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<document id="2AC7C7217294132F8C76E108B74A9D47" ID-DOI="10.11646/zootaxa.3664.2.1" ID-GBIF-Dataset="a50b71f5-48a6-464f-aea1-f29bbc8ee322" ID-ISSN="1175-5326" ID-Zenodo-Dep="220503" ID-ZooBank="105D178E-6DBF-401D-85BB-CDB041B67E91" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460615203542" checkinUser="plazi" docAuthor="Shi, Hongliang, Sciaky, Riccardo, Liang, Hongbin &amp; Zhou, Hongzhang" docDate="2013" docId="03AA10085655FFE6C1D8FD69FDEFF8D2" docLanguage="en" docName="zt03664p135.pdf" docOrigin="Zootaxa 3664 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pterostichus (Wraseiellus) pseudodiversus Shi &amp; Sciaky, new species" docType="treatment" docVersion="9" lastPageNumber="116" masterDocId="FF936870565BFFE9C14FFF95FFA9FF9E" masterDocTitle="A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions" masterLastPageNumber="135" masterPageNumber="101" pageNumber="115" updateTime="1698330049886" updateUser="plazi">
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<mods:title id="41B2BB228CB1D7967455AC836CE6680A">A new subgenus Wraseiellus of the genus Pterostichus Bonelli (Coleoptera, Carabidae, Pterostichini) and new species descriptions</mods:title>
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<mods:namePart id="A4EAF4E409E4C18F1C7EA63F3479E584">Shi, Hongliang</mods:namePart>
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<mods:namePart id="2A1AAA38AC1FE233AF5238080E927D4B">Sciaky, Riccardo</mods:namePart>
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<mods:namePart id="E5CE711EB05DCE3EBF2A385523A5759E">Liang, Hongbin</mods:namePart>
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<mods:namePart id="1378AF19BA58C5B1E5FB053095020AD6">Zhou, Hongzhang</mods:namePart>
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<heading id="D0F416725655FFE7C1D8FD69FC70FC8B" bold="true" box="[151,985,763,790]" fontSize="11" level="1" pageId="14" pageNumber="115" reason="1">
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<taxonomicName id="4C03DA9D5655FFE7C1D8FD69FC95FC8B" ID-CoL="4Q4HR" authority="Shi &amp; Sciaky" authorityName="Shi &amp; Sciaky" box="[151,828,763,790]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="pseudodiversus" status="sp. nov." subGenus="Wraseiellus">
<emphasis id="B9777D0C5655FFE7C1D8FD69FE84FC88" bold="true" box="[151,301,764,790]" italics="true" pageId="14" pageNumber="115">Pterostichus</emphasis>
(
<emphasis id="B9777D0C5655FFE7C072FD69FE61FC88" bold="true" box="[317,456,764,790]" italics="true" pageId="14" pageNumber="115">Wraseiellus</emphasis>
)
<emphasis id="B9777D0C5655FFE7C095FD69FD3BFC88" bold="true" box="[474,658,764,790]" italics="true" pageId="14" pageNumber="115">pseudodiversus</emphasis>
Shi &amp; Sciaky
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,
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(
<figureCitation id="1338BD9B5655FFE7C1D1FC8BFE3EFCA8" box="[158,407,798,822]" captionStart="FIGURES 13 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="14" pageNumber="115">Figures 13, 14, 15, 16</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C0ECFC8BFE68FCA8" box="[419,449,798,822]" captionStart="FIGURES 19 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="14" pageNumber="115">26</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C082FC8BFDE9FCA8" box="[461,576,798,822]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="14" pageNumber="115">37, 38, 39</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C304FC8BFD3DFCA8" box="[587,660,798,822]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="14" pageNumber="115">4556</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C3D0FC8BFD14FCA8" box="[671,701,798,822]" captionStart="FIGURES 57 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="14" pageNumber="115">62</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C386FC8BFCB8FCA8" box="[713,785,798,822]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">7984</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C252FC8BFCCCFCA8" box="[797,869,798,822]" captionStart="FIGURES 85 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="14" pageNumber="115">85, 89</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C23EFC8BFC26FCA8" box="[881,911,798,822]" captionStart="FIGURES 90 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="14" pageNumber="115">92</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C2D4FC8BFC6FFCA8" box="[923,966,798,822]" captionStart="FIGURES 98 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="14" pageNumber="115">103</figureCitation>
)
</paragraph>
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locality
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.
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: Yunnan province, Ninglang County, Luguhu, Gouzuandong pass,
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.
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(IZAS), male, body length =
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, pinned, genitalia dissected and glued on plastic film pinned under specimen, &quot;
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, Yunnan, / Lijiang, Ninglang, / Luguhu, Gouzuandong /
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; 2009.
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/ SHI H.L. coll. PTF trap / Inst. Of Zoology, CAS / &quot;; &quot;IOZ(E)1891200&quot;; &quot;
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3/
<taxonomicName id="4C03DA9D5655FFE7C307FB8FFCDAFBAC" authority="Wraseiellus" authorityName="Wraseiellus" box="[584,883,1050,1074]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="genus">Pterostichus (Wraseiellus)</taxonomicName>
/
<taxonomicName id="4C03DA9D5655FFE7C2DEFB8FFB97FBAC" box="[913,1086,1050,1074]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="pseudodiversus" status="sp. nov.">pseudodiversus</taxonomicName>
<taxonomicNameLabel id="A244C0775655FFE7C506FB8EFB7DFBAC" box="[1097,1236,1050,1074]" pageId="14" pageNumber="115" rank="species">new species</taxonomicNameLabel>
/ Des. SHI H.L. 2012&quot; [red label] [
<figureCitation id="1338BD9B5655FFE7C02CFBABFE3EFBD2" box="[355,407,1086,1100]" captionStart="FIGURES 13 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="14" pageNumber="115">Figs. 13</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C0D0FBABFE19FBD2" box="[415,432,1086,1100]" captionStart="FIGURES 19 28" captionStartId="24.[151,264,1308,1331]" captionTargetBox="[166,1426,208,1270]" captionTargetId="figure@24.[151,1436,193,1299]" captionTargetPageId="24" captionText="FIGURES 19 28. Labels of name-bearing type specimen: 19, lectotype of Pterostichus andrewesi Jedlička, 1931; 20, holotype of Pterostichus meyeri Jedlička, 1934; 21, holotype of Feronia crebrepunctata Straneo, 1937; 22, holotype of Pterostichus comatus n. sp.; 23, holotype of Pterostichus crassiapex n. sp.; 24, a syntype of Omaseus diversus Fairmaire, 1886; 25, holotype of Pterostichus stictopleurus cangshanensis n. ssp.; 26, holotype of Pterostichus pseudodiversus n. sp.; 27, holotype of Feronia kambaiti Andrewes, 1947 [photoed by Dr. J. Bergsten]; 28, holotype of Pterostichus kambaiti Jedlička, 1965." httpUri="https://zenodo.org/record/220507/files/figure.png" pageId="14" pageNumber="115">26</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C0F8FBABFE61FBD2" box="[439,456,1086,1100]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="14" pageNumber="115">37</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C081FBABFE76FBD2" box="[462,479,1086,1100]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="14" pageNumber="115">45</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C0A9FBABFE5EFBD2" box="[486,503,1086,1100]" captionStart="FIGURES 57 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="14" pageNumber="115">62</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C0B2FBABFDA7FBD2" box="[509,526,1086,1100]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">79</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C35AFBABFD84FBD2" box="[533,557,1086,1100]" captionStart="FIGURES 98 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="14" pageNumber="115">103</figureCitation>
].
<emphasis id="B9777D0C5655FFE7C30EFBD7FD10FBC4" bold="true" box="[577,697,1090,1114]" pageId="14" pageNumber="115">
<typeStatus id="54B81FBC5655FFE7C30EFBD7FD10FBC4" box="[577,697,1090,1114]" pageId="14" pageNumber="115" type="paratype">Paratypes</typeStatus>
</emphasis>
(total
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):
<emphasis id="B9777D0C5655FFE7C2FAFBD7FBBDFBC4" bold="true" box="[949,1044,1090,1114]" pageId="14" pageNumber="115">Luguhu</emphasis>
:
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(IZAS), the same label data as the
<typeStatus id="54B81FBC5655FFE7C06BFBFDFE20FB1E" box="[292,393,1128,1152]" pageId="14" pageNumber="115" type="holotype">holotype</typeStatus>
, but IOZ(E) database number 18912011891203 [
<figureCitation id="1338BD9B5655FFE7C298FBF0FB81FBED" box="[983,1064,1125,1139]" captionStart="FIGURES 85 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="14" pageNumber="115">Figs. 85, 89</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C57EFBF0FBEBFBED" box="[1073,1090,1125,1139]" captionStart="FIGURES 90 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="14" pageNumber="115">92</figureCitation>
].
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(SCAU), &quot;Yunnan, Luguhu, 2002.08, Tian Mingyi leg.&quot;.
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(IZAS), &quot;
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, Yunnan prov., Ninglang county, south of Luguhu lake, Gouzuandong pass,
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day,
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<figureCitation id="1338BD9B5655FFE7C29BFB41FBADFB7C" box="[980,1028,1236,1250]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="14" pageNumber="115">Fig. 47</figureCitation>
].
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(
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,
<specimenCount id="9D056A975655FFE7C4CFFB42FF47FA8D" pageId="14" pageNumber="115" type="female">11 females</specimenCount>
in IZAS;
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in CRS), &quot;
<collectingCountry id="F314E18E5655FFE7C3F4FB6EFCB0FA8D" box="[699,793,1275,1299]" name="China" pageId="14" pageNumber="115">CHINA</collectingCountry>
, Yunnan prov., Ninglang county, south of Luguhu lake, Gouzuandong pass,
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&quot;; &quot;mixed forest with pine and rhododendra, by pitfall trap, SHI Hongliang, LIU Ye leg. Institute of Zoology, CAS&quot; [
<figureCitation id="1338BD9B5655FFE7C28CFAD6FBECFACF" box="[963,1093,1347,1361]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="14" pageNumber="115">Figs. 46, 48, 49, 50</figureCitation>
].
<emphasis id="B9777D0C5655FFE7C515FAD0FB61FAC0" bold="true" box="[1114,1224,1349,1374]" pageId="14" pageNumber="115">Lanping:</emphasis>
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(SCAU), &quot;Yunnan, Nujiang, Lanping,
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, 2002.7.20, Tian Mingyi leg.&quot; [
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,
<figureCitation id="1338BD9B5655FFE7C2A8FAFCFC51FAE9" box="[999,1016,1385,1399]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">83</figureCitation>
].
<emphasis id="B9777D0C5655FFE7C541FAF9FBFCFA1B" bold="true" box="[1038,1109,1388,1413]" pageId="14" pageNumber="115">Deqin</emphasis>
:
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,
<specimenCount id="9D056A975655FFE7C59BFAF9FA91FA1A" box="[1236,1336,1388,1412]" pageId="14" pageNumber="115" type="female">1 female</specimenCount>
(IZAS), &quot;Yunnan, Deqin, Baimangxueshan, (E.slope)
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&quot;; &quot;1987.
<date id="FFBD87DE5655FFE7C21CFA06FC03FA35" box="[851,938,1427,1451]" pageId="14" pageNumber="115" value="1926-08">VIII.26</date>
, Liao Subai leg.&quot; [
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,
<figureCitation id="1338BD9B5655FFE7C5F3FA05FB64FA00" box="[1212,1229,1424,1438]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="14" pageNumber="115">55</figureCitation>
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].
<emphasis id="B9777D0C5655FFE7C5B5FA06FAD4FA32" bold="true" box="[1274,1405,1427,1452]" pageId="14" pageNumber="115">Zhongdian</emphasis>
:
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(CRS), &quot;CHINA-YUNNAN, 16
<date id="FFBD87DE5655FFE7C301FA2FFD0DFA4C" box="[590,676,1466,1490]" pageId="14" pageNumber="115" value="94-06-21">21.6.94</date>
ZHONGDIAN, lgt. E. Kučera&quot; [
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,
<figureCitation id="1338BD9B5655FFE7C511FA22FBC7FA5B" box="[1118,1134,1463,1477]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">81</figureCitation>
].
<emphasis id="B9777D0C5655FFE7C5CCFA2FFB14FA4D" bold="true" box="[1155,1213,1466,1491]" pageId="14" pageNumber="115">Muli</emphasis>
:
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,
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(IZAS), &quot;
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, Sichuan, Mianbu pass btw. Yanyuan and Muli,
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mixed forest,
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&quot;; &quot;2012.
<date id="FFBD87DE5655FFE7C1ABF990FEBFF983" box="[228,278,1541,1565]" pageId="14" pageNumber="115">VI.7</date>
, pitfall trap, SHI Hongliang, LIU Ye leg. Institute of Zoology, CAS &quot; [
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,
<figureCitation id="1338BD9B5655FFE7C52BF997FBDCF98E" box="[1124,1141,1538,1552]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="14" pageNumber="115">38</figureCitation>
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,
<figureCitation id="1338BD9B5655FFE7C5E6F997FB13F98E" box="[1193,1210,1538,1552]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">80</figureCitation>
].
</paragraph>
<paragraph id="8BBCA11E5655FFE7C188F9BCFAE7F9F9" blockId="14.[151,1436,870,2036]" pageId="14" pageNumber="115">
<emphasis id="B9777D0C5655FFE7C188F9BCFD8CF9DF" bold="true" box="[199,549,1576,1601]" pageId="14" pageNumber="115">Non-type materials examined</emphasis>
.
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(IZAS), &quot;
<collectingCountry id="F314E18E5655FFE7C212F9BCFC12F9DF" box="[861,955,1577,1601]" name="China" pageId="14" pageNumber="115">CHINA</collectingCountry>
, Yunnan, Diqing, Weixi, Pantiange, Zhazi;
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,
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&quot;; &quot;
<quantity id="4CFB0CFB5655FFE7C0F0F9DAFDBAF9F9" box="[447,531,1615,1639]" metricMagnitude="3" metricUnit="m" metricValue="3.029" pageId="14" pageNumber="115" unit="m" value="3029.0">3029m</quantity>
; 2009.
<date id="FFBD87DE5655FFE7C32FF9DAFD03F9F9" box="[608,682,1615,1639]" pageId="14" pageNumber="115" value="2012-07">VII.12</date>
, Shi H.L. coll. PTF trap Inst. of Zoology, CAS&quot; [
<figureCitation id="1338BD9B5655FFE7C585F9D8FB56F9C5" box="[1226,1279,1613,1627]" captionStart="FIGURES 13 18" captionStartId="23.[151,264,1749,1772]" captionTargetBox="[158,1427,204,1715]" captionTargetId="figure@23.[151,1436,193,1728]" captionTargetPageId="23" captionText="FIGURES 13 18. Habitus of Pterostichus (Wraseiellus) spp.: scale bar = 2.0 mm; 13, Pt. pseudodiversus n. sp., holotype; 14, Pt. pseudodiversus n. sp., a female paratype from Muli; 15, Pt. pseudodiversus n. sp., a male paratype from Deqin; 16, Pt. pseudodiversus n. sp., a male from Weixi; 17, Pt. kambaiti (Andrewes), holotype [photoed by Dr. J. Bergsten]; 18, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220506/files/figure.png" pageId="14" pageNumber="115">Figs. 16</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C44AF9D8FABFF9C5" box="[1285,1302,1613,1627]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="14" pageNumber="115">39</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C453F9D8FA84F9C5" box="[1308,1325,1613,1627]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="14" pageNumber="115">56</figureCitation>
,
<figureCitation id="1338BD9B5655FFE7C47CF9D8FAEDF9C5" box="[1331,1348,1613,1627]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">84</figureCitation>
].
</paragraph>
</subSubSection>
<subSubSection id="C319F2955655FFE6C188F9E6FB5DFF69" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115" type="diagnosis">
<paragraph id="8BBCA11E5655FFE7C188F9E6FB8BF969" blockId="14.[151,1436,870,2036]" pageId="14" pageNumber="115">
<emphasis id="B9777D0C5655FFE7C188F9E6FE93F912" bold="true" box="[199,314,1651,1676]" pageId="14" pageNumber="115">Diagnosis</emphasis>
. From the shape of pronotum and male genital characters, this new species is very similar to
<taxonomicName id="4C03DA9D5655FFE7C1D8F90DFDF6F931" authority="Fairmaire. From" authorityName="Fairmaire. From" box="[151,607,1687,1712]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C1D8F90DFE39F92E" box="[151,400,1688,1712]" italics="true" pageId="14" pageNumber="115">Pterostichus diversus</emphasis>
Fairmaire. From
</taxonomicName>
all the other members of this subgenus, these two species can be distinguished by: (1) 5th tarsomere usually without seta beneath; (2) a distinct ridge present between outer basal foveal groove and lateral margin; (3) right paramere long and curved, apex wide.
</paragraph>
<paragraph id="8BBCA11E5655FFE6C188F891FB5DFF69" blockId="14.[151,1436,870,2036]" lastBlockId="15.[151,1437,151,1868]" lastPageId="15" lastPageNumber="116" pageId="14" pageNumber="115">
The new species is difficult to be distinguished from
<taxonomicName id="4C03DA9D5655FFE7C25BF890FC31F882" box="[788,920,1796,1820]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C25BF890FC31F882" box="[788,920,1796,1820]" italics="true" pageId="14" pageNumber="115">Pt. diversus</emphasis>
</taxonomicName>
in external characters, but their male genitalia show that they are different species. Compared with
<taxonomicName id="4C03DA9D5655FFE7C3A4F8BCFCD9F8DE" box="[747,880,1832,1856]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C3A4F8BCFCD9F8DE" box="[747,880,1832,1856]" italics="true" pageId="14" pageNumber="115">Pt. diversus</emphasis>
</taxonomicName>
, the new species is different in: (1) area between inner and outer basal foveal grooves usually less convex than in
<taxonomicName id="4C03DA9D5655FFE7C225F8D8FC47F8FA" box="[874,1006,1868,1892]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C225F8D8FC47F8FA" box="[874,1006,1868,1892]" italics="true" pageId="14" pageNumber="115">Pt. diversus</emphasis>
</taxonomicName>
(
<figureCitation id="1338BD9B5655FFE7C2B1F8D9FB0BF8FA" box="[1022,1186,1868,1892]" captionStart="FIGURES 98 106" captionStartId="32.[151,264,1472,1495]" captionTargetBox="[155,1435,216,1425]" captionTargetId="figure@32.[151,1436,215,1427]" captionTargetPageId="32" captionText="FIGURES 98 106. Pronotum hind angle and basal area (right side) of Pterostichus (Wraseiellus) spp.; scale bar = 0.5 mm; 98, Pt. andrewesi Jedlička, a female from Jinfoshan; 99, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 100, Pt. crassiapex n. sp., holotype; 101, Pt. comatus n. sp., holotype; 102, Pt. diversus (Fairmaire), a male from Yulongshan; 103, Pt. pseudodiversus n. sp., holotype; 104, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 105, Pt. stictopleurus cangshanensis n. ssp., holotype; 106, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220515/files/figure.png" pageId="14" pageNumber="115">Figs. 102, 103</figureCitation>
); (2) 3rd antennomere with few additional setae, while in
<taxonomicName id="4C03DA9D5655FFE7C379F8E4FD14F816" box="[566,701,1904,1928]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C379F8E4FD14F816" box="[566,701,1904,1928]" italics="true" pageId="14" pageNumber="115">Pt. diversus</emphasis>
</taxonomicName>
3rd antennomere usually glabrous except apical ring; (3) right paramere shorter than in
<taxonomicName id="4C03DA9D5655FFE7C0FEF800FD9CF832" box="[433,565,1940,1964]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C0FEF800FD9CF832" box="[433,565,1940,1964]" italics="true" pageId="14" pageNumber="115">Pt. diversus</emphasis>
</taxonomicName>
, apex rounded or slightly pointed, never truncate (
<figureCitation id="1338BD9B5655FFE7C53FF801FB67F832" box="[1136,1230,1940,1964]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">Figs. 79</figureCitation>
B84B), but in
<taxonomicName id="4C03DA9D5655FFE7C433F800FF5DF84E" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5655FFE7C433F800FF5DF84E" italics="true" pageId="14" pageNumber="115">Pt. diversus</emphasis>
</taxonomicName>
, right paramere longer and apex usually truncate (truncate in
<figureCitation id="1338BD9B5655FFE7C2EEF82DFC55F84E" box="[929,1020,1976,2000]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">Figs. 74</figureCitation>
B76B, rarely slightly pointed in
<figureCitation id="1338BD9B5655FFE7C43FF82DFF1FF86A" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="14" pageNumber="115">Fig. 77</figureCitation>
B); (4) in
<taxonomicName id="4C03DA9D5655FFE7C06EF848FE5DF86A" box="[289,500,2012,2036]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="115" phylum="Arthropoda" rank="species" species="pseudodiversus">
<emphasis id="B9777D0C5655FFE7C06EF848FE5DF86A" box="[289,500,2012,2036]" italics="true" pageId="14" pageNumber="115">Pt. pseudodiversus</emphasis>
</taxonomicName>
, median lobe of aedeagus stouter (
<figureCitation id="1338BD9B5655FFE7C237F849FC7AF86A" box="[888,979,2012,2036]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="14" pageNumber="115">Figs. 37</figureCitation>
A39A), in lateral view, apex not bent to the dorsal side (
<figureCitation id="1338BD9B5654FFE6C008FF02FE33FF31" box="[327,410,151,175]" captionStart="FIGURES 57 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="15" pageNumber="116">Fig. 62</figureCitation>
B), apex of apical lamella narrow or slightly truncate, slightly bent to the left side (
<figureCitation id="1338BD9B5654FFE6C47CFF02FF1EFF4A" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Figs. 45 56</figureCitation>
); but in
<taxonomicName id="4C03DA9D5654FFE6C05EFF28FE3CFF4A" box="[273,405,188,212]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5654FFE6C05EFF28FE3CFF4A" box="[273,405,188,212]" italics="true" pageId="15" pageNumber="116">Pt. diversus</emphasis>
</taxonomicName>
, median lobe of aedeagus slenderer (
<figureCitation id="1338BD9B5654FFE6C27CFF29FC26FF4A" box="[819,911,188,212]" captionStart="FIGURES 29 34" captionStartId="25.[151,264,1854,1877]" captionTargetBox="[176,1418,197,1824]" captionTargetId="figure@25.[151,1436,193,1833]" captionTargetPageId="25" captionText="FIGURES 29 34. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe. 29, Pt. andrewesi Jedlička, a male from Jinfoshan; 30, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 31, Pt. crassiapex n. sp., holotype; 32, Pt. comatus n. sp., holotype; 33, Pt. diversus (Fairmaire), a male from Yulongshan; 34, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;." httpUri="https://zenodo.org/record/220508/files/figure.png" pageId="15" pageNumber="116">Figs. 33</figureCitation>
A, 34A), in lateral view, apex distinctly bent to the dorsal side (
<figureCitation id="1338BD9B5654FFE6C007FF4AFE33FF69" box="[328,410,223,247]" captionStart="FIGURES 57 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="15" pageNumber="116">Fig. 61</figureCitation>
B), apex of apical lamella capitate and not bent to the left side (
<figureCitation id="1338BD9B5654FFE6C511FF4AFB4EFF69" box="[1118,1255,223,247]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Figs. 41, 42</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955654FFE6C188FE96FEFFFB5A" pageId="15" pageNumber="116" type="description">
<paragraph id="8BBCA11E5654FFE6C188FE96FEFFFB5A" blockId="15.[151,1437,151,1868]" pageId="15" pageNumber="116">
<emphasis id="B9777D0C5654FFE6C188FE96FEF8FE82" bold="true" box="[199,337,259,284]" pageId="15" pageNumber="116">Description</emphasis>
. Body length
<quantity id="4CFB0CFB5654FFE6C0B3FE91FD0FFE82" box="[508,678,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.335" metricValueMax="1.49" metricValueMin="1.18" pageId="15" pageNumber="116" unit="mm" value="13.35" valueMax="14.9" valueMin="11.8">11.814.9 mm</quantity>
; dorsal side black or reddish brown, shining in males; legs dark brown, tarsi brown; ventral side black or dark brown. Males with elytral microsculpture isodiametric.
<emphasis id="B9777D0C5654FFE6C441FEBDFAE7FEDE" bold="true" box="[1294,1358,296,320]" pageId="15" pageNumber="116">Head</emphasis>
. Frons and vertex impunctate or sparsely punctate; antennae reaching elytra basal eighth; inner margin of 3rd antennomere usually with few additional setae subapically, but glabrous in some specimens; males with the terminal labial palpomere similar as in females, slightly expanded and truncate.
<emphasis id="B9777D0C5654FFE6C220FE01FC41FE32" bold="true" box="[879,1000,404,428]" pageId="15" pageNumber="116">Pronotum</emphasis>
rounded, widest a little before middle, PW/PL = 1.261.46; lateral margins fully rounded near the middle, slightly sinuate just before hind angles; hind angles slightly pointed; one mid-lateral seta present at about anterior third; basal fovea moderately deep, faintly defined; inner groove of basal fovea subparallel to median line, slightly curved to the outside posteriorly, ended at a distance from the posterior margin of pronotum; outer groove of basal fovea about half length as inner one, as deep as inner one, reaching or almost reaching the basal margin of pronotum; ridge between outer groove and lateral margin distinct; area between inner and outer basal foveal grooves feebly convex and punctate; basal fovea area punctate, some sparser punctures usually present on the area between two basal foveae.
<emphasis id="B9777D0C5654FFE6C598FD05FA8AFD36" bold="true" box="[1239,1315,656,680]" pageId="15" pageNumber="116">Elytra</emphasis>
with base slightly depressed between 3rd and 5th intervals; elytral shoulder slightly narrowed, humeral teeth small but distinct; intervals feebly convex; striae moderately deep, with very faint punctures; scutellar striae long, usually incomplete, located between 1st stria and elytral suture; umbilical pore series on 9th interval sparse in the middle, composed of 1618 pores (67, 23, 78).
<emphasis id="B9777D0C5654FFE6C331FCB5FCB8FCA6" bold="true" box="[638,785,799,824]" pageId="15" pageNumber="116">Ventral side</emphasis>
. Proepisterna and metaepisterna finely punctate; terminal sternum of males without special structure.
<emphasis id="B9777D0C5654FFE6C331FCD1FD1FFCC2" bold="true" box="[638,694,836,860]" pageId="15" pageNumber="116">Legs</emphasis>
. 5th tarsomere without seta beneath.
<emphasis id="B9777D0C5654FFE6C518FCD1FAA8FCC2" bold="true" box="[1111,1281,835,860]" pageId="15" pageNumber="116">Male genitalia</emphasis>
. Median lobe of male genitalia bent at a 90 degrees angle at about basal two-fifths; ventral margin almost straight in the middle, apical part not bent to the ventral side (
<figureCitation id="1338BD9B5654FFE6C31CFC19FD06FC3A" box="[595,687,908,932]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="15" pageNumber="116">Figs. 37</figureCitation>
A39A); apical orifice large, slightly turned to the left side, apical lamella slightly twisted to the left (
<figureCitation id="1338BD9B5654FFE6C36DFC3AFDD6FC59" box="[546,639,943,967]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="15" pageNumber="116">Figs. 37</figureCitation>
B39B); apical lamella laminar, 1.11.6 times as the basal width, apex bent to the left, rounded or slightly truncate (
<figureCitation id="1338BD9B5654FFE6C3ECFC41FC98FC72" box="[675,817,980,1004]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Figs. 4556</figureCitation>
); apex of apical lamella not turned to the dorsal side, ventral surface slightly thickened (
<figureCitation id="1338BD9B5654FFE6C36DFC62FDD1FB91" box="[546,632,1015,1039]" captionStart="FIGURES 57 66" captionStartId="28.[151,264,1811,1834]" captionTargetBox="[167,1430,195,1775]" captionTargetId="figure@28.[151,1436,179,1792]" captionTargetPageId="28" captionText="FIGURES 57 66. Apex of median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, dorsal view of apical lamella; B, left lateral view of apical lamella; C, right lateral view of apical lamella; 57, Pt. andrewesi Jedlička, a male from Jinfoshan; 58, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 59, Pt. crassiapex n. sp., holotype; 60, Pt. comatus n. sp., holotype; 61, Pt. diversus (Fairmaire), a male from Yulongshan; 62, Pt. pseudodiversus n. sp., holotype; 63, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 64, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 65, Pt. stictopleurus cangshanensis n. ssp., holotype; 66, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan." httpUri="https://zenodo.org/record/220511/files/figure.png" pageId="15" pageNumber="116">Fig. 62</figureCitation>
B). Right paramere laminar, elongate and curved, right margin curved, narrowed apically, apex rounded or slightly pointed (
<figureCitation id="1338BD9B5654FFE6C3BCFB89FCFBFBAA" box="[755,850,1052,1076]" captionStart="FIGURES 67 84" captionStartId="29.[151,264,1584,1607]" captionTargetBox="[166,1426,201,1530]" captionTargetId="figure@29.[151,1436,193,1545]" captionTargetPageId="29" captionText="FIGURES 67 84. Parameres of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, left paramere; B, right paramere: 67, Pt. andrewesi Jedlička, a male from Jinfoshan; 68, Pt. andrewesi Jedlička, a paralectotype from Jinfoshan; 69, Pt. meyeri Jedlička, a male from &quot; Yunnan &quot;; 70, Pt. comatus n. sp., holotype; 71, Pt. crassiapex n. sp., holotype; 72, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 73, Pt. stictopleurus cangshanensis n. ssp., holotype; 74 75, Pt. diversus (Fairmaire), males from Yulongshan; 76, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 77, Pt. diversus (Fairmaire), a male from &quot; Yunnan &quot;; 78, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička; 79, Pt. pseudodiversus n. sp., holotype; 80, Pt. pseudodiversus n. sp., a paratype from Muli; 81, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 82, Pt. pseudodiversus n. sp., a paratype from Deqin; 83, Pt. pseudodiversus n. sp.., a paratype from Lanping; 84, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220512/files/figure.png" pageId="15" pageNumber="116">Figs. 79</figureCitation>
B84B).
<emphasis id="B9777D0C5654FFE6C2F8FB89FBD7FBAA" bold="true" box="[951,1150,1051,1076]" pageId="15" pageNumber="116">Female genitalia</emphasis>
similar with
<taxonomicName id="4C03DA9D5654FFE6C459FB88FEADFBC9" authority="Fairmaire" authorityName="Fairmaire" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5654FFE6C459FB88FA35FBAA" box="[1302,1436,1052,1076]" italics="true" pageId="15" pageNumber="116">Pt. diversus</emphasis>
Fairmaire
</taxonomicName>
(
<figureCitation id="1338BD9B5654FFE6C05AFBAAFEC2FBC9" box="[277,363,1087,1111]" captionStart="FIGURES 85 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="15" pageNumber="116">Fig. 89</figureCitation>
). Spermatheca with the seminal canal about ten times long as the receptaculum; spermathecal gland strongly expanded; the seminal canal inserted at the base of common oviduct, base of seminal canal sclerotized (
<figureCitation id="1338BD9B5654FFE6C06EFB1DFEDFFB3E" box="[289,374,1160,1184]" captionStart="FIGURES 85 89" captionStartId="30.[151,264,1685,1708]" captionTargetBox="[183,1411,216,1645]" captionTargetId="figure@30.[151,1436,193,1664]" captionTargetPageId="30" captionText="FIGURES 85 89. Female reproductive system of Pterostichus (Wraseiellus) spp.: scale bar A = 1.0 mm; scale bar B = 0.5 mm; A, spermatheca; B, receptaculum: 85, internal reproductive system structures (weakly-stained) of Pt. pseudodiversus n. sp., a paratype from Luguhu; 86, Pt. andrewesi Jedlička, a female from Jinfoshan; 87, Pt. diversus (Fairmaire), a female from Yulongshan; 88, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 89, Pt. pseudodiversus n. sp., a paratype from Luguhu. Abbreviations: bc, bursa copulatrix; co, common oviduct; rc, receptaculum; s 1, stylomere 1; s 2, stylomere 2; sc, seminal canal; sg, spermathecal gland; sp, spermatheca; spc, spermathecal canal." httpUri="https://zenodo.org/record/220513/files/figure.png" pageId="15" pageNumber="116">Fig. 85</figureCitation>
). Stylomere 2 with two or three ensiform setae at the outer margin and three or four at inner margin (
<figureCitation id="1338BD9B5654FFE6C1BAFB39FEE3FB5A" box="[245,330,1196,1220]" captionStart="FIGURES 90 93" captionStartId="31.[151,264,1640,1663]" captionTargetBox="[161,1418,216,1605]" captionTargetId="figure@31.[151,1436,193,1619]" captionTargetPageId="31" captionText="FIGURES 90 93. Left stylomere of ovipositor of Pterostichus (Wraseiellus) spp.: scale bar = 0.5 mm; A, ventral view; B, dorsal view: 90, Pt. diversus (Fairmaire), a female from Yulongshan; 91, Pt. stictopleurus cangshanensis n. ssp., a paratype from Cangshan; 92, Pt. pseudodiversus n. sp., a paratype from Luguhu. 93, Pt. andrewesi Jedlička, a female from Jinfoshan. Abbreviations: es, ensiform setae; ns, nematiform setae; s 1, stylomere 1; s 2, stylomere 2." httpUri="https://zenodo.org/record/220514/files/figure.png" pageId="15" pageNumber="116">Fig. 92</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955654FFE6C188FB5AFE69FAE6" pageId="15" pageNumber="116" type="distribution">
<paragraph id="8BBCA11E5654FFE6C188FB5AFE69FAE6" blockId="15.[151,1437,151,1868]" pageId="15" pageNumber="116">
<emphasis id="B9777D0C5654FFE6C188FB5AFEF0FB76" bold="true" box="[199,345,1231,1256]" pageId="15" pageNumber="116">Distribution</emphasis>
. The distribution range of this species is the widest in the subgenus, including many localities from northwest part of Yunnan: Luguhu (
<geoCoordinate id="EE37C7D95654FFE6C3C1FB61FD47FA92" box="[654,750,1268,1292]" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="555" value="27.63">N27.63º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95654FFE6C3B0FB61FCC3FA92" box="[767,874,1268,1292]" direction="east" orientation="longitude" pageId="15" pageNumber="116" precision="555" value="100.82">E100.82º</geoCoordinate>
), Lanping (
<geoCoordinate id="EE37C7D95654FFE6C2B9FB61FBFFFA92" box="[1014,1110,1268,1292]" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="555" value="26.46">N26.46º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95654FFE6C527FB61FB6DFA92" box="[1128,1220,1268,1292]" direction="east" orientation="longitude" pageId="15" pageNumber="116" precision="555" value="99.25">E99.25º</geoCoordinate>
), Weixi (
<geoCoordinate id="EE37C7D95654FFE6C478FB61FA3EFA92" box="[1335,1431,1268,1292]" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="555" value="27.35">N27.35º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95654FFE6C1D8FA8DFF5DFAAE" box="[151,244,1304,1328]" direction="east" orientation="longitude" pageId="15" pageNumber="116" precision="555" value="99.28">E99.28º</geoCoordinate>
), Deqin (
<geoCoordinate id="EE37C7D95654FFE6C014FA8DFE12FAAE" box="[347,443,1304,1328]" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="555" value="28.37">N28.37º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95654FFE6C088FA8DFD8DFAAE" box="[455,548,1304,1328]" direction="east" orientation="longitude" pageId="15" pageNumber="116" precision="555" value="99.02">E99.02º</geoCoordinate>
), Zhongdian (
<geoCoordinate id="EE37C7D95654FFE6C38EFA8DFC88FAAE" box="[705,801,1304,1328]" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="555" value="27.82">N27.82º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95654FFE6C262FA8DFC23FAAE" box="[813,906,1304,1328]" direction="east" orientation="longitude" pageId="15" pageNumber="116" precision="555" value="99.7">E99.70º</geoCoordinate>
), and Mianbu pass between Muli and Yanyuan (
<geoCoordinate id="EE37C7D95654FFE6C1D1FAA9FF56FACA" box="[158,255,1340,1364]" direction="north" orientation="latitude" pageId="15" pageNumber="116" precision="555" value="27.69">N27.69º</geoCoordinate>
,
<geoCoordinate id="EE37C7D95654FFE6C044FAA9FEDDFACA" box="[267,372,1340,1364]" direction="east" orientation="longitude" pageId="15" pageNumber="116" precision="555" value="101.22">E101.22º</geoCoordinate>
) from south Sichuan.
<taxonomicName id="4C03DA9D5654FFE6C33FFAA8FCEDFACA" box="[624,836,1340,1364]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="pseudodiversus">
<emphasis id="B9777D0C5654FFE6C33FFAA8FCEDFACA" box="[624,836,1340,1364]" italics="true" pageId="15" pageNumber="116">Pt. pseudodiversus</emphasis>
</taxonomicName>
is strictly allopartic with all the other species of this subgenus (
<figureCitation id="1338BD9B5654FFE6C041FAF5FE1AFAE6" box="[270,435,1376,1400]" captionStart="FIGURES 110 111" captionStartId="33.[151,264,1885,1908]" captionTargetBox="[224,1312,207,1827]" captionTargetId="figure@33.[224,1343,193,1844]" captionTargetPageId="33" captionText="FIGURES 110 111. Distribution maps for species of subgenus Pterostichus (Wraseiellus): - Pt. andrewesi Jedlička; « Pt. meyeri Jedlička; ˑ Pt. crassiapex n. sp.; ¿ Pt. comatus n. sp.; ● Pt. diversus (Fairmaire); Ο Pt. pseudodiversus n. sp.; ˔ Pt. stictopleurus stictopleurus (Fairmaire); r Pt. stictopleurus cangshanensis n. ssp.; Pt. kambaiti (Andrewes). 111. shows an enlarged scale of the area in figure 110 marked by the dotted line square." httpUri="https://zenodo.org/record/220516/files/figure.png" pageId="15" pageNumber="116">Figs. 110, 111</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C319F2955654FFE6C188FA11FBE3FA5E" pageId="15" pageNumber="116" type="etymology">
<paragraph id="8BBCA11E5654FFE6C188FA11FBE3FA5E" blockId="15.[151,1437,151,1868]" pageId="15" pageNumber="116">
<emphasis id="B9777D0C5654FFE6C188FA11FEE1FA02" bold="true" box="[199,328,1412,1436]" pageId="15" pageNumber="116">Etymology</emphasis>
. The name &quot;
<taxonomicName id="4C03DA9D5654FFE6C0AEFA11FD27FA02" box="[481,654,1412,1436]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="pseudodiversus">
<emphasis id="B9777D0C5654FFE6C0AEFA11FD27FA02" box="[481,654,1412,1436]" italics="true" pageId="15" pageNumber="116">pseudodiversus</emphasis>
</taxonomicName>
&quot; is the combination of the Greek epithets &quot;
<emphasis id="B9777D0C5654FFE6C5C2FA11FB67FA02" box="[1165,1230,1412,1436]" italics="true" pageId="15" pageNumber="116">pseud</emphasis>
-&quot; and &quot;
<taxonomicName id="4C03DA9D5654FFE6C463FA11FA20FA02" box="[1324,1417,1412,1436]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5654FFE6C463FA11FA20FA02" box="[1324,1417,1412,1436]" italics="true" pageId="15" pageNumber="116">diversus</emphasis>
</taxonomicName>
&quot;, referring to the similarity of this new species with
<taxonomicName id="4C03DA9D5654FFE6C383FA3DFBEEFA5E" authority="Fairmaire" authorityName="Fairmaire" box="[716,1095,1448,1472]" class="Insecta" family="Carabidae" genus="Pterostichus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="116" phylum="Arthropoda" rank="species" species="diversus">
<emphasis id="B9777D0C5654FFE6C383FA3DFC15FA5E" box="[716,956,1448,1472]" italics="true" pageId="15" pageNumber="116">Pterostichus diversus</emphasis>
(Fairmaire)
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C319F2955654FFE6C188FA59FDEFF8D2" pageId="15" pageNumber="116" type="discussion">
<paragraph id="8BBCA11E5654FFE6C188FA59FDEFF8D2" blockId="15.[151,1437,151,1868]" pageId="15" pageNumber="116">
<emphasis id="B9777D0C5654FFE6C188FA59FE9DFA7A" bold="true" box="[199,308,1484,1508]" pageId="15" pageNumber="116">Remarks</emphasis>
. In this species, the shape of apical lamella of male genitalia varies from individual to individual. In general, specimens from Zhongdian and Deqin have the apex more rounded (
<figureCitation id="1338BD9B5654FFE6C564FA65FB15F996" box="[1067,1212,1520,1544]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Figs. 54, 55</figureCitation>
), while those from Lanping, Luguhu and Muli more truncate (
<figureCitation id="1338BD9B5654FFE6C334F981FCACF9B2" box="[635,773,1556,1580]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Figs. 4553</figureCitation>
). Moreover, the only examined male from Weixi, which is somewhat different from the others, has the slenderest apical lamella (
<figureCitation id="1338BD9B5654FFE6C2E5F9ADFC57F9CE" box="[938,1022,1592,1616]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Fig. 56</figureCitation>
). But even in the same locality (such as specimens from Luguhu), the shape of apical lamella is not constant, from rounded to slightly truncate, and from rather narrow to somewhat wider (
<figureCitation id="1338BD9B5654FFE6C355F915FD0DF906" box="[538,676,1664,1688]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">Figs. 4550</figureCitation>
). Similar variation also presents in the shape of pronotum, narrower or wider, varying both in population and individual level. Since there is no constant difference both in male genital and external characters among specimens from different localities, we do not try to divide this species into different subspecies in the present paper, and include the majority of examined specimens from five localities (see above) in the
<typeStatus id="54B81FBC5654FFE6C18CF884FF5AF8B7" box="[195,243,1809,1833]" pageId="15" pageNumber="116">type</typeStatus>
series except those three specimens (
<specimenCount id="9D056A975654FFE6C3E6F885FD5CF8B6" box="[681,757,1808,1832]" pageId="15" pageNumber="116" type="male">1 male</specimenCount>
and
<specimenCount id="9D056A975654FFE6C27EF885FC08F8B6" box="[817,929,1808,1832]" pageId="15" pageNumber="116" type="female">2 females</specimenCount>
) from Weixi which were excluded for their slenderer apical lamella (
<figureCitation id="1338BD9B5654FFE6C0FFF8A1FDA4F8D2" box="[432,525,1844,1868]" captionStart="FIGURES 35 40" captionStartId="26.[151,264,1759,1782]" captionTargetBox="[168,1419,197,1735]" captionTargetId="figure@26.[151,1436,193,1738]" captionTargetPageId="26" captionText="FIGURES 35 40. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp.: scale bar = 1.0 mm; A, left lateral view of median lobe; B, dorsal view of median lobe; 35, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 36, Pt. stictopleurus cangshanensis n. ssp., holotype; 37, Pt. pseudodiversus n. sp., holotype; 38, Pt. pseudodiversus n. sp., a paratype from Muli; 39, Pt. pseudodiversus n. sp., a male from Weixi; 40, Pt. kambaiti (Andrewes), holotype of Pterostichus kambaiti Jedlička." httpUri="https://zenodo.org/record/220509/files/figure.png" pageId="15" pageNumber="116">Figs. 39</figureCitation>
,
<figureCitation id="1338BD9B5654FFE6C356F8A1FD90F8D2" box="[537,569,1844,1868]" captionStart="FIGURES 41 56" captionStartId="27.[151,264,1385,1408]" captionTargetBox="[153,1417,210,1351]" captionTargetId="figure@27.[151,1436,193,1364]" captionTargetPageId="27" captionText="FIGURES 41 56. Median lobe of male genitalia of Pterostichus (Wraseiellus) spp., dorsal view on apical lamella: scale bar = 0.5 mm; 41, Pt. diversus (Fairmaire), a male from Yulongshan; 42, Pt. diversus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 43, Pt. stictopleurus (Fairmaire), a male from &quot; Ta-pin-tze &quot;; 44, Pt. stictopleurus cangshanensis n. ssp., holotype; 45, Pt. pseudodiversus n. sp., holotype; 46 50, Pt. pseudodiversus n. sp., paratypes from Luguhu; 51, Pt. pseudodiversus n. sp., a paratype from Lanping; 52 53, Pt. pseudodiversus n. sp., paratypes from Muli; 54, Pt. pseudodiversus n. sp., a paratype from Zhongdian; 55, Pt. pseudodiversus n. sp., a paratype from Deqin; 56, Pt. pseudodiversus n. sp., a male from Weixi." httpUri="https://zenodo.org/record/220510/files/figure.png" pageId="15" pageNumber="116">56</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>