="133867DF4A466D7556C6083CF7E1FAE5" box="[936,1001,1346,1370]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
b) with 2 long ridges on lower portion of outer surface; lower one longer, extending from tip of immovable finger to near articulation with carpus; upper margin with large granules, distal one long, tooth-shaped; lower margin with long ridges, becoming round tubercles distally to base of immovable finger; inner surface smooth in middle, transverse ridges on upper part, tubercles proximally on lower portion. Movable finger with large tubercles proximally on upper margin; cutting edge with remarkably large, round tooth on proximal part; cutting edge of immovable finger with 3 similar teeth on proximal part, short setae along inner surface of teeth; black tip of fingers not spoon-shaped.
</paragraph>
<paragraph id="8BBC7B5A4A466D7555AB0B39F046F880" blockId="2.[151,1436,151,1855]" pageId="2" pageNumber="3">Walking legs relatively strong, flattened. Merus relatively large; posterior surface with transverse ridges; anterior margin serrated, with subdistal spine; small spine between subdistal spines, distal notch of merus in third walking leg; 4 spines on posterodistal lobe, becoming smaller distally. Carpus with 2 carinae on posterior surface, anterior carina more prominent, short plumose setae along posterior one; anterior surface with inconspicuous carina. Propodus with strong, retractile bristle along anterior, posterior margins; plumose setae on posterior surface; one posterodistal spine; distal lobe with supplementary tooth. Dactylus with 2 rows of 6 spines along anterior margin; 2 rows of 4 or 5 spines along posterior margin; dactylus ending with large spine.</paragraph>
<caption id="DF7C2BD24A476D7455F90A71F75DF8DD" httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="3" pageNumber="4" targetBox="[224,1382,283,1701]" targetPageId="3">
<paragraph id="8BBC7B5A4A476D7455F90A71F75DF8DD" blockId="3.[151,1436,1807,1890]" pageId="3" pageNumber="4">
<emphasis id="B977A7484A476D7455F90A71F51CF89A" bold="true" box="[151,276,1807,1830]" pageId="3" pageNumber="4">FIGURE 1.</emphasis>
<taxonomicName id="4C0300D94A476D7454750A6EF65AF89A" authority="Latreille" authorityName="Latreille" box="[283,594,1807,1829]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="albolineatus">
<emphasis id="B977A7484A476D7454750A6EF5FDF89A" box="[283,501,1807,1829]" italics="true" pageId="3" pageNumber="4">Grapsus albolineatus</emphasis>
Latreille
</taxonomicName>
in Milbert, 1812, male (ZUTC Brach1175): ae; female (ZUTC Brach1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G1, lateral surface (right): f, apical part of the same G1; g, female gonopore (right).
</paragraph>
</caption>
<paragraph id="8BBC7B5A4A406D7355AB0DE9F6EAFF6B" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
<collectingCountry id="F3143BCA4A406D7355AB0DE9F4F7FF10" box="[197,255,151,175]" name="Malta" pageId="4" pageNumber="5">Male</collectingCountry>
abdomen (
<figureCitation id="133867DF4A406D7354EE0DE9F5CAFF10" box="[384,450,151,175]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
d) triangular; fifth somite longest, about 2.3 times as broad as long, sixth with lateral margin converging distally; telson clearly triangular.
</paragraph>
<paragraph id="8BBC7B5A4A406D7355AB0DA1F72EFE80" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
G1 (
<figureCitation id="133867DF4A406D7355960DA1F531FF48" box="[248,313,223,247]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
e) remarkably stout, nearly straight; outer margin concave; inner margin convex, posterior surface proximally elevated in middle part; apical chitinous process (
<figureCitation id="133867DF4A406D7356290C7AF78DFEA3" box="[839,901,260,284]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="4" pageNumber="5">Fig. 1</figureCitation>
f) small, directed outwards; gonopore terminal; long setae around apical part, obscuring chitinous process.
</paragraph>
<paragraph id="8BBC7B5A4A406D7355AB0C32F0C2FEDB" blockId="4.[151,1436,151,1040]" box="[197,1226,332,357]" pageId="4" pageNumber="5">
Female gonopore (
<figureCitation id="133867DF4A406D7354F90C32F5DEFEDB" box="[407,470,332,356]" captionStart="FIGURE 1" captionStartId="3.[151,250,1807,1830]" captionTargetBox="[224,1382,283,1701]" captionTargetId="figure@3.[164,1422,193,1786]" captionTargetPageId="3" captionText="FIGURE 1. Grapsus albolineatus Latreille in Milbert, 1812, male (ZUTC Brach 1175): a e; female (ZUTC Brach 1175): g. a, posterior surface of carapace; b, cheliped of male, outer surface; c, third maxilliped (right); d, male abdomen; e, G 1, lateral surface (right): f, apical part of the same G 1; g, female gonopore (right)." httpUri="https://zenodo.org/record/205388/files/figure.png" pageId="4" pageNumber="5">
Fig.
<quantity id="4CFBD6BF4A406D7354A50C32F5DEFEDB" box="[459,470,332,356]" metricMagnitude="-3" metricUnit="kg" metricValue="1.0" pageId="4" pageNumber="5" unit="g" value="1.0">1</quantity>
</figureCitation>
g) with elevated finger-like operculum, directed obliquely forward.
</paragraph>
</subSubSection>
<subSubSection id="C31928D14A406D7355AB0C0EF4D5FE70" pageId="4" pageNumber="5" type="discussion">
<paragraph id="8BBC7B5A4A406D7355AB0C0EF4D5FE70" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7355AB0C0EF531FE37" bold="true" box="[197,313,368,392]" pageId="4" pageNumber="5">Remarks.</emphasis>
<taxonomicName id="4C0300D94A406D73542E0C0EF639FE37" box="[320,561,368,392]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="albolineatus">
<emphasis id="B977A7484A406D73542E0C0EF639FE37" box="[320,561,368,392]" italics="true" pageId="4" pageNumber="5">Grapsus albolineatus</emphasis>
</taxonomicName>
has been referred to
<bibRefCitation id="EF9206AB4A406D73564E0C11F7D0FE38" author="Lamarck" box="[800,984,367,391]" pageId="4" pageNumber="5" refString="Lamarck, M. de (1818) Hombranches Brachyures. In: Les caracteres generaux et particulaires des ces animaux, leur distribution, leurs classes, leurs familles, leurs genres, et la citation des principales especes qui s'y rapportent. Tome 5. Deuxieme section, 266 - 273." type="journal article" year="1818">Lamarck (1818)</bibRefCitation>
, but Ng
<emphasis id="B977A7484A406D7351550C0FF065FE37" box="[1083,1133,368,392]" italics="true" pageId="4" pageNumber="5">et al</emphasis>
. (2008) noted that the correct authority should be Latreille, in Milbert (1812).
<bibRefCitation id="EF9206AB4A406D7357920CEAF7B9FE13" author="Holthuis" box="[764,945,404,428]" pageId="4" pageNumber="5" refString="Holthuis, L. B. (1977) The Grapsidae, Gecarcinidae and Palicidae (Crustacea: Decapoda: Brachyura) of the Red Sea. Israel Journal of Zoology, 26 (3 - 4), 141 - 192." type="journal article" year="1977">Holthuis (1977)</bibRefCitation>
has discussed the validity of this name in detail.
</paragraph>
</subSubSection>
<subSubSection id="C31928D14A406D7355AB0CA5F0A5FBB0" pageId="4" pageNumber="5" type="biology_ecology">
<paragraph id="8BBC7B5A4A406D7355AB0CA5F69BFD18" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7355AB0CA5F52CFE4B" bold="true" box="[197,292,475,500]" pageId="4" pageNumber="5">Biology.</emphasis>
<taxonomicName id="4C0300D94A406D7354450CA3F613FE4B" box="[299,539,476,500]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="albolineatus">
<emphasis id="B977A7484A406D7354450CA3F613FE4B" box="[299,539,476,500]" italics="true" pageId="4" pageNumber="5">Grapsus albolineatus</emphasis>
</taxonomicName>
is a relatively large-size species (largest male CL =
<quantity id="4CFBD6BF4A406D73510C0CA2F0DEFE4B" box="[1122,1238,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="5.027" pageId="4" pageNumber="5" unit="mm" value="50.27">50.27 mm</quantity>
, CB =
<quantity id="4CFBD6BF4A406D73504A0CA2F19FFE4B" box="[1316,1431,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="5.477" pageId="4" pageNumber="5" unit="mm" value="54.77">54.77 mm</quantity>
, largest female CL =
<quantity id="4CFBD6BF4A406D7354EE0C81F5FDFDA8" box="[384,501,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="5.914" pageId="4" pageNumber="5" unit="mm" value="59.14">59.14 mm</quantity>
, CB =
<quantity id="4CFBD6BF4A406D7357280C81F6B1FDA8" box="[582,697,511,535]" metricMagnitude="-2" metricUnit="m" metricValue="6.38" pageId="4" pageNumber="5" unit="mm" value="63.8">63.80 mm</quantity>
) and is the most common species on exposed rocky shores in the Gulf of
<collectingCountry id="F3143BCA4A406D73559F0F5AF53DFD83" box="[241,309,548,572]" name="Oman" pageId="4" pageNumber="5">Oman</collectingCountry>
and the Persian Gulf, where it is present in large populations on the high rocky intertidal. This species is sympatric with
<taxonomicName id="4C0300D94A406D7354FF0F37F69DFDDF" box="[401,661,585,608]" class="Malacostraca" family="Grapsidae" genus="Metopograpsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="messor">
<emphasis id="B977A7484A406D7354FF0F37F69DFDDF" box="[401,661,585,608]" italics="true" pageId="4" pageNumber="5">Metopograpsus messor</emphasis>
</taxonomicName>
in sheltered rocky shores in the Persian Gulf and in some locations it is replaced by the latter.
<taxonomicName id="4C0300D94A406D7354C50F13F694FD3B" box="[427,668,620,644]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="albolineatus">
<emphasis id="B977A7484A406D7354C50F13F694FD3B" box="[427,668,620,644]" italics="true" pageId="4" pageNumber="5">Grapsus albolineatus</emphasis>
</taxonomicName>
is typical of exposed rocky shores and is normally seen foraging on algae, disappearing into crevices if disturbed.
</paragraph>
<paragraph id="8BBC7B5A4A406D7355AB0FCDF55BFCAB" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7355AB0FCDF515FD73" bold="true" box="[197,285,691,716]" pageId="4" pageNumber="5">Colour.</emphasis>
Brownish red (
<figureCitation id="133867DF4A406D7354A40FCAF606FD73" box="[458,526,692,716]" captionStart="FIGURE 4" captionStartId="11.[151,250,1918,1941]" captionTargetBox="[199,1365,236,1846]" captionTargetId="figure@11.[151,1436,193,1897]" captionTargetPageId="11" captionText="FIGURE 4. Metopograpsus thukuhar (Owen, 1839), male (ZUTC Brach 1162), a f; female (ZUTC Brach 1162): g. a, posterior surface of carapace; b, lower orbital margin of male; c, cheliped of male, outer surface; d, third walking leg of male (left); e, male abdomen; f, G 1, lateral surface (right); g, female gonopore (right)." httpUri="https://zenodo.org/record/205391/files/figure.png" pageId="4" pageNumber="5">Fig. 4</figureCitation>
a), sometimes bluish carapace, walking legs light red or orange, ventral side light, mottled with reddish orange, anterior surface of the palms light. The colour pattern was described in detail by
<bibRefCitation id="EF9206AB4A406D7355F90F82F547FCAB" author="Banerjee" box="[151,335,764,788]" pageId="4" pageNumber="5" refString="Banerjee, S. K. (1960) Biological results of Snellius Expedition 18. The genera Grapsus, Geograpsus and Metopograpsus (Crustacea: Brachyura). Temminckia, 10, 132 - 199." type="journal article" year="1960">Banerjee (1960)</bibRefCitation>
.
</paragraph>
<paragraph id="8BBC7B5A4A406D7355AB0E61F7BCFC1B" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7355AB0E61F59FFC87" bold="true" box="[197,407,799,824]" pageId="4" pageNumber="5">Regional records.</emphasis>
PERSIAN GULF:
<collectingCountry id="F3143BCA4A406D73571C0E61F6A5FC87" box="[626,685,799,824]" name="United Arab Emirates" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D73571C0E61F6A5FC87" bold="true" box="[626,685,799,824]" pageId="4" pageNumber="5">UAE</emphasis>
</collectingCountry>
[
<bibRefCitation id="EF9206AB4A406D7357D50E61F750FC88" author="Titgen" box="[699,856,799,823]" pageId="4" pageNumber="5" refString="Titgen, R. H. (1982) The Systematics and Ecology of the Decapods of Dubai, and their Zoogeographic Relationships to the Persian Gulf and the Western Indian Ocean. Doctoral Dissertation, Texas A &amp; M University." type="book" year="1982">Titgen (1982)</bibRefCitation>
from Dubai;
<bibRefCitation id="EF9206AB4A406D73569F0E61F071FC88" author="Apel" box="[1009,1145,799,823]" pageId="4" pageNumber="5" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes, pp. 1 - 268. Unpublished doctoral dissertation, Johann Wolfgang Goethe-Universitat, Frankfurt am Main, Germany, 260 pp." type="book chapter" year="2001">Apel (2001)</bibRefCitation>
from Abu Dhabi and Ras al-Khaimah],
<collectingCountry id="F3143BCA4A406D73545B0E3DF5DFFCE3" box="[309,471,835,860]" name="Saudi Arabia" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D73545B0E3DF5DFFCE3" bold="true" box="[309,471,835,860]" pageId="4" pageNumber="5">Saudi Arabia</emphasis>
</collectingCountry>
(
<bibRefCitation id="EF9206AB4A406D7354860E3AF66DFCE3" author="Apel" box="[488,613,836,860]" pageId="4" pageNumber="5" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes, pp. 1 - 268. Unpublished doctoral dissertation, Johann Wolfgang Goethe-Universitat, Frankfurt am Main, Germany, 260 pp." type="book chapter" year="2001">Apel 2001</bibRefCitation>
),
<collectingCountry id="F3143BCA4A406D7357150E3AF6B8FCE3" box="[635,688,836,860]" name="Iran" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7357150E3AF6B8FCE3" bold="true" box="[635,688,836,860]" pageId="4" pageNumber="5">Iran</emphasis>
</collectingCountry>
[
<bibRefCitation id="EF9206AB4A406D7357AE0E3AF79EFCE3" author="Stephensen" box="[704,918,836,860]" pageId="4" pageNumber="5" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen (1945)</bibRefCitation>
as
<taxonomicName id="4C0300D94A406D7356AE0E3BF094FCE3" box="[960,1180,836,860]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B977A7484A406D7356AE0E3BF094FCE3" box="[960,1180,836,860]" italics="true" pageId="4" pageNumber="5">Grapsus maculatus</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="4C0300D94A406D73519B0E3BF19CFCE3" box="[1269,1428,837,860]" class="Malacostraca" family="Cancridae" genus="Cancer" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A406D73519B0E3BF19CFCE3" box="[1269,1428,837,860]" italics="true" pageId="4" pageNumber="5">tenuicrustatus</emphasis>
</taxonomicName>
;
<bibRefCitation id="EF9206AB4A406D7355F90E17F557FCC0" author="Pretzmann" box="[151,351,871,896]" pageId="4" pageNumber="5" refString="Pretzmann, G. (1971) Ergebnisse einiger Sammelreisen nach Vorderasien. 2. Teil: Marine Brachyura. Annalen des Naturhistorischen Museums Wien, 75, 477 - 487." type="journal article" year="1971">Pretzmann (1971)</bibRefCitation>
as
<taxonomicName id="4C0300D94A406D7354EA0E16F68FFC3F" box="[388,647,872,896]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A406D7354EA0E16F68FFC3F" box="[388,647,872,896]" italics="true" pageId="4" pageNumber="5">Grapsus tenuicrustatus</emphasis>
</taxonomicName>
; present study]. GULF OF
<collectingCountry id="F3143BCA4A406D7356D50E19F01DFCC0" box="[955,1045,871,895]" name="Oman" pageId="4" pageNumber="5">OMAN</collectingCountry>
:
<collectingCountry id="F3143BCA4A406D7351700E16F05BFC3F" box="[1054,1107,872,896]" name="Iran" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7351700E16F05BFC3F" bold="true" box="[1054,1107,872,896]" pageId="4" pageNumber="5">Iran</emphasis>
</collectingCountry>
[
<bibRefCitation id="EF9206AB4A406D73510E0E19F13BFCC0" author="Stephensen" box="[1120,1331,871,895]" pageId="4" pageNumber="5" refString="Stephensen, K. (1945) The Brachyura of the Iranian Gulf. Danish Scientific Investigations in Iran, Part IV. E. Munksgaard, Copenhagen, 57 - 237." type="book chapter" year="1945">Stephensen (1945)</bibRefCitation>
as
<taxonomicName id="4C0300D94A406D7350390E16F53DFC1B" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="maculatus">
<emphasis id="B977A7484A406D7350390E16F53DFC1B" italics="true" pageId="4" pageNumber="5">Grapsus maculatus</emphasis>
</taxonomicName>
subsp.
<taxonomicName id="4C0300D94A406D7354E70EF3F620FC1B" box="[393,552,909,932]" class="Malacostraca" family="Grapsidae" genus="Grapsus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="tenuicrustatus">
<emphasis id="B977A7484A406D7354E70EF3F620FC1B" box="[393,552,909,932]" italics="true" pageId="4" pageNumber="5">tenuicrustatus</emphasis>
</taxonomicName>
; present study],
<collectingCountry id="F3143BCA4A406D73578D0EF5F717FC1B" box="[739,799,907,932]" name="United Arab Emirates" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D73578D0EF5F717FC1B" bold="true" box="[739,799,907,932]" pageId="4" pageNumber="5">UAE</emphasis>
</collectingCountry>
(
<bibRefCitation id="EF9206AB4A406D7356420EF2F7A0FC1B" author="Apel" box="[812,936,908,932]" pageId="4" pageNumber="5" refString="Apel, M. (2001) Taxonomie und Zoogeographie der Brachyura, Paguridea und Porcellanidae (Crustacea: Decapoda) des Persisch-Arabischen Golfes, pp. 1 - 268. Unpublished doctoral dissertation, Johann Wolfgang Goethe-Universitat, Frankfurt am Main, Germany, 260 pp." type="book chapter" year="2001">Apel 2001</bibRefCitation>
).
</paragraph>
<paragraph id="8BBC7B5A4A406D7355AB0ED1F0A5FBB0" blockId="4.[151,1436,151,1040]" pageId="4" pageNumber="5">
<emphasis id="B977A7484A406D7355AB0ED1F60DFC77" bold="true" box="[197,517,943,968]" pageId="4" pageNumber="5">Geographical distribution.</emphasis>
Indo-West Pacific:
<collectingCountry id="F3143BCA4A406D7357870ED1F751FC78" box="[745,857,943,967]" name="Mauritius" pageId="4" pageNumber="5">Mauritius</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D7356090ED1F7CDFC78" box="[871,965,943,967]" name="Somalia" pageId="4" pageNumber="5">Somalia</collectingCountry>
, Red Sea, Gulf of Aden,
<collectingCountry id="F3143BCA4A406D7351850ED1F14BFC78" box="[1259,1347,943,967]" name="Yemen" pageId="4" pageNumber="5">Socotra</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D73503E0ED1F19FFC78" box="[1360,1431,943,967]" name="Oman" pageId="4" pageNumber="5">Oman</collectingCountry>
, Persian Gulf, Gulf of
<collectingCountry id="F3143BCA4A406D7354FD0EAAF5D2FC53" box="[403,474,980,1004]" name="Oman" pageId="4" pageNumber="5">Oman</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D7354860EAAF642FC53" box="[488,586,980,1004]" name="Pakistan" pageId="4" pageNumber="5">Pakistan</collectingCountry>
, Arabian Sea coast of
<collectingCountry id="F3143BCA4A406D73563E0EAAF784FC53" box="[848,908,980,1004]" name="India" pageId="4" pageNumber="5">India</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D7356F70EAAF004FC53" box="[921,1036,980,1004]" name="Sri Lanka" pageId="4" pageNumber="5">Sri Lanka</collectingCountry>
, Bay of Bengal, Andaman Islands,
<collectingCountry id="F3143BCA4A406D7355F90E89F542FBB0" box="[151,330,1015,1039]" name="India" pageId="4" pageNumber="5">Nicobar Islands</collectingCountry>
, Mergui Archipelago,
<collectingCountry id="F3143BCA4A406D7357280E89F6BEFBB0" box="[582,694,1015,1039]" name="Indonesia" pageId="4" pageNumber="5">Indonesia</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D7357AF0E89F73DFBB0" box="[705,821,1015,1039]" name="Singapore" pageId="4" pageNumber="5">Singapore</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D73562F0E89F78FFBB0" box="[833,903,1015,1039]" name="China" pageId="4" pageNumber="5">China</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D7356FC0E87F7DDFBAF" box="[914,981,1017,1040]" name="Japan" pageId="4" pageNumber="5">Japan</collectingCountry>
,
<collectingCountry id="F3143BCA4A406D7356B10E89F040FBB0" box="[991,1096,1015,1039]" name="Australia" pageId="4" pageNumber="5">Australia</collectingCountry>
, Hawaii.
</paragraph>
</subSubSection>
</treatment>
</document>