treatments-xml/data/03/AA/CA/03AACA29C625FF848154FC6BFC3E04F2.xml
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<document id="8CFD53E9CE303C0155C973D66773604A" ID-DOI="10.5281/zenodo.5312352" ID-ISSN="0374-1036" ID-Zenodo-Dep="5312352" ID-ZooBank="2C4DD1B7-A32F-41FF-B866-3AC74B3083EC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630119354752" checkinUser="marcus" docAuthor="Kočárek, Petr" docDate="2014" docId="03AACA29C625FF848154FC6BFC3E04F2" docLanguage="en" docName="ActaEntMusNatPra.2014.54.suppl.1-22.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 54" docStyle="DocumentStyle:E9BE312A6EDF9B84FC5211AAA80D80BF.10:ActaEntMusNatPra.2005-2016.journal_article" docStyleId="E9BE312A6EDF9B84FC5211AAA80D80BF" docStyleName="ActaEntMusNatPra.2005-2016.journal_article" docStyleVersion="10" docTitle="Guanchia bituberculata" docType="treatment" docVersion="4" lastPageNumber="16" masterDocId="FF93B251C628FF8B8069FFFBFFAF0541" masterDocTitle="Earwigs (Dermaptera) of Socotra Island: checklist, distribution, and description of a new genus and four new species" masterLastPageNumber="21" masterPageNumber="1" pageNumber="14" updateTime="1699159095649" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="D0BA980AC0BF7D405012462E8DC76C01">Earwigs (Dermaptera) of Socotra Island: checklist, distribution, and description of a new genus and four new species</mods:title>
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<mods:namePart id="4C36B78B6C87F1D986E4BE21AB9686D0">Kočárek, Petr</mods:namePart>
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<mods:title id="A05F32298D3945400262F8A1E459312A">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="9F90990DE4B8762C0765015B1A369E8F">2014</mods:date>
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<treatment id="03AACA29C625FF848154FC6BFC3E04F2" ID-DOI="http://doi.org/10.5281/zenodo.5449251" ID-Zenodo-Dep="5449251" LSID="urn:lsid:plazi:treatment:03AACA29C625FF848154FC6BFC3E04F2" httpUri="http://treatment.plazi.org/id/03AACA29C625FF848154FC6BFC3E04F2" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<subSubSection id="C31928B4C625FF868154FC6BFCDA06E9" box="[317,885,912,936]" pageId="13" pageNumber="14" type="nomenclature">
<paragraph id="8BBC7B3FC625FF868154FC6BFCDA06E9" blockId="13.[317,885,912,936]" box="[317,885,912,936]" pageId="13" pageNumber="14">
<heading id="D0F4CC53C625FF868154FC6BFCDA06E9" bold="true" box="[317,885,912,936]" centered="true" fontSize="8" level="2" pageId="13" pageNumber="14" reason="3">
<emphasis id="B977A72DC625FF868154FC6BFCDA06E9" bold="true" box="[317,885,912,936]" pageId="13" pageNumber="14">
<taxonomicName id="4C0300BCC625FF868154FC6BFD5A06E9" authority="(Brindle, 1966)" baseAuthorityName="Brindle" baseAuthorityYear="1966" box="[317,757,912,936]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF868154FC6BFDEA06E9" bold="true" box="[317,581,912,936]" italics="true" pageId="13" pageNumber="14">Guanchia bituberculata</emphasis>
(
<bibRefCitation id="EF9206CEC625FF86823BFC6AFD4106E9" author="BRINDLE A." box="[594,750,913,936]" pageId="13" pageNumber="14" pagination="29 - 49" refId="ref10820" refString="BRINDLE A. 1966: Dermaptera from Central and East Africa. Revue de Zoologie et de Botanique Africaine 74: 29 - 49." type="journal article" year="1966">Brindle, 1966</bibRefCitation>
)
</taxonomicName>
stat. restit.
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF86824BFC54FD3F0683" box="[546,656,943,962]" pageId="13" pageNumber="14" type="description">
<paragraph id="8BBC7B3FC625FF86824BFC54FD3F0683" blockId="13.[113,656,943,1069]" box="[546,656,943,962]" pageId="13" pageNumber="14">
(
<figureCitation id="133867BAC625FF868241FC54FD250683" box="[552,650,943,962]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="13" pageNumber="14">Figs 2426</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF868018FC31FE59016C" pageId="13" pageNumber="14" type="reference_group">
<paragraph id="8BBC7B3FC625FF868018FC31FE72069C" blockId="13.[113,656,943,1069]" box="[113,477,970,989]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB135C11C625FF868018FC31FE72069C" box="[113,477,970,989]" pageId="13" pageNumber="14">
<treatmentCitation id="0AA25D2EC625FF868018FC31FE75069C" author="BRINDLE A." box="[113,474,970,989]" page="43" pageId="13" pageNumber="14" year="1966">
<taxonomicName id="4C0300BCC625FF868018FC31FE75069C" authority="Brindle, 1966: 43" authorityName="Brindle" authorityPageNumber="43" authorityYear="1966" box="[113,474,970,989]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF868018FC31FE93069C" box="[113,316,970,989]" italics="true" pageId="13" pageNumber="14">Anechura bituberculata</emphasis>
<bibRefCitation id="EF9206CEC625FF868128FC31FE75069C" author="BRINDLE A." box="[321,474,970,989]" pageId="13" pageNumber="14" pagination="29 - 49" refId="ref10820" refString="BRINDLE A. 1966: Dermaptera from Central and East Africa. Revue de Zoologie et de Botanique Africaine 74: 29 - 49." type="journal article" year="1966">Brindle, 1966: 43</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBC7B3FC625FF868018FC1EFE5706B9" blockId="13.[113,656,943,1069]" box="[113,504,997,1016]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB135C11C625FF868018FC1EFE5706B9" box="[113,504,997,1016]" pageId="13" pageNumber="14">
<treatmentCitation id="0AA25D2EC625FF868018FC1EFE5A06B9" author="BRINDLE A." box="[113,501,997,1016]" page=")" pageId="13" pageNumber="14" year="1973">
<taxonomicName id="4C0300BCC625FF868018FC1EFE5A06B9" authority=": BRINDLE (1973: 261)" authorityName="BRINDLE" authorityPageNumber="261" authorityYear="1973" box="[113,501,997,1016]" class="Insecta" family="Forficulidae" genus="Forficula" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF868018FC1EFE9706B9" box="[113,312,997,1016]" italics="true" pageId="13" pageNumber="14">Forficula bituberculata</emphasis>
:
<bibRefCitation id="EF9206CEC625FF86812AFC1EFE5A06B9" author="BRINDLE A." box="[323,501,997,1016]" pageId="13" pageNumber="14" pagination="1 - 335" refId="ref10845" refString="BRINDLE A. 1973: The Dermaptera of Africa. Pt. 1. Annales du Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques 205: 1 - 335." type="journal article" year="1973">BRINDLE (1973: 261)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BBC7B3FC625FF868018FC04FDBB0153" blockId="13.[113,656,943,1069]" box="[113,532,1023,1042]" pageId="13" pageNumber="14">
<taxonomicName id="4C0300BCC625FF868018FC04FDBF0153" authority=": STEINMANN (1993: 560)" authorityName="STEINMANN" authorityPageNumber="560" authorityYear="1993" box="[113,528,1023,1042]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF868018FC04FE920153" box="[113,317,1023,1042]" italics="true" pageId="13" pageNumber="14">Guanchia bituberculata</emphasis>
: STEINMANN (1993: 560)
</taxonomicName>
.
</paragraph>
<paragraph id="8BBC7B3FC625FF868018FBE1FE59016C" blockId="13.[113,656,943,1069]" box="[113,502,1050,1069]" pageId="13" pageNumber="14">
<treatmentCitationGroup id="AB135C11C625FF868018FBE1FE59016C" box="[113,502,1050,1069]" pageId="13" pageNumber="14">
<treatmentCitation id="0AA25D2EC625FF868018FBE1FE5D016C" author="HAAS F. &amp; POHL H. &amp; WRANIK W." box="[113,498,1050,1069]" page=")" pageId="13" pageNumber="14" year="2004">
<taxonomicName id="4C0300BCC625FF868018FBE1FE5D016C" ID-CoL="DVVJ" authority=": HAAS et al. (2004: 413)" authorityName="HAAS" authorityPageNumber="413" authorityYear="2004" box="[113,498,1050,1069]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868018FBE1FEB0016C" box="[113,287,1050,1069]" italics="true" pageId="13" pageNumber="14">Anechura sokotrana</emphasis>
:
<bibRefCitation id="EF9206CEC625FF868143FBE1FE5D016C" author="HAAS F. &amp; POHL H. &amp; WRANIK W." box="[298,498,1050,1069]" pageId="13" pageNumber="14" pagination="409 - 419" refId="ref11078" refString="HAAS F., POHL H. &amp; WRANIK W. 2004: Dermaptera of the Socotra Archipelago with the description of a new species. Fauna of Arabia 20: 409 - 419." type="journal article" year="2004">HAAS et al. (2004: 413)</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF868018FBBFFCD4006D" pageId="13" pageNumber="14" type="materials_examined">
<paragraph id="8BBC7B3FC625FF868018FBBFFD1B0116" blockId="13.[113,1089,1092,1324]" box="[113,692,1092,1111]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FBBFFF450116" bold="true" box="[113,234,1092,1111]" pageId="13" pageNumber="14">
<typeStatus id="54B8C59DC625FF868018FBBFFF310116" box="[113,158,1092,1111]" pageId="13" pageNumber="14">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F3143BAFC625FF868086FBBFFE870116" box="[239,296,1092,1111]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">Congo</collectingCountry>
[=
<collectingCountry id="F3143BAFC625FF86812FFBBFFDDA0116" box="[326,629,1092,1111]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">Democratic Republic of the Congo</collectingCountry>
],
<collectingRegion id="49C7B5DDC625FF8682EBFBBFFD000116" box="[642,687,1092,1111]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="13" pageNumber="14">Kivu</collectingRegion>
.
</paragraph>
<paragraph id="8BBC7B3FC625FF868018FBA5FCD4006D" blockId="13.[113,1089,1092,1324]" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FBA5FF590130" bold="true" box="[113,246,1118,1137]" pageId="13" pageNumber="14">Type material.</emphasis>
<typeStatus id="54B8C59DC625FF868092FBA5FEE10131" box="[251,334,1118,1137]" pageId="13" pageNumber="14" type="holotype">HOLOTYPE</typeStatus>
: J,
<typeStatus id="54B8C59DC625FF86811EFBA5FE550130" box="[375,506,1118,1137]" pageId="13" pageNumber="14" type="holotype">HOLOTYPUS</typeStatus>
// I.
<collectionCode id="ED12E3FAC625FF868272FBA5FD830130" box="[539,556,1118,1137]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="14">R</collectionCode>
.S.A.C.-MUS.
<collectingCountry id="F3143BAFC625FF8682C0FBA5FD5A0130" box="[681,757,1118,1137]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">CONGO</collectingCountry>
/
<collectingRegion id="49C7B5DDC625FF86836FFBA5FC9B0130" box="[774,820,1118,1137]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="13" pageNumber="14">Kivu</collectingRegion>
: Terr. Uvira, Ht / Luvubu,
<quantity id="4CFBD6DAC625FF86847FFBA5FF2B01CD" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="14" unit="m" value="2750.0">2750 m</quantity>
. (humus) / N. Leleup
<date id="FFBD5DFFC625FF868129FB82FED001CD" box="[320,383,1145,1164]" pageId="13" pageNumber="14" value="1954-05">V-1954</date>
//
<taxonomicName id="4C0300BCC625FF8681FCFB82FE4701CD" authorityName="Scudder" authorityYear="1876" box="[405,488,1145,1164]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Anechura</taxonomicName>
/
<taxonomicName id="4C0300BCC625FF868193FB82FD0001CD" authority="Brindle" authorityName="Brindle" box="[506,687,1145,1164]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">bituberculata Brindle</taxonomicName>
/ det. A. Brindle /
<typeStatus id="54B8C59DC625FF868326FB82FC3501CD" box="[847,922,1145,1164]" pageId="13" pageNumber="14" type="holotype">holotype</typeStatus>
// Type // Digitised by / F. Haas for / SYNTHESIS / Aug. 06 //
<collectionCode id="ED12E3FAC625FF86818FFB6FFD8A01E6" box="[486,549,1172,1191]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="14" type="Museum">MRAC</collectionCode>
ENT 000017410 (
<collectionCode id="ED12E3FAC625FF8682A5FB6FFCBF01E6" box="[716,784,1172,1191]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="14" type="Museum">MRAC</collectionCode>
).
<typeStatus id="54B8C59DC625FF868377FB6FFCD901E4" box="[798,886,1172,1191]" pageId="13" pageNumber="14" type="paratype">PARATYPES</typeStatus>
:
<specimenCount id="9D05B0B6C625FF8683E8FB6FFC0F01E7" box="[897,928,1165,1191]" pageId="13" pageNumber="14" type="female">1 ♀</specimenCount>
,
<typeStatus id="54B8C59DC625FF8683C6FB6FFB8001E6" box="[943,1071,1172,1191]" pageId="13" pageNumber="14" type="allotype">ALLOTYPUS</typeStatus>
// I.
<collectionCode id="ED12E3FAC625FF868015FB55FF220180" box="[124,141,1198,1217]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="14">R</collectionCode>
.S.A.C.-MUS.
<collectingCountry id="F3143BAFC625FF868162FB55FEF80180" box="[267,343,1198,1217]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">CONGO</collectingCountry>
/
<collectingRegion id="49C7B5DDC625FF868100FB55FE380180" box="[361,407,1198,1217]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="13" pageNumber="14">Kivu</collectingRegion>
: Terr. Uvira, Ht / Luvubu,
<quantity id="4CFBD6DAC625FF868215FB55FD6F0180" box="[636,704,1198,1217]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="14" unit="m" value="2750.0">2750 m</quantity>
. (humus) / N. Leleup
<date id="FFBD5DFFC625FF868314FB55FC130180" box="[893,956,1198,1217]" pageId="13" pageNumber="14" value="1954-05">V-1954</date>
// Allo / type //
<taxonomicName id="4C0300BCC625FF868018FB32FF6B019D" authorityName="Scudder" authorityYear="1876" box="[113,196,1225,1244]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Anechura</taxonomicName>
/
<taxonomicName id="4C0300BCC625FF8680BCFB32FE26019D" authority="Brindle" authorityName="Brindle" box="[213,393,1225,1244]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">bituberculata Brindle</taxonomicName>
/ det. A. Brindle /
<typeStatus id="54B8C59DC625FF86824EFB32FDC3019D" box="[551,620,1225,1244]" pageId="13" pageNumber="14" type="allotype">allotype</typeStatus>
// Digitised by / F. Haas for / SYNTHESIS / Aug. 06 //
<collectionCode id="ED12E3FAC625FF868018FB1FFF1F01B6" box="[113,176,1252,1271]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="14" type="Museum">MRAC</collectionCode>
ENT 000017410 (
<collectionCode id="ED12E3FAC625FF86813CFB1FFE3601B6" box="[341,409,1252,1271]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="14" type="Museum">MRAC</collectionCode>
); 1 J,
<typeStatus id="54B8C59DC625FF8681BDFB1FFD8301B6" box="[468,556,1252,1271]" pageId="13" pageNumber="14" type="paratype">Paratypus</typeStatus>
// I.
<collectionCode id="ED12E3FAC625FF868224FB1FFDF101B6" box="[589,606,1252,1271]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="14">R</collectionCode>
.S.A.C.-MUS.
<collectingCountry id="F3143BAFC625FF8682B2FB1FFC8801B6" box="[731,807,1252,1271]" name="Democratic Republic of the Congo" pageId="13" pageNumber="14">CONGO</collectingCountry>
/
<collectingRegion id="49C7B5DDC625FF86835EFB1FFCCA01B6" box="[823,869,1252,1271]" country="Democratic Republic of the Congo" name="Sud-Kivu" pageId="13" pageNumber="14">Kivu</collectingRegion>
: Terr. Uvira, Ht / Luvubu,
<quantity id="4CFBD6DAC625FF868018FB05FF150050" box="[113,186,1278,1297]" metricMagnitude="3" metricUnit="m" metricValue="2.75" pageId="13" pageNumber="14" unit="m" value="2750.0">2,750 m</quantity>
. (humus) / N. Leleup
<date id="FFBD5DFFC625FF86811DFB05FE1D0050" box="[372,434,1278,1297]" pageId="13" pageNumber="14" value="1954-05">V-1954</date>
// Para- / type //
<taxonomicName id="4C0300BCC625FF868228FB05FD3B0050" authorityName="Scudder" authorityYear="1876" box="[577,660,1278,1297]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="genus">Anechura</taxonomicName>
/
<taxonomicName id="4C0300BCC625FF8682CDFB05FCF60050" authority="Brindle" authorityName="Brindle" box="[676,857,1278,1297]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">bituberculata Brindle</taxonomicName>
/ det. A. Brindle /
<typeStatus id="54B8C59DC625FF86839EFB05FBEF0050" box="[1015,1088,1278,1297]" pageId="13" pageNumber="14" type="paratype">paratype</typeStatus>
// Digitised by / F. Haas for / SYNTHESIS / Aug. 06 //
<collectionCode id="ED12E3FAC625FF868221FAE2FD28006D" box="[584,647,1305,1324]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="14" type="Museum">MRAC</collectionCode>
ENT 000017410 (
<collectionCode id="ED12E3FAC625FF868344FAE2FCDE006D" box="[813,881,1305,1324]" country="Belgium" httpUri="http://grbio.org/cool/z14b-bb31" name="Musée Royal de lAfrique Centrale" pageId="13" pageNumber="14" type="Museum">MRAC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C31928B4C625FF848018FABEFE9D04D2" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14" type="discussion">
<paragraph id="8BBC7B3FC625FF848018FABEFE9D04D2" blockId="13.[113,1090,1349,1596]" lastBlockId="15.[113,1089,220,435]" lastPageId="15" lastPageNumber="16" pageId="13" pageNumber="14">
<emphasis id="B977A72DC625FF868018FABEFF41001D" bold="true" box="[113,238,1349,1372]" pageId="13" pageNumber="14">Comments.</emphasis>
<bibRefCitation id="EF9206CEC625FF86809BFABEFE0B001D" author="HAAS F. &amp; POHL H. &amp; WRANIK W." box="[242,420,1349,1372]" pageId="13" pageNumber="14" pagination="409 - 419" refId="ref11078" refString="HAAS F., POHL H. &amp; WRANIK W. 2004: Dermaptera of the Socotra Archipelago with the description of a new species. Fauna of Arabia 20: 409 - 419." type="journal article" year="2004">HAAS et al. (2004)</bibRefCitation>
synonymised
<taxonomicName id="4C0300BCC625FF86825AFABEFC8C001D" authorityName="Brindle" authorityYear="1966" box="[563,803,1349,1372]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF86825AFABEFC8C001D" box="[563,803,1349,1372]" italics="true" pageId="13" pageNumber="14">Anechura bituberculata</emphasis>
</taxonomicName>
with
<taxonomicName id="4C0300BCC625FF868331FABEFC76001D" box="[856,985,1349,1372]" class="Insecta" family="Forficulidae" genus="Anechura" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868331FABEFC76001D" box="[856,985,1349,1372]" italics="true" pageId="13" pageNumber="14">A. sokotrana</emphasis>
</taxonomicName>
(currently
<taxonomicName id="4C0300BCC625FF868018FA9EFE92003D" authorityName="Kočárek" authorityYear="2014" baseAuthorityName="Burr" baseAuthorityYear="1905" box="[113,317,1381,1404]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868018FA9EFE92003D" box="[113,317,1381,1404]" italics="true" pageId="13" pageNumber="14">Guanchia sokotrana</emphasis>
</taxonomicName>
). Based on the comparison of the
<typeStatus id="54B8C59DC625FF8682ECFA9EFD1E003D" box="[645,689,1381,1404]" pageId="13" pageNumber="14">type</typeStatus>
material of
<taxonomicName id="4C0300BCC625FF86834AFA9EFBBF003D" box="[803,1040,1381,1404]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF86834AFA9EFBBF003D" box="[803,1040,1381,1404]" italics="true" pageId="13" pageNumber="14">Guanchia bituberculata</emphasis>
</taxonomicName>
with specimens of
<taxonomicName id="4C0300BCC625FF868095FA7EFE2E00DD" authorityName="Kočárek" authorityYear="2014" baseAuthorityName="Burr" baseAuthorityYear="1905" box="[252,385,1413,1436]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868095FA7EFE2E00DD" box="[252,385,1413,1436]" italics="true" pageId="13" pageNumber="14">G. sokotrana</emphasis>
</taxonomicName>
(see above) and the detailed photographs of the male of
<taxonomicName id="4C0300BCC625FF8683D2FA7EFBEF00DD" authorityName="Kočárek" authorityYear="2014" baseAuthorityName="Burr" baseAuthorityYear="1905" box="[955,1088,1413,1436]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF8683D2FA7EFBEF00DD" box="[955,1088,1413,1436]" italics="true" pageId="13" pageNumber="14">G. sokotrana</emphasis>
</taxonomicName>
published by
<bibRefCitation id="EF9206CEC625FF868092FA5EFE1800FD" author="HAAS F. &amp; POHL H. &amp; WRANIK W." box="[251,439,1445,1468]" pageId="13" pageNumber="14" pagination="409 - 419" refId="ref11078" refString="HAAS F., POHL H. &amp; WRANIK W. 2004: Dermaptera of the Socotra Archipelago with the description of a new species. Fauna of Arabia 20: 409 - 419." type="journal article" year="2004">HAAS et al. (2004)</bibRefCitation>
, these two species were recognised to be distinct. Therefore,
<taxonomicName id="4C0300BCC625FF86844EFA5EFF56009D" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC625FF86844EFA5EFF56009D" italics="true" pageId="13" pageNumber="14">G. bituberculata</emphasis>
</taxonomicName>
is removed from the synonymy and considered to be a valid species. These two species can be distinguished based on the following combination of characters:
<taxonomicName id="4C0300BCC625FF868319FA1EFBEE00BD" authorityName="Kočárek" authorityYear="2014" baseAuthorityName="Burr" baseAuthorityYear="1905" box="[880,1089,1509,1532]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="13" pageNumber="14" phylum="Arthropoda" rank="species" species="sokotrana">
<emphasis id="B977A72DC625FF868319FA1EFBEE00BD" box="[880,1089,1509,1532]" italics="true" pageId="13" pageNumber="14">Guanchia sokotrana</emphasis>
</taxonomicName>
(
<figureCitation id="133867BAC625FF868010F9FEFF5F035D" box="[121,240,1541,1564]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="13" pageNumber="14">Figs 2023</figureCitation>
): the legs, pronotum, abdomen and forceps are pubescent; the middle antennomeres are eight times longer than wide; the pronotum and tegmina are blackish brown, the pronotum has yellowish sides, each tegmen has a yellowish central spot; the pronotum is as long as it is wide; the ultimate tergite has two narrow conical tubercules, the distal end of the tergite between them is straight (
<figureCitation id="133867BAC626FF858129F9FFFE23035A" box="[320,396,1540,1563]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="14" pageNumber="15">Fig. 22</figureCitation>
); the forceps are rounded, regularly tapering from the base to the tip; the male genitalia (
<figureCitation id="133867BAC626FF85815DF9DFFED1037A" box="[308,382,1572,1595]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="14" pageNumber="15">Fig. 23</figureCitation>
) have long parameres, 5.56.0 times longer than wide, regularly tapering to the tip.
<taxonomicName id="4C0300BCC627FF8480BDFF27FE6905B2" box="[212,454,220,243]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC627FF8480BDFF27FE6905B2" box="[212,454,220,243]" italics="true" pageId="15" pageNumber="16">Guanchia bituberculata</emphasis>
</taxonomicName>
(
<figureCitation id="133867BAC627FF8481BAFF27FDE505B2" box="[467,586,220,243]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="15" pageNumber="16">Figs 2426</figureCitation>
): smooth, not a pubescent body; the middle antennomeres are 45 times longer than wide; the pronotum and tegmina are entirely reddish brown; the pronotum is wider than long; the ultimate tergite has two large conical tubercles with a weak concavity between them (
<figureCitation id="133867BAC627FF84811FFEC7FE6E0412" box="[374,449,316,339]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="15" pageNumber="16">Fig. 25</figureCitation>
); the branches of the forceps are dilated and angled basally; the male genitalia are robust (
<figureCitation id="133867BAC627FF848113FEA7FE6B0432" box="[378,452,348,371]" captionStart="Figs 2026" captionStartId="14.[151,188,1317,1336]" captionTargetBox="[174,1106,219,1286]" captionTargetId="figure-11@14.[164,1113,219,1294]" captionTargetPageId="14" captionText="Figs 2026. Guanchia sokotrana (Burr, 1905) comb. nov. (2023) and Guanchia bituberculata (Brindle, 1966) stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype." figureDoi="http://doi.org/10.5281/zenodo.5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="15" pageNumber="16">Fig. 26</figureCitation>
) with short parameres that are 3.03.5 times longer than wide, with a rounded tip.
</paragraph>
</subSubSection>
<caption id="DF7C2BB7C626FF8580FEFADEFE4600C9" ID-DOI="http://doi.org/10.5281/zenodo.5312364" ID-Zenodo-Dep="5312364" httpUri="https://zenodo.org/record/5312364/files/figure.png" pageId="14" pageNumber="15" startId="14.[151,188,1317,1336]" targetBox="[174,1106,219,1286]" targetPageId="14">
<paragraph id="8BBC7B3FC626FF8580FEFADEFE4600C9" blockId="14.[151,1127,1317,1416]" pageId="14" pageNumber="15">
Figs 2026.
<taxonomicName id="4C0300BCC626FF85816FFADEFD850079" authority="(Burr, 1905)" authorityName="Kočárek" authorityYear="2014" baseAuthorityName="Burr" baseAuthorityYear="1905" box="[262,554,1317,1336]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="sokotrana" status="comb. nov.">
<emphasis id="B977A72DC626FF85816FFADEFE190079" box="[262,438,1317,1336]" italics="true" pageId="14" pageNumber="15">Guanchia sokotrana</emphasis>
(
<bibRefCitation id="EF9206CEC626FF8581ADFADEFD8A0079" author="BURR M." box="[452,549,1317,1336]" pageId="14" pageNumber="15" pagination="468 - 496" refId="ref10949" refString="BURR M. 1905: Notes on Forficularia. IX. One new species with synonymic notes. Annals and Magazine of Natural History, Series 7 16: 468 - 496." type="journal article" year="1905">Burr, 1905</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A2441A56C626FF85825BFADEFD3C0079" box="[562,659,1317,1336]" pageId="14" pageNumber="15" rank="species">comb. nov.</taxonomicNameLabel>
(2023) and
<taxonomicName id="4C0300BCC626FF858362FADEFBC90079" authority="(Brindle, 1966)" baseAuthorityName="Brindle" baseAuthorityYear="1966" box="[779,1126,1317,1336]" class="Insecta" family="Forficulidae" genus="Guanchia" kingdom="Animalia" order="Dermaptera" pageId="14" pageNumber="15" phylum="Arthropoda" rank="species" species="bituberculata">
<emphasis id="B977A72DC626FF858362FADEFC760079" box="[779,985,1317,1336]" italics="true" pageId="14" pageNumber="15">Guanchia bituberculata</emphasis>
(
<bibRefCitation id="EF9206CEC626FF85838FFADEFBCF0079" author="BRINDLE A." box="[998,1120,1317,1336]" pageId="14" pageNumber="15" pagination="29 - 49" refId="ref10820" refString="BRINDLE A. 1966: Dermaptera from Central and East Africa. Revue de Zoologie et de Botanique Africaine 74: 29 - 49." type="journal article" year="1966">Brindle, 1966</bibRefCitation>
)
</taxonomicName>
stat. restit. (2426). 20 habitus of male, dorsal view; 21 male sternal plates, ventral view; 22 detail of the end of male ultimate tergite; 23 male genitalia; 24 habitus of male holotype, 25 detail of the end of male ultimate tergite; 26 male genitalia of holotype.
</paragraph>
</caption>
<subSubSection id="C31928B4C627FF848018FE67FC3E04F2" box="[113,913,412,435]" pageId="15" pageNumber="16" type="distribution">
<paragraph id="8BBC7B3FC627FF848018FE67FC3E04F2" blockId="15.[113,1089,220,435]" box="[113,913,412,435]" pageId="15" pageNumber="16">
<emphasis id="B977A72DC627FF848018FE67FEAB04F2" bold="true" box="[113,260,412,435]" pageId="15" pageNumber="16">Distribution.</emphasis>
<collectingCountry id="F3143BAFC627FF848163FE67FD2D04F2" box="[266,642,412,435]" name="Democratic Republic of the Congo" pageId="15" pageNumber="16">Democratic Republic of the Congo</collectingCountry>
,
<collectingCountry id="F3143BAFC627FF8482E4FE67FD4F04F2" box="[653,736,412,435]" name="Uganda" pageId="15" pageNumber="16">Uganda</collectingCountry>
(
<bibRefCitation id="EF9206CEC627FF848287FE67FC2A04F2" author="BRINDLE A." box="[750,901,412,435]" pageId="15" pageNumber="16" pagination="1 - 335" refId="ref10845" refString="BRINDLE A. 1973: The Dermaptera of Africa. Pt. 1. Annales du Musee Royal de l'Afrique Centrale Tervuren, Sciences Zoologiques 205: 1 - 335." type="journal article" year="1973">BRINDLE 1973</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>