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<document id="10B5CE0192CC085BC8EA66C9E2E94780" ID-DOI="10.11646/zootaxa.5514.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="13849492" ID-ZooBank="BF26E06E-7719-4542-ACF5-F8DBE4015F58" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1727441078522" checkinUser="plazi" docAuthor="He, Liang &amp; Zhou, Hong-Zhang" docDate="2024" docId="03AA87A8FFC5CE66FF51FB86498AFD80" docLanguage="en" docName="zootaxa.5514.1.6.pdf" docOrigin="Zootaxa 5514 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5514.1.6" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Miobdelus motuoensis He and Zhou 2024, sp. nov." docType="treatment" docVersion="2" lastPageNumber="95" masterDocId="FF93FFD0FFC4CE62FFC6FFB54963FFE1" masterDocTitle="A new species of Miobdelus Sharp and two new records of Eucibdelus Kraatz (Coleoptera, Staphylinidae, Staphylininae) from China" masterLastPageNumber="100" masterPageNumber="91" pageNumber="92" updateTime="1727442566679" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="696D74A7424EC4624DD4AC352F0083F2">A new species of Miobdelus Sharp and two new records of Eucibdelus Kraatz (Coleoptera, Staphylinidae, Staphylininae) from China</mods:title>
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<mods:name id="EC9918837473923EDCFC143EBE91A099" type="personal">
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<mods:roleTerm id="3964AE21AAA355A521BAC010BB8A3FB6">Author</mods:roleTerm>
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<mods:namePart id="37B3315BB8BD26EFF145EA589C0AF729">He, Liang</mods:namePart>
<mods:affiliation id="2F7CD6674F92AD8FA5D7466552FD30FD">Institute of Plant Protection, Tianjin Academy of Agricultural Sciences, 17 km in Jinjing Road, Xiqing District, Tianjin, 300384, China</mods:affiliation>
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<mods:namePart id="E165ACBEE2E80575819A149955774E31">Zhou, Hong-Zhang</mods:namePart>
<mods:affiliation id="DBF4A55CC3ECD8EBC1F951A041900BFB">Key Laboratory of Zoological Systematics and Evolution, Institute of Zoology, Chinese Academy of Sciences, 1 Beichen West Rd., Chaoyang District, Beijing 100101, P. R. China &amp; University of the Chinese Academy of Sciences, 19 A Yuquan Rd., Shijingshan District, Beijing 100049, P. R. China</mods:affiliation>
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<mods:title id="957872956D748AD09248E4677423691A">Zootaxa</mods:title>
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<mods:date id="BF709ED5DDB8431433D7404403C75800">2024</mods:date>
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<mods:number id="35DB94760593882A4634C3F97608A524">2024-09-25</mods:number>
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<mods:number id="0B8E0E192A1B5E356A4DAE15D88719C0">5514</mods:number>
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<mods:identifier id="15F9697A64E391989718E27998C1BB36" type="DOI">10.11646/zootaxa.5514.1.6</mods:identifier>
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<treatment id="03AA87A8FFC5CE66FF51FB86498AFD80" LSID="urn:lsid:plazi:treatment:03AA87A8FFC5CE66FF51FB86498AFD80" httpUri="http://treatment.plazi.org/id/03AA87A8FFC5CE66FF51FB86498AFD80" lastPageId="4" lastPageNumber="95" pageId="1" pageNumber="92">
<subSubSection id="C3196535FFC5CE63FF51FB864BDCFBAF" box="[151,703,1075,1102]" pageId="1" pageNumber="92" type="nomenclature">
<paragraph id="8BBC36BEFFC5CE63FF51FB864BDCFBAF" blockId="1.[151,703,1075,1138]" box="[151,703,1075,1102]" pageId="1" pageNumber="92">
<heading id="D0F481D2FFC5CE63FF51FB864BDCFBAF" bold="true" box="[151,703,1075,1102]" fontSize="11" level="1" pageId="1" pageNumber="92" reason="1">
<emphasis id="B977EAACFFC5CE63FF51FB864BDCFBAF" bold="true" box="[151,703,1075,1102]" pageId="1" pageNumber="92">
<taxonomicName id="4C034D3DFFC5CE63FF51FB864B36FBAF" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[151,597,1075,1102]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC5CE63FF51FB8648C4FBAF" bold="true" box="[151,423,1075,1102]" italics="true" pageId="1" pageNumber="92">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
,
<taxonomicNameLabel id="A24457D7FFC5CE63FDA7FB814BDCFBAF" box="[609,703,1076,1102]" pageId="1" pageNumber="92" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3196535FFC5CE63FF51FBED4829FB93" box="[151,330,1112,1138]" pageId="1" pageNumber="92" type="description">
<paragraph id="8BBC36BEFFC5CE63FF51FBED4829FB93" blockId="1.[151,703,1075,1138]" box="[151,330,1112,1138]" pageId="1" pageNumber="92">
(
<figureCitation id="13382A3BFFC5CE63FF59FBED4822FB93" box="[159,321,1112,1138]" captionStart-0="FIGURE 1–1" captionStart-1="FIGURE 1–2" captionStartId-0="2.[152,255,1808,1832]" captionStartId-1="3.[152,255,1905,1929]" captionTargetBox-0="[152,1431,388,1780]" captionTargetBox-1="[151,1436,273,1882]" captionTargetId-0="figure-104@2.[151,1436,378,1782]" captionTargetId-1="figure-76@3.[151,1436,273,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–1. Miobdelus motuoensis He and Zhou sp. nov., morphology A habitus, dorsal view B aedeagus, lateral view C aedeagus, ventral view D male sternite 9 E male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E)." captionText-1="FIGURE 1–2. Miobdelus motuoensis He and Zhou sp. nov., morphology A male sternite 9 B &amp; D aedeagus, lateral view C aedeagus, ventral view E male tergite 10 F male sternite 8 G underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.13849496" figureDoi-1="http://doi.org/10.5281/zenodo.13849498" httpUri-0="https://zenodo.org/record/13849496/files/figure.png" httpUri-1="https://zenodo.org/record/13849498/files/figure.png" pageId="1" pageNumber="92">Figs. 1–1, 1–2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3196535FFC5CE63FF51FB15486FFAC7" pageId="1" pageNumber="92" type="materials_examined">
<paragraph id="8BBC36BEFFC5CE63FF51FB154BD8FB5B" blockId="1.[151,1437,1184,2039]" box="[151,699,1184,1211]" pageId="1" pageNumber="92">
<materialsCitation id="3B6B3CE3FFC5CE63FF51FB154BD5FB5B" box="[151,694,1184,1211]" country="China" county="Motuo County" location="Motuo County" pageId="1" pageNumber="92" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="B977EAACFFC5CE63FF51FB154B61FB5B" bold="true" box="[151,514,1184,1210]" pageId="1" pageNumber="92">
<typeStatus id="54B8881CFFC5CE63FF51FB1549B1FB5B" box="[151,210,1184,1210]" pageId="1" pageNumber="92">Type</typeStatus>
locality:
<collectingCountry id="F314762EFFC5CE63FE86FB1548C1FB5B" box="[320,418,1184,1210]" name="China" pageId="1" pageNumber="92">CHINA</collectingCountry>
:
<collectingRegion id="49C7F85CFFC5CE63FE68FB154B61FB5B" box="[430,514,1184,1210]" country="China" name="Xizang" pageId="1" pageNumber="92">Xizang</collectingRegion>
</emphasis>
:
<collectingCounty id="62DD4E32FFC5CE63FDD7FB144BD5FB5B" box="[529,694,1184,1211]" pageId="1" pageNumber="92">Motuo County</collectingCounty>
</materialsCitation>
.
</paragraph>
<paragraph id="8BBC36BEFFC5CE63FF01FB71486FFAC7" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="92">
<emphasis id="B977EAACFFC5CE63FF01FB71480CFB3F" bold="true" box="[199,367,1220,1246]" pageId="1" pageNumber="92">Type material</emphasis>
.
<materialsCitation id="3B6B3CE3FFC5CE63FEBBFB71486FFAC7" collectingDate="2021-06-11" collectionCode="IOZ, IZ-CAS" collectorName="Hong Liu &amp; Man Qin" country="China" county="Motuo County" elevation="1408" latitude="29.2337" location="Gelin Village" longLatPrecision="6" longitude="95.1771" municipality="Beibeng Town" pageId="1" pageNumber="92" specimenCount="1" stateProvince="Xizang" typeStatus="holotype">
<emphasis id="B977EAACFFC5CE63FEBBFB714889FB3F" bold="true" box="[381,490,1220,1246]" pageId="1" pageNumber="92">
<typeStatus id="54B8881CFFC5CE63FEBBFB714889FB3F" box="[381,490,1220,1246]" pageId="1" pageNumber="92" type="holotype">Holotype</typeStatus>
</emphasis>
male,
<emphasis id="B977EAACFFC5CE63FDF1FB714B99FB3F" bold="true" box="[567,762,1220,1246]" pageId="1" pageNumber="92">
<collectingCountry id="F314762EFFC5CE63FDF1FB714BFAFB3F" box="[567,665,1220,1246]" name="China" pageId="1" pageNumber="92">CHINA</collectingCountry>
:
<collectingRegion id="49C7F85CFFC5CE63FD60FB714B99FB3F" box="[678,762,1220,1246]" country="China" name="Xizang" pageId="1" pageNumber="92">Xizang</collectingRegion>
</emphasis>
:
<collectingCounty id="62DD4E32FFC5CE63FCCFFB704AD3FB3F" box="[777,944,1220,1247]" pageId="1" pageNumber="92">Motuo County</collectingCounty>
,
<collectingMunicipality id="6BD8ACC4FFC5CE63FC7DFB714D01FB3E" box="[955,1122,1220,1247]" pageId="1" pageNumber="92">Beibeng Town</collectingMunicipality>
,
<quantity id="4CFB9B5BFFC5CE63FBA8FB714DD5FB3F" box="[1134,1206,1220,1246]" metricMagnitude="4" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="92" unit="km" value="17.0">17 km</quantity>
in the road of
<location id="8EDC6065FFC5CE63FA98FB714988FAE3" LSID="urn:lsid:plazi:treatment:03AA87A8FFC5CE66FF51FB86498AFD80:8EDC6065FFC5CE63FA98FB714988FAE3" country="China" county="Motuo County" latitude="29.2337" longLatPrecision="6" longitude="95.1771" municipality="Beibeng Town" name="Gelin Village" pageId="1" pageNumber="92" stateProvince="Xizang">Gelin Village</location>
,
<geoCoordinate id="EE375079FFC5CE63FF31FB5D4817FAE3" box="[247,372,1256,1282]" degrees="29.2337" direction="north" orientation="latitude" pageId="1" pageNumber="92" precision="5" value="29.2337">29.2337°N</geoCoordinate>
,
<geoCoordinate id="EE375079FFC5CE63FE46FB5D4899FAE2" box="[384,506,1256,1283]" degrees="95.1771" direction="east" orientation="longitude" pageId="1" pageNumber="92" precision="5" value="95.1771">95.1771°E</geoCoordinate>
,
<quantity id="4CFB9B5BFFC5CE63FDC0FB5D4B1AFAE3" box="[518,633,1256,1282]" metricMagnitude="3" metricUnit="m" metricValue="1.4080099999999998" pageId="1" pageNumber="92" unit="m" value="1408.01">
<elevation id="002ED18DFFC5CE63FDC0FB5D4B1AFAE3" box="[518,633,1256,1282]" metricMagnitude="3" metricUnit="m" metricValue="1.4080099999999998" pageId="1" pageNumber="92" unit="m" value="1408.01">1408.01m</elevation>
</quantity>
,
<collectingMethod id="52424EA9FFC5CE63FD43FB5D4B8DFAE2" box="[645,750,1256,1283]" pageId="1" pageNumber="92">light trap</collectingMethod>
,
<date id="FFBD107EFFC5CE63FD3CFB5D4A1FFAE3" box="[762,892,1256,1282]" pageId="1" pageNumber="92" value="2021-06-11">
<collectingDate id="EFF9E996FFC5CE63FD3CFB5D4A1FFAE3" box="[762,892,1256,1282]" pageId="1" pageNumber="92" value="2021-06-11">2021.VI.11</collectingDate>
</date>
N1,
<collectionCode id="ED12AE7BFFC5CE63FC6CFB5D4AB8FAE2" box="[938,987,1256,1283]" pageId="1" pageNumber="92">IOZ</collectionCode>
(E)2562097, coll.
<collectorName id="26F65368FFC5CE63FB63FB5C4C71FAE3" box="[1189,1298,1256,1283]" pageId="1" pageNumber="92">Hong Liu</collectorName>
&amp;
<collectorName id="26F65368FFC5CE63FAF1FB5C4CFFFAE3" box="[1335,1436,1256,1283]" pageId="1" pageNumber="92">Man Qin</collectorName>
(
<collectionCode id="ED12AE7BFFC5CE63FF59FAB94863FAC7" box="[159,256,1292,1318]" pageId="1" pageNumber="92">IZ-CAS</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3196535FFC5CE66FF01FA8548C1FF50" lastPageId="4" lastPageNumber="95" pageId="1" pageNumber="92" type="description">
<paragraph id="8BBC36BEFFC5CE63FF01FA8548D5FA8E" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="92">
<emphasis id="B977EAACFFC5CE63FF01FA85481DFAAB" bold="true" box="[199,382,1328,1354]" pageId="1" pageNumber="92">Measurements.</emphasis>
Body length:
<quantity id="4CFB9B5BFFC5CE63FDE7FA854BEAFAAA" box="[545,649,1328,1355]" metricMagnitude="-2" metricUnit="m" metricValue="2.06" pageId="1" pageNumber="92" unit="mm" value="20.6">20.6 mm</quantity>
. CL: 1.10; EL: 3.67; ELS: 2.01; EW: 3.83; HL: 2.93; HW: 3.31; PO: 0.99; PL: 3.58; PW: 3.15.
</paragraph>
<paragraph id="8BBC36BEFFC5CE63FF01FACD4BF3F96F" blockId="1.[151,1437,1184,2039]" pageId="1" pageNumber="92">
<emphasis id="B977EAACFFC5CE63FF01FACD483AFA73" bold="true" box="[199,345,1400,1426]" pageId="1" pageNumber="92">Description.</emphasis>
Relatively large sized species among members of
<taxonomicName id="4C034D3DFFC5CE63FC4EFACD4A9DFA73" authorityName="Sharp" authorityYear="1889" box="[904,1022,1400,1426]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAACFFC5CE63FC4EFACD4A9DFA73" box="[904,1022,1400,1426]" italics="true" pageId="1" pageNumber="92">Miobdelus</emphasis>
</taxonomicName>
. Body elongate, almost totally black; head, pronotum and elytra totally black; visible abdominal tergites black; apical half of visible abdomen tergites 2 and 3 each with one inconspicuous black tomentose patch; middle portion of visible abdominal tergite 4 covered with a large black tomentose patch, with two golden-yellowish tomentose patches on lateral sides; almost entire surface of visible abdominal tergite 5 covered with pale black pubescence; maxillary and labial palpi black to dark brunneous; antennae with segments 1 to 3 dark brown; segments 4–7 reddish-brown, with basal and apical portion distinctly paler, gradually becoming darker towards apex; segments 8 to 11 yellowish-white; legs black, with tarsus distinctly paler, tending to be reddish-brown.
</paragraph>
<paragraph id="8BBC36BEFFC5CE60FF01F92D488CFE84" blockId="1.[151,1437,1184,2039]" lastBlockId="2.[151,1437,151,357]" lastPageId="2" lastPageNumber="93" pageId="1" pageNumber="92">
Head of rounded quadrangular shape, with obtusely rounded posterior angles, slightly wider than long (ratio 1.12); eyes relatively larger and more protruding than most
<taxonomicName id="4C034D3DFFC5CE63FCFEF9094ACDF937" authorityName="Sharp" authorityYear="1889" box="[824,942,1724,1750]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="92" phylum="Arthropoda" rank="genus">
<emphasis id="B977EAACFFC5CE63FCFEF9094ACDF937" box="[824,942,1724,1750]" italics="true" pageId="1" pageNumber="92">Miobdelus</emphasis>
</taxonomicName>
, slightly longer than tempora (ratio 1.11) in dorsal view; punctation and pubescence on dorsal surface of head dense, punctation robust, gradually becoming sparser toward clypeus, with shiny interspaces; interspaces without microsculpture. Y-shaped epicranial line almost totally disappeared on head disc. Antennae long and slender, segment 3 longer than segment 2 (ratio 1.35), segments 4 to 7 distinctly longer than wide, segments 8 to 10 about as long as wide to vaguely wider than long, last segment distinctly wider than segment 10. Pronotum longer than wide (ratio 1.13), lateral sides almost parallel, slightly curved, gradually narrowed towards widely rounded posterior angles; narrow marginal groove disappearing downwards at posterior fourth of pronotal length; impunctate midline almost totally disappeared, only vaguely present on posterior fifth of pronotal disc; punctation and pubescence on disc finer and denser than that on dorsal surface of head, interspace without microsculpture. Scutellum finely punctate and setose on entire surface, appearing very dull. Elytra moderately long, elytral length distinctly shorter (ratio 0.56) along suture but vaguely longer (ratio 1.02) along sides than pronotum along midline, slightly dilated posteriad; elytral punctation fine and dense, granulose. Wings fully developed. Abdomen elongate, with almost parallel lateral sides, gradually narrowed posteriorly; all visible abdominal tergites finely and densely punctate and pubescent; posterior margin of visible tergite 5 without narrow seam of palisade setae.
</paragraph>
<caption id="DF7C6636FFC6CE60FF5EF8A54DC2F8AC" ID-DOI="http://doi.org/10.5281/zenodo.13849496" ID-Zenodo-Dep="13849496" httpUri="https://zenodo.org/record/13849496/files/figure.png" pageId="2" pageNumber="93" startId="2.[152,255,1808,1832]" targetBox="[152,1431,388,1780]" targetPageId="2" targetType="figure">
<paragraph id="8BBC36BEFFC6CE60FF5EF8A54DC2F8AC" blockId="2.[151,1436,1808,1869]" pageId="2" pageNumber="93">
<emphasis id="B977EAACFFC6CE60FF5EF8A54850F8C9" bold="true" box="[152,307,1808,1832]" pageId="2" pageNumber="93">FIGURE 1–1.</emphasis>
<taxonomicName id="4C034D3DFFC6CE60FEFCF8A44BCEF8C8" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[314,685,1809,1833]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="93" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC6CE60FEFCF8A44B7CF8C9" box="[314,543,1809,1833]" italics="true" pageId="2" pageNumber="93">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
<emphasis id="B977EAACFFC6CE60FD73F8A44A65F8C8" bold="true" box="[693,774,1809,1833]" pageId="2" pageNumber="93">
<taxonomicNameLabel id="A24457D7FFC6CE60FD73F8A44A65F8C8" box="[693,774,1809,1833]" pageId="2" pageNumber="93" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, morphology
<emphasis id="B977EAACFFC6CE60FC5EF8A54AC9F8C9" bold="true" box="[920,938,1808,1832]" pageId="2" pageNumber="93">A</emphasis>
habitus, dorsal view
<emphasis id="B977EAACFFC6CE60FB41F8A44DFBF8C8" bold="true" box="[1159,1176,1809,1833]" pageId="2" pageNumber="93">B</emphasis>
aedeagus, lateral view
<emphasis id="B977EAACFFC6CE60FA4CF8A54CFFF8C9" bold="true" box="[1418,1436,1808,1832]" pageId="2" pageNumber="93">C</emphasis>
aedeagus, ventral view
<emphasis id="B977EAACFFC6CE60FE43F88048F4F8AC" bold="true" box="[389,407,1845,1869]" pageId="2" pageNumber="93">D</emphasis>
male sternite 9
<emphasis id="B977EAACFFC6CE60FDFFF8804B29F8AC" bold="true" box="[569,586,1845,1869]" pageId="2" pageNumber="93">E</emphasis>
male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E).
</paragraph>
</caption>
<paragraph id="8BBC36BEFFC6CE60FF01F82A4CFFF858" blockId="2.[151,1436,1951,2013]" box="[199,1436,1951,1977]" pageId="2" pageNumber="93">
<emphasis id="B977EAACFFC6CE60FF01F82A4868F858" bold="true" box="[199,267,1951,1977]" pageId="2" pageNumber="93">Male.</emphasis>
Abdominal sternite 8 narrow, with narrow and deep, obtuse medioapical emargination (
<figureCitation id="13382A3BFFC6CE60FAC9F82A4CECF858" box="[1295,1423,1951,1977]" captionStart-0="FIGURE 1–1" captionStart-1="FIGURE 1–2" captionStartId-0="2.[152,255,1808,1832]" captionStartId-1="3.[152,255,1905,1929]" captionTargetBox-0="[152,1431,388,1780]" captionTargetBox-1="[151,1436,273,1882]" captionTargetId-0="figure-104@2.[151,1436,378,1782]" captionTargetId-1="figure-76@3.[151,1436,273,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–1. Miobdelus motuoensis He and Zhou sp. nov., morphology A habitus, dorsal view B aedeagus, lateral view C aedeagus, ventral view D male sternite 9 E male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E)." captionText-1="FIGURE 1–2. Miobdelus motuoensis He and Zhou sp. nov., morphology A male sternite 9 B &amp; D aedeagus, lateral view C aedeagus, ventral view E male tergite 10 F male sternite 8 G underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.13849496" figureDoi-1="http://doi.org/10.5281/zenodo.13849498" httpUri-0="https://zenodo.org/record/13849496/files/figure.png" httpUri-1="https://zenodo.org/record/13849498/files/figure.png" pageId="2" pageNumber="93">Fig. 1–2 F</figureCitation>
).
</paragraph>
<paragraph id="8BBC36BEFFC6CE61FF51F8764C12FF18" blockId="2.[151,1436,1951,2013]" lastBlockId="3.[151,1436,151,249]" lastPageId="3" lastPageNumber="94" pageId="2" pageNumber="93">
Sternite 9 narrow and elongate, with markedly shallow apical emargination; tapered basal portion wide, markedly elongate; apical half very densely setose (
<figureCitation id="13382A3BFFC7CE61FDBDFF224A02FF50" box="[635,865,151,177]" captionStart-0="FIGURE 1–1" captionStart-1="FIGURE 1–2" captionStartId-0="2.[152,255,1808,1832]" captionStartId-1="3.[152,255,1905,1929]" captionTargetBox-0="[152,1431,388,1780]" captionTargetBox-1="[151,1436,273,1882]" captionTargetId-0="figure-104@2.[151,1436,378,1782]" captionTargetId-1="figure-76@3.[151,1436,273,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–1. Miobdelus motuoensis He and Zhou sp. nov., morphology A habitus, dorsal view B aedeagus, lateral view C aedeagus, ventral view D male sternite 9 E male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E)." captionText-1="FIGURE 1–2. Miobdelus motuoensis He and Zhou sp. nov., morphology A male sternite 9 B &amp; D aedeagus, lateral view C aedeagus, ventral view E male tergite 10 F male sternite 8 G underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.13849496" figureDoi-1="http://doi.org/10.5281/zenodo.13849498" httpUri-0="https://zenodo.org/record/13849496/files/figure.png" httpUri-1="https://zenodo.org/record/13849498/files/figure.png" pageId="3" pageNumber="94">Figs. 1–1 D, 1–2 A</figureCitation>
). Tergite 10 elongate, shovel-like, sparsely setose (
<figureCitation id="13382A3BFFC7CE61FF59FF0E481AFF34" box="[159,377,187,213]" captionStart-0="FIGURE 1–1" captionStart-1="FIGURE 1–2" captionStartId-0="2.[152,255,1808,1832]" captionStartId-1="3.[152,255,1905,1929]" captionTargetBox-0="[152,1431,388,1780]" captionTargetBox-1="[151,1436,273,1882]" captionTargetId-0="figure-104@2.[151,1436,378,1782]" captionTargetId-1="figure-76@3.[151,1436,273,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–1. Miobdelus motuoensis He and Zhou sp. nov., morphology A habitus, dorsal view B aedeagus, lateral view C aedeagus, ventral view D male sternite 9 E male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E)." captionText-1="FIGURE 1–2. Miobdelus motuoensis He and Zhou sp. nov., morphology A male sternite 9 B &amp; D aedeagus, lateral view C aedeagus, ventral view E male tergite 10 F male sternite 8 G underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.13849496" figureDoi-1="http://doi.org/10.5281/zenodo.13849498" httpUri-0="https://zenodo.org/record/13849496/files/figure.png" httpUri-1="https://zenodo.org/record/13849498/files/figure.png" pageId="3" pageNumber="94">Figs. 1–1 E, 1–2 E</figureCitation>
). Aedeagus elongate, apical portion with a pair of vertical lip like protrusion (
<figureCitation id="13382A3BFFC7CE61FB28FF0E4862FF18" captionStart-0="FIGURE 1–1" captionStart-1="FIGURE 1–2" captionStartId-0="2.[152,255,1808,1832]" captionStartId-1="3.[152,255,1905,1929]" captionTargetBox-0="[152,1431,388,1780]" captionTargetBox-1="[151,1436,273,1882]" captionTargetId-0="figure-104@2.[151,1436,378,1782]" captionTargetId-1="figure-76@3.[151,1436,273,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–1. Miobdelus motuoensis He and Zhou sp. nov., morphology A habitus, dorsal view B aedeagus, lateral view C aedeagus, ventral view D male sternite 9 E male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E)." captionText-1="FIGURE 1–2. Miobdelus motuoensis He and Zhou sp. nov., morphology A male sternite 9 B &amp; D aedeagus, lateral view C aedeagus, ventral view E male tergite 10 F male sternite 8 G underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.13849496" figureDoi-1="http://doi.org/10.5281/zenodo.13849498" httpUri-0="https://zenodo.org/record/13849496/files/figure.png" httpUri-1="https://zenodo.org/record/13849498/files/figure.png" pageId="3" pageNumber="94">Figs. 1–1 B, C, 1–2 B–D</figureCitation>
); apical portion of paramere elongate and simple, without sensory peg setae, shaped as in
<figureCitation id="13382A3BFFC7CE61FB37FF6A4C0FFF18" box="[1265,1388,223,249]" captionStart-0="FIGURE 1–1" captionStart-1="FIGURE 1–2" captionStartId-0="2.[152,255,1808,1832]" captionStartId-1="3.[152,255,1905,1929]" captionTargetBox-0="[152,1431,388,1780]" captionTargetBox-1="[151,1436,273,1882]" captionTargetId-0="figure-104@2.[151,1436,378,1782]" captionTargetId-1="figure-76@3.[151,1436,273,1882]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1–1. Miobdelus motuoensis He and Zhou sp. nov., morphology A habitus, dorsal view B aedeagus, lateral view C aedeagus, ventral view D male sternite 9 E male tergite 10. Scale bars: 2.00 mm (A); 0.50 mm (B−E)." captionText-1="FIGURE 1–2. Miobdelus motuoensis He and Zhou sp. nov., morphology A male sternite 9 B &amp; D aedeagus, lateral view C aedeagus, ventral view E male tergite 10 F male sternite 8 G underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G)." figureDoi-0="http://doi.org/10.5281/zenodo.13849496" figureDoi-1="http://doi.org/10.5281/zenodo.13849498" httpUri-0="https://zenodo.org/record/13849496/files/figure.png" httpUri-1="https://zenodo.org/record/13849498/files/figure.png" pageId="3" pageNumber="94">Fig. 1–2 G</figureCitation>
.
</paragraph>
<caption id="DF7C6636FFC7CE61FF5EF8C44813F833" ID-DOI="http://doi.org/10.5281/zenodo.13849498" ID-Zenodo-Dep="13849498" httpUri="https://zenodo.org/record/13849498/files/figure.png" pageId="3" pageNumber="94" startId="3.[152,255,1905,1929]" targetBox="[151,1436,273,1882]" targetPageId="3" targetType="figure">
<paragraph id="8BBC36BEFFC7CE61FF5EF8C44813F833" blockId="3.[151,1436,1905,2002]" pageId="3" pageNumber="94">
<emphasis id="B977EAACFFC7CE61FF5EF8C44850F86B" bold="true" box="[152,307,1905,1930]" pageId="3" pageNumber="94">FIGURE 1–2.</emphasis>
<taxonomicName id="4C034D3DFFC7CE61FEFCF8C74BCEF86B" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[314,685,1906,1930]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="94" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC7CE61FEFCF8C74B7CF868" box="[314,543,1906,1930]" italics="true" pageId="3" pageNumber="94">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
<emphasis id="B977EAACFFC7CE61FD73F8C74A65F86B" bold="true" box="[693,774,1906,1930]" pageId="3" pageNumber="94">
<taxonomicNameLabel id="A24457D7FFC7CE61FD73F8C74A65F86B" box="[693,774,1906,1930]" pageId="3" pageNumber="94" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, morphology
<emphasis id="B977EAACFFC7CE61FC5EF8C44AC9F868" bold="true" box="[920,938,1905,1929]" pageId="3" pageNumber="94">A</emphasis>
male sternite 9
<emphasis id="B977EAACFFC7CE61FB97F8C74DFBF86B" bold="true" box="[1105,1176,1905,1930]" pageId="3" pageNumber="94">B &amp; D</emphasis>
aedeagus, lateral view
<emphasis id="B977EAACFFC7CE61FA4CF8C44CFFF868" bold="true" box="[1418,1436,1905,1929]" pageId="3" pageNumber="94">C</emphasis>
aedeagus, ventral view
<emphasis id="B977EAACFFC7CE61FE4DF82348FFF84F" bold="true" box="[395,412,1942,1966]" pageId="3" pageNumber="94">E</emphasis>
male tergite 10
<emphasis id="B977EAACFFC7CE61FD8EF8234B34F84F" bold="true" box="[584,599,1942,1966]" pageId="3" pageNumber="94">F</emphasis>
male sternite 8
<emphasis id="B977EAACFFC7CE61FCC4F8204A75F84C" bold="true" box="[770,790,1941,1965]" pageId="3" pageNumber="94">G</emphasis>
underside of apical portion of paramere. Scale bars: 0.50 mm (A−E); 0.25 mm (G).
</paragraph>
</caption>
<paragraph id="8BBC36BEFFC0CE66FF01FF2248C1FF50" blockId="4.[151,1436,151,609]" box="[199,418,151,177]" pageId="4" pageNumber="95">
<emphasis id="B977EAACFFC0CE66FF01FF224846FF50" bold="true" box="[199,293,151,177]" pageId="4" pageNumber="95">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="C3196535FFC0CE66FF01FF0F4D51FDF8" pageId="4" pageNumber="95" type="distribution">
<paragraph id="8BBC36BEFFC0CE66FF01FF0F4C18FF34" blockId="4.[151,1436,151,609]" box="[199,1403,186,213]" pageId="4" pageNumber="95">
<emphasis id="B977EAACFFC0CE66FF01FF0F4802FF35" bold="true" box="[199,353,186,212]" pageId="4" pageNumber="95">Distribution.</emphasis>
<taxonomicName id="4C034D3DFFC0CE66FEA1FF0E4B9CFF34" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[359,767,187,213]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC0CE66FEA1FF0E4B00FF35" box="[359,611,187,213]" italics="true" pageId="4" pageNumber="95">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
,
<emphasis id="B977EAACFFC0CE66FCCCFF0E4A00FF34" bold="true" box="[778,867,187,213]" pageId="4" pageNumber="95">
<taxonomicNameLabel id="A24457D7FFC0CE66FCCCFF0E4A00FF34" box="[778,867,187,213]" pageId="4" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is at present known only from the
<typeStatus id="54B8881CFFC0CE66FB2CFF0E4C79FF34" box="[1258,1306,187,213]" pageId="4" pageNumber="95">type</typeStatus>
locality.
</paragraph>
<paragraph id="8BBC36BEFFC0CE66FF01FF6A4D51FDF8" blockId="4.[151,1436,151,609]" pageId="4" pageNumber="95">
<emphasis id="B977EAACFFC0CE66FF01FF6A4872FF18" bold="true" box="[199,273,223,249]" pageId="4" pageNumber="95">Notes.</emphasis>
<taxonomicName id="4C034D3DFFC0CE66FEDFFF6A4BD6FF18" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[281,693,223,249]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC0CE66FEDFFF6A4B76FF19" box="[281,533,223,249]" italics="true" pageId="4" pageNumber="95">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
,
<emphasis id="B977EAACFFC0CE66FD07FF6A4A7FFF18" bold="true" box="[705,796,223,249]" pageId="4" pageNumber="95">
<taxonomicNameLabel id="A24457D7FFC0CE66FD07FF6A4A7FFF18" box="[705,796,223,249]" pageId="4" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
can be easily distinguished from other congeners by the uniquely shaped aedeagus, and by the white apical segments of antennae.
<taxonomicName id="4C034D3DFFC0CE66FC11FEB64C0CFEFC" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[983,1391,259,285]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC0CE66FC11FEB64DB1FEFD" box="[983,1234,259,285]" italics="true" pageId="4" pageNumber="95">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
,
<emphasis id="B977EAACFFC0CE66FABDFEB649ABFEA0" bold="true" pageId="4" pageNumber="95">
<taxonomicNameLabel id="A24457D7FFC0CE66FABDFEB649ABFEA0" pageId="4" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is most similar to
<taxonomicName id="4C034D3DFFC0CE66FE5AFE924ADBFEA0" authority="He and Zhou 2018" authorityName="He and Zhou" authorityYear="2018" box="[412,952,295,321]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="chrysochromatus">
<emphasis id="B977EAACFFC0CE66FE5AFE924BB9FEA0" box="[412,730,295,321]" italics="true" pageId="4" pageNumber="95">Miobdelus chrysochromatus</emphasis>
<bibRefCitation id="EF924B4FFFC0CE66FD24FE924ADBFEA0" author="He, L. &amp; Zhou, H. - Z." box="[738,952,295,321]" pageId="4" pageNumber="95" pagination="301 - 353" refId="ref2925" refString="He, L. &amp; Zhou, H. - Z. (2018) Taxonomy of the genus Miobdelus Sharp, 1889 (Coleoptera: Staphylinidae: Staphylinini) and five new species from China. Zootaxa, 4377 (3), 301 - 353. https: // doi. org / 10.11646 / zootaxa. 4377.3.1" type="journal article" year="2018">He and Zhou 2018</bibRefCitation>
</taxonomicName>
; both species have pale apical segments of antennae, and they both have elongate apical portion of paramere, lacking sensory peg setae. However, they can be easily distinguished from each other:
<taxonomicName id="4C034D3DFFC0CE66FDF3FEDA4AA4FE68" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[565,967,367,393]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC0CE66FDF3FEDA4A4CFE69" box="[565,815,367,393]" italics="true" pageId="4" pageNumber="95">Miobdelus motuoensis</emphasis>
He and Zhou
</taxonomicName>
,
<emphasis id="B977EAACFFC0CE66FC17FEDA4D4AFE68" bold="true" box="[977,1065,367,393]" pageId="4" pageNumber="95">
<taxonomicNameLabel id="A24457D7FFC0CE66FC17FEDA4D4AFE68" box="[977,1065,367,393]" pageId="4" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is much duller, and not decorated with any bright pubescence on elytra and abdominal tergites like
<taxonomicName id="4C034D3DFFC0CE66FCA8FE264D30FE4C" authorityName="He and Zhou" authorityYear="2018" box="[878,1107,403,429]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="chrysochromatus">
<emphasis id="B977EAACFFC0CE66FCA8FE264D30FE4C" box="[878,1107,403,429]" italics="true" pageId="4" pageNumber="95">M. chrysochromatus</emphasis>
</taxonomicName>
(except on visible abdominal tergite 4); the apical antennae segments of
<taxonomicName id="4C034D3DFFC0CE66FDBDFE024A02FE30" authorityName="He and Zhou" authorityYear="2018" box="[635,865,439,465]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="chrysochromatus">
<emphasis id="B977EAACFFC0CE66FDBDFE024A02FE30" box="[635,865,439,465]" italics="true" pageId="4" pageNumber="95">M. chrysochromatus</emphasis>
</taxonomicName>
are gradually becoming paler towards apex, while in
<taxonomicName id="4C034D3DFFC0CE66FF72FE6E4890FE14" authority="He and Zhou" authorityName="He and Zhou" authorityYear="2024" box="[180,499,475,501]" class="Insecta" family="Staphylinidae" genus="Miobdelus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="95" phylum="Arthropoda" rank="species" species="motuoensis" status="sp. nov.">
<emphasis id="B977EAACFFC0CE66FF72FE6E4834FE15" box="[180,343,475,500]" italics="true" pageId="4" pageNumber="95">M. motuoensis</emphasis>
He and Zhou
</taxonomicName>
,
<emphasis id="B977EAACFFC0CE66FE3BFE6E4B34FE14" bold="true" box="[509,599,475,501]" pageId="4" pageNumber="95">
<taxonomicNameLabel id="A24457D7FFC0CE66FE3BFE6E4B34FE14" box="[509,599,475,501]" pageId="4" pageNumber="95" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
antennal segments 8 to 11 are distinctly paler than other segments; and the median lobe of the aedeagus is completely differently shaped in these two species.
</paragraph>
</subSubSection>
<subSubSection id="C3196535FFC0CE66FF01FD96498AFD80" pageId="4" pageNumber="95" type="etymology">
<paragraph id="8BBC36BEFFC0CE66FF01FD96498AFD80" blockId="4.[151,1436,151,609]" pageId="4" pageNumber="95">
<emphasis id="B977EAACFFC0CE66FF01FD96482DFDDC" bold="true" box="[199,334,547,573]" pageId="4" pageNumber="95">Etymology.</emphasis>
The species epithet is derived from the Chinese name of the
<typeStatus id="54B8881CFFC0CE66FBF4FD964D01FDDC" box="[1074,1122,547,573]" pageId="4" pageNumber="95">type</typeStatus>
locality in
<collectingCountry id="F314762EFFC0CE66FB29FD964C51FDDC" box="[1263,1330,547,573]" name="China" pageId="4" pageNumber="95">China</collectingCountry>
,
<collectingRegion id="49C7F85CFFC0CE66FA82FD964CFBFDDC" box="[1348,1432,547,573]" country="China" name="Xizang" pageId="4" pageNumber="95">Xizang</collectingRegion>
, Motuo.
</paragraph>
</subSubSection>
</treatment>
</document>
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