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<document id="37051759A60994DAB737A13A1AF61BF3" ID-DOI="10.5281/zenodo.12764872" ID-ISSN="1989-6581" ID-Zenodo-Dep="12764872" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1721243510350" checkinUser="felipe" docAuthor="Fa ̇ ndez, Eduardo I., Rider, David A. &amp; Carvajal, Mariom A." docDate="2017" docId="03AAB33B8E3FFF9427CAFAB0902DFC5E" docLanguage="en" docName="ArqEntomol.17.285-288.pdf" docOrigin="Arquivos Entomolóxicos 17" docStyle="DocumentStyle:DF23C849EA72F80F6DF49FA5382909D5.2:ArqEntomol.2008-.journal_article.1column" docStyleId="DF23C849EA72F80F6DF49FA5382909D5" docStyleName="ArqEntomol.2008-.journal_article.1column" docStyleVersion="2" docTitle="Graphorn bicornutus ndez, Rider &amp; Carvajal, 2017, sp. nov." docType="treatment" docVersion="3" lastPageNumber="287" masterDocId="FF93CB438E3EFF96275AFF9E914FFF9B" masterDocTitle="Graphorn bicornutus, a new genus and a new species from Argentina (Heteroptera: Pentatomidae)." masterLastPageNumber="288" masterPageNumber="285" pageNumber="286" updateTime="1721249399668" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="1FA85FF00C67032B22ACCCFBBCAE8980" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D1909F4D6C47313B2FE86E15A2F62DF3">
<mods:title id="E8C3B688EEBE36F2063D71D077F07366">Graphorn bicornutus, a new genus and a new species from Argentina (Heteroptera: Pentatomidae).</mods:title>
</mods:titleInfo>
<mods:name id="5E25A768F0A03D5EEB4273E89FBA30A8" type="personal">
<mods:role id="C3E789AA41273AECF2E109805D4935A1">
<mods:roleTerm id="710FD25F867D785B8AB4C1A193F6FD6A">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6E0456A6B1EE4B39AE9E5E27B8ABB58F">Fa ̇ ndez, Eduardo I.</mods:namePart>
</mods:name>
<mods:name id="8E2AA03431AB81EC92F589344DAA8D14" type="personal">
<mods:role id="C514BF659A4DB49916B0F481D44E293F">
<mods:roleTerm id="6FC7C3D6D6950D725E696127784D313B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FD1269B5880A222B8F2587BFD8581D08">Rider, David A.</mods:namePart>
</mods:name>
<mods:name id="19FF5751277C27B4BE3339EC3827D764" type="personal">
<mods:role id="49DC7B9CA08F2067D19D5FAD183D4D92">
<mods:roleTerm id="9B61BB9097BCA4503AE8A413A800BB29">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="FEF221CB46928A452C31BE2CC6EF5B5A">Carvajal, Mariom A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="9C0530B6DE0907DE25E1444D520B2C51">text</mods:typeOfResource>
<mods:relatedItem id="6620CD1DAB37D59D8178E1014AC99CA1" type="host">
<mods:titleInfo id="1C0A7409CEBC83C7EC389249FD164DDD">
<mods:title id="CE0B143033B973BB9AB154FAAA4592D4">Arquivos Entomolóxicos</mods:title>
</mods:titleInfo>
<mods:part id="026FF071971BD8D45C663F5C66FEF30A">
<mods:date id="91996609DC99BB209D1A9761CA6FC86A">2017</mods:date>
<mods:detail id="5D564E8EA2AB356DCF83E207C569FF47" type="pubDate">
<mods:number id="194BFAD803510F83B35E42CB78231C2F">2017-04-10</mods:number>
</mods:detail>
<mods:detail id="74A87160B382648D672FC34525C4EE1D" type="volume">
<mods:number id="1773AFC63F853CFB3C2E44347507BF0D">17</mods:number>
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<mods:start id="804BBADC34C03B610EFFD5730153693D">285</mods:start>
<mods:end id="91B640EF1D42ABD64130C6ADBB920962">288</mods:end>
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<mods:classification id="FEF863FE85B4CEEAA7EA39FE9B538BF8">journal article</mods:classification>
<mods:identifier id="C5706D83C458DD4593730CDA8C4EBFF3" type="DOI">10.5281/zenodo.12764872</mods:identifier>
<mods:identifier id="E86B2FF651F3BAED4BAF7B0AEDB68491" type="ISSN">1989-6581</mods:identifier>
<mods:identifier id="87989FD2B2CF769CC668B114806064E3" type="Zenodo-Dep">12764872</mods:identifier>
</mods:mods>
<treatment id="03AAB33B8E3FFF9427CAFAB0902DFC5E" ID-DOI="http://doi.org/10.5281/zenodo.12764402" ID-Zenodo-Dep="12764402" LSID="urn:lsid:plazi:treatment:03AAB33B8E3FFF9427CAFAB0902DFC5E" httpUri="http://treatment.plazi.org/id/03AAB33B8E3FFF9427CAFAB0902DFC5E" lastPageId="2" lastPageNumber="287" pageId="1" pageNumber="286">
<subSubSection id="C31951A68E3FFF9727CAFAB093C5FADC" box="[144,650,1326,1352]" pageId="1" pageNumber="286" type="nomenclature">
<paragraph id="8BBC022D8E3FFF9727CAFAB093C5FADC" blockId="1.[144,650,1326,1352]" box="[144,650,1326,1352]" pageId="1" pageNumber="286">
<heading id="D0F4B5418E3FFF9727CAFAB093C5FADC" bold="true" box="[144,650,1326,1352]" fontSize="10" level="2" pageId="1" pageNumber="286" reason="6">
<emphasis id="B977DE3F8E3FFF9727CAFAB093C5FADC" bold="true" box="[144,650,1326,1352]" pageId="1" pageNumber="286">
<taxonomicName id="4C0379AE8E3FFF9727CAFAB090DCFADC" authorityName="Fȧndez &amp; Rider &amp; Carvajal" authorityYear="2017" box="[144,403,1326,1352]" class="Insecta" family="Pentatomidae" genus="Graphorn" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="286" phylum="Arthropoda" rank="species" species="bicornutus" status="sp. nov.">
<emphasis id="B977DE3F8E3FFF9727CAFAB090DCFADC" bold="true" box="[144,403,1326,1352]" italics="true" pageId="1" pageNumber="286">Graphorn bicornutus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24463448E3FFF9726C4FAB0934AFADC" box="[414,517,1326,1351]" pageId="1" pageNumber="286" rank="species">sp. nov.</taxonomicNameLabel>
(
<figureCitation id="13381EA88E3FFF97254FFAB09330FADC" box="[533,639,1326,1351]" captionStart="Figs" captionStartId="3.[251,298,2006,2026]" captionTargetBox="[246,1305,864,1978]" captionTargetId="figure-223@3.[239,1311,857,1978]" captionTargetPageId="3" captionText="Figs. 1-5.- Graphorn bicornutus gen. &amp; sp. nov. 1.- Habitus. 2.- Metapleuron. 3.- Female terminalia. 4.- Head and pronotum, detailed dorsal view. 5.- Head and pronotum, detailed lateral view." figureDoi="http://doi.org/10.5281/zenodo.12764874" httpUri="https://zenodo.org/record/12764874/files/figure.png" pageId="1" pageNumber="286">Figs. 1-5</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31951A68E3FFF9427ECFAF093FAFCF0" lastPageId="2" lastPageNumber="287" pageId="1" pageNumber="286" type="description">
<paragraph id="8BBC022D8E3FFF9727ECFAF0901EFA1C" blockId="1.[182,1444,1390,2046]" box="[182,337,1390,1415]" pageId="1" pageNumber="286">
<emphasis id="B977DE3F8E3FFF9727ECFAF0901EFA1C" bold="true" box="[182,337,1390,1415]" pageId="1" pageNumber="286">Description.</emphasis>
</paragraph>
<paragraph id="8BBC022D8E3FFF9727ECFA0D9068F92B" blockId="1.[182,1444,1390,2046]" pageId="1" pageNumber="286">
<emphasis id="B977DE3F8E3FFF9727ECFA0D9068F92B" bold="true" pageId="1" pageNumber="286">
Head: Elongate (
<figureCitation id="13381EA88E3FFF9726D7FA0D9094FA37" box="[397,475,1427,1452]" captionStart="Figs" captionStartId="3.[251,298,2006,2026]" captionTargetBox="[246,1305,864,1978]" captionTargetId="figure-223@3.[239,1311,857,1978]" captionTargetPageId="3" captionText="Figs. 1-5.- Graphorn bicornutus gen. &amp; sp. nov. 1.- Habitus. 2.- Metapleuron. 3.- Female terminalia. 4.- Head and pronotum, detailed dorsal view. 5.- Head and pronotum, detailed lateral view." figureDoi="http://doi.org/10.5281/zenodo.12764874" httpUri="https://zenodo.org/record/12764874/files/figure.png" pageId="1" pageNumber="286">Fig. 4</figureCitation>
), longer than wide, dull cream colored, with dense, dark punctation. Eyes prominent, reddish. Ocelli small, oval, located near base of head, behind an imaginary line drawn through the posterior margins of compound eyes, but not contiguous with pronotum. Paraclypei enlarged, punctate, apices of paraclypei narrowly rounded, but not acute. Lateral margins of head concave for basal two-thirds, then curving mesad for distal third. Antennae, dull cream colored basally, becoming light orange apically, first antennomere not extending beyond apex of head. Rostrum reaching metathoracic coxae, first rostral segment extending beyond hind margins of bucculae.
</emphasis>
</paragraph>
<paragraph id="8BBC022D8E3FFF9427ECF9229361FE13" blockId="1.[182,1444,1390,2046]" lastBlockId="2.[182,1444,219,875]" lastPageId="2" lastPageNumber="287" pageId="1" pageNumber="286">
<emphasis id="B977DE3F8E3FFF9427ECF9229361FE13" bold="true" lastPageId="2" lastPageNumber="287" pageId="1" pageNumber="286">
Thorax: Pronotum hexagonal, dull cream colored, densely punctate, disk abruptly elevated; anterolateral margins crenulate; cicatrices immaculate, each with a small ivory callus at mesial limit; humeral angles projected, acute, each forming a strong, apically dark spine (
<figureCitation id="13381EA88E3FFF97233EF8989596F884" box="[1124,1241,1798,1823]" captionStart="Figs" captionStartId="3.[251,298,2006,2026]" captionTargetBox="[246,1305,864,1978]" captionTargetId="figure-223@3.[239,1311,857,1978]" captionTargetPageId="3" captionText="Figs. 1-5.- Graphorn bicornutus gen. &amp; sp. nov. 1.- Habitus. 2.- Metapleuron. 3.- Female terminalia. 4.- Head and pronotum, detailed dorsal view. 5.- Head and pronotum, detailed lateral view." figureDoi="http://doi.org/10.5281/zenodo.12764874" httpUri="https://zenodo.org/record/12764874/files/figure.png" pageId="1" pageNumber="286">Figs. 4, 5</figureCitation>
); posterolateral margins somewhat irregular. Scutellum subtriangular (
<figureCitation id="13381EA88E3FFF9724D6F8B29295F8DE" box="[908,986,1836,1861]" captionStart="Figs" captionStartId="3.[251,298,2006,2026]" captionTargetBox="[246,1305,864,1978]" captionTargetId="figure-223@3.[239,1311,857,1978]" captionTargetPageId="3" captionText="Figs. 1-5.- Graphorn bicornutus gen. &amp; sp. nov. 1.- Habitus. 2.- Metapleuron. 3.- Female terminalia. 4.- Head and pronotum, detailed dorsal view. 5.- Head and pronotum, detailed lateral view." figureDoi="http://doi.org/10.5281/zenodo.12764874" httpUri="https://zenodo.org/record/12764874/files/figure.png" pageId="1" pageNumber="286">Fig. 1</figureCitation>
), dull cream colored with nearly concolorous, dense punctures, with two relatively large ivory calluses, one on each side near basal angles, extending little beyond base of scutellum; lateral margins slightly concave mesially; apex broadly rounded. Coria dull cream colored, densely, dark punctate, somewhat darker near clavus, radial vein simple, not carinate or calloused; membrane smokey with conspicuous longitudinal veins. Thoracic sterna yellowish with dense light-brown punctures, a weak oval ivory callus present in middle of mesosternum. Metapleura dull cream colored with dark dense punctation, each ostiolar peritreme short, not reaching halfway to lateral margin of metapleuron, evaporatoria relatively small (
<figureCitation id="13381EA88E3CFF9427E4FEBB9046FEA5" box="[190,265,293,318]" captionStart="Figs" captionStartId="3.[251,298,2006,2026]" captionTargetBox="[246,1305,864,1978]" captionTargetId="figure-223@3.[239,1311,857,1978]" captionTargetPageId="3" captionText="Figs. 1-5.- Graphorn bicornutus gen. &amp; sp. nov. 1.- Habitus. 2.- Metapleuron. 3.- Female terminalia. 4.- Head and pronotum, detailed dorsal view. 5.- Head and pronotum, detailed lateral view." figureDoi="http://doi.org/10.5281/zenodo.12764874" httpUri="https://zenodo.org/record/12764874/files/figure.png" pageId="2" pageNumber="287">Fig. 2</figureCitation>
), each restricted to a very small rugose area around ostiolar peritreme and along posterior margin of mesopleuron, ostiolar plate weakly punctured. Legs yellow, with strong dark setae, tibiae and femora subequal in length.
</emphasis>
</paragraph>
<paragraph id="8BBC022D8E3CFF9427ECFE0B951EFD86" blockId="2.[182,1444,219,875]" pageId="2" pageNumber="287">
<emphasis id="B977DE3F8E3CFF9427ECFE0B951EFD86" bold="true" pageId="2" pageNumber="287">Abdomen: Yellowish dorsally; connexiva dull, cream colored, posterolateral angles prominent, acute; each connexival segment with brownish markings mesially. Abdominal venter dull, cream colored, with dense, concolorous punctatures; transverse rows of small ivory calluses along abdominal sterna; spiracles yellowish to caramel, each placed on distinct circular ivory callus.</emphasis>
</paragraph>
<paragraph id="8BBC022D8E3CFF9427ECFDB79021FD2A" blockId="2.[182,1444,219,875]" pageId="2" pageNumber="287">
<emphasis id="B977DE3F8E3CFF9427ECFDB79021FD2A" bold="true" pageId="2" pageNumber="287">
Female genitalia (
<figureCitation id="13381EA88E3CFF9426CFFDB790ADFDD9" box="[405,482,553,578]" captionStart="Figs" captionStartId="3.[251,298,2006,2026]" captionTargetBox="[246,1305,864,1978]" captionTargetId="figure-223@3.[239,1311,857,1978]" captionTargetPageId="3" captionText="Figs. 1-5.- Graphorn bicornutus gen. &amp; sp. nov. 1.- Habitus. 2.- Metapleuron. 3.- Female terminalia. 4.- Head and pronotum, detailed dorsal view. 5.- Head and pronotum, detailed lateral view." figureDoi="http://doi.org/10.5281/zenodo.12764874" httpUri="https://zenodo.org/record/12764874/files/figure.png" pageId="2" pageNumber="287">Fig. 3</figureCitation>
): First gonocoxae finely punctate. Each gonocoxite triangular-shaped, with convex sides and slightly elongated apices. Second gonocoxae small, rectangular shaped with rounded apices. Paratergites 8 rhomboid with pointed apices. Paratergites 9 triangular, swollen distally, with pointed apices.
</emphasis>
</paragraph>
<paragraph id="8BBC022D8E3CFF9427ECFD209029FD4C" blockId="2.[182,1444,219,875]" box="[182,358,702,727]" pageId="2" pageNumber="287">
<emphasis id="B977DE3F8E3CFF9427ECFD209029FD4C" bold="true" box="[182,358,702,727]" pageId="2" pageNumber="287">Male unknown.</emphasis>
</paragraph>
<paragraph id="8BBC022D8E3CFF9427ECFD7D93FAFCF0" blockId="2.[182,1444,219,875]" pageId="2" pageNumber="287">
<emphasis id="B977DE3F8E3CFF9427ECFD7D93FAFCF0" bold="true" pageId="2" pageNumber="287">Measurements: Total length: 9.45, transhumeral pronotal width: 6.95, medial pronotal length: 2.25, medial head length: 2.11, head width across compound eyes: 1.49, medial scutellar length: 3.43, basal scutellar width: 3.43, rostral segments: I: 1.37 II: 1.34 III: 0.62 IV: 0.56, antennal segments: I: 0.52 II: 0.90 III: 0.99, IV: 0.86, V: 1.22.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C31951A68E3CFF9427ECFC19902DFC5E" pageId="2" pageNumber="287" type="materials_examined">
<paragraph id="8BBC022D8E3CFF9427ECFC19902DFC5E" blockId="2.[182,1444,902,965]" pageId="2" pageNumber="287">
<materialsCitation id="3B6B08708E3CFF9427ECFC19902DFC5E" collectingDate="1989-01-25" collectorName="L. O'Brien &amp; J. Gibmer" country="Argentina" location="Chaco Province" pageId="2" pageNumber="287" specimenCount="1" specimenCount-female="1" stateProvince="Chaco" typeStatus="holotype">
<emphasis id="B977DE3F8E3CFF9427ECFC1995D2FC3B" bold="true" box="[182,1181,902,928]" pageId="2" pageNumber="287">
<typeStatus id="54B8BC8F8E3CFF9427ECFC199062FC3B" box="[182,301,903,928]" pageId="2" pageNumber="287" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F31442BD8E3CFF942660FC1890ACFC3B" box="[314,483,902,928]" name="Argentina" pageId="2" pageNumber="287">ARGENTINA</collectingCountry>
,
<collectingRegion id="49C7CCCF8E3CFF9426AAFC1893E3FC3B" box="[496,684,902,928]" country="Argentina" name="Chaco" pageId="2" pageNumber="287">Chaco Province</collectingRegion>
,
<quantity id="4CFBAFC88E3CFF9425E3FC199249FC3B" box="[697,774,903,928]" metricMagnitude="4" metricUnit="m" metricValue="6.0" pageId="2" pageNumber="287" unit="km" value="60.0">60 km</quantity>
NE of
<collectingRegion id="49C7CCCF8E3CFF94243BFC1992BEFC3B" box="[865,1009,903,928]" country="Argentina" name="Resistencia" pageId="2" pageNumber="287">Resistencia</collectingRegion>
,
<date id="FFBD24ED8E3CFF9424A4FC1995CBFC3B" box="[1022,1156,903,928]" pageId="2" pageNumber="287" value="1989-01-25">
<collectingDate id="EFF9DD058E3CFF9424A4FC1995CBFC3B" box="[1022,1156,903,928]" pageId="2" pageNumber="287" value="1989-01-25">25-I-1989</collectingDate>
</date>
,
<specimenCount id="9D05C9A48E3CFF9423CBFC1995D2FC3B" box="[1169,1181,903,928]" count="1" pageId="2" pageNumber="287" type="female">1</specimenCount>
</emphasis>
♀,
<emphasis id="B977DE3F8E3CFF94239BFC189449FC3B" bold="true" box="[1217,1286,902,928]" pageId="2" pageNumber="287">CW &amp;</emphasis>
<collectorName id="26F667FB8E3CFF94224AFC1994C5FC3B" box="[1296,1418,902,928]" pageId="2" pageNumber="287">L. O’Brien</collectorName>
&amp;
<emphasis id="B977DE3F8E3CFF9427ECFC32902DFC5E" bold="true" box="[182,354,939,965]" pageId="2" pageNumber="287">
<collectorName id="26F667FB8E3CFF9427ECFC32907FFC5E" box="[182,304,939,965]" pageId="2" pageNumber="287">J. Gibmer</collectorName>
leg.
</emphasis>
</materialsCitation>
</paragraph>
</subSubSection>
</treatment>
</document>
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