treatments-xml/data/FA/86/E5/FA86E567BEE65F9082EC1D397E8C6468.xml
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1150.93760" ID-Pensoft-Pub="1313-2970-1150-1" ID-Pensoft-UUID="2D0F861C78665B9BABB241437CA5ED53" ID-ZooBank="A38C5FB69F664F858788AAA53D21704D" ModsDocID="1313-2970-1150-1" checkinTime="1677664009133" checkinUser="pensoft" docAuthor="Crews, Sarah C." docDate="2023" docId="FA86E567BEE65F9082EC1D397E8C6468" docLanguage="en" docName="ZooKeys 1150: 1-189" docOrigin="ZooKeys 1150" docPubDate="2023-02-27" docSource="http://dx.doi.org/10.3897/zookeys.1150.93760" docTitle="Karaops karrawarla Crews &amp; Harvey 2011" docType="treatment" docVersion="1" id="2D0F861C78665B9BABB241437CA5ED53" lastPageNumber="1" masterDocId="2D0F861C78665B9BABB241437CA5ED53" masterDocTitle="But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia" masterLastPageNumber="189" masterPageNumber="1" pageNumber="1" updateTime="1677664009133" updateUser="pensoft">
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<mods:titleInfo>
<mods:title>But wait, there's more! Descriptions of new species and undescribed sexes of flattie spiders (Araneae, Selenopidae, Karaops) from Australia</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Crews, Sarah C.</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-9360-6236</mods:nameIdentifier>
<mods:affiliation>California Academy of Sciences, Department of Entomology, 55 Music Concourse Drive, San Francisco, CA, 94118, USA</mods:affiliation>
<mods:nameIdentifier type="email">screwsemail@gmail.com</mods:nameIdentifier>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2023</mods:date>
<mods:detail type="pubDate">
<mods:number>2023-02-27</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>1150</mods:number>
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<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>189</mods:end>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1150.93760</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1150.93760</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-1150-1</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:FA86E567BEE65F9082EC1D397E8C6468" httpUri="http://treatment.plazi.org/id/FA86E567BEE65F9082EC1D397E8C6468" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName ID-CoL="3R2MD" LSID="FA86E567-BEE6-5F90-82EC-1D397E8C6468" authority="Crews &amp; Harvey, 2011" authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops karrawarla" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="karrawarla">Karaops karrawarla Crews &amp; Harvey, 2011</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 55" captionStartId="F64" captionText="Figure 55. Members of the Pilbara-Gascoyne species group, Western Australia A Karaops joehaeneri sp. nov., paratype male, Escarpment Trail, Kennedy Range National Park (sel _ 1131, WAM T 155507) B same, holotype female, epigyne (sel _ 1130 WAM T 155506) C same, endogyne D Karaops karrawarla, female paratype, epigyne, Bush Bay, Western Australia (WAM T 76700) E same, endogyne. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure55" httpUri="https://binary.pensoft.net/fig/816444" pageId="0" pageNumber="1">Figs 55D, E</figureCitation>
<figureCitation captionStart="Figure 58" captionStartId="F67" captionText="Figure 58. Karaops karrawarla, Bush Bay, Western Australia A male holotype, palp, ventral (WAM T 55001) B female paratype (WAM T 76700) C male holotype (WAM T 55001) D close up of embolus tip as indicated by arrow in E, below E palp, retrolateral F palp, expanded, light gray = tip of conductor, dark gray = median apophysis. Scale bars: 0.5 mm (A, E, F); 2 mm (B, C)." figureDoi="10.3897/zookeys.1150.93760.figure58" httpUri="https://binary.pensoft.net/fig/816447" pageId="0" pageNumber="1">, 58A-F</figureCitation>
<figureCitation captionStart="Map 1" captionStartId="F3" captionText="Map 1. Map of Australia showing areas that are expanded in subsequent figures. CD = Central Desert group, D = Karaops dawara group, F = Karaops francesae group, K = Kimberley group, UA = species that are not assigned to a group, PG = Pilbara-Gascoyne group." figureDoi="10.3897/zookeys.1150.93760.map1" httpUri="https://binary.pensoft.net/fig/816383" pageId="0" pageNumber="1">, Maps 1</figureCitation>
<figureCitation captionStart="Map 9" captionStartId="F57" captionText="Map 9. Species of the Pilbara-Gascoyne species group A diamonds (juveniles): blue = Erawandoo Hill, orange = Mt. Augustus, red = Nammuldi, yellow = Hillside-Corunna-Glenn Herring, black = Marble Bar-Glacier Valley, pink = Orebody 19, 31, teal = Wheelara North, orange with thick outline = Harding Dam (maybe Karaops ngarluma, see text); Circles: dark blue = Karaops julianneae, dark green = Karaops badgeradda, red = Karaops joehaeneri sp. nov., teal = Karaops karrawarla, blue = Karaops durrantorum sp. nov., brown = Karaops burbidgei, green = Karaops martamarta, purple = Karaops nyangumarta, black = Karaops banyjima, yellow = Karaops morganoconnelli sp. nov., gray = Karaops nyiyaparli, red-brown = Karaops feedtime, bright pink = Karaops nyamal (probably, see text), white = Karaops kariyarra, bright green = Karaops yurlburr, orange = Karaops ngarluma, tan = Karaops jaburrara, peach = Karaops forteyi, teal = new species yet undescribed; small white diamonds are unidentifiable immatures for which no molecular data are available B boxed area expanded for clarity." figureDoi="10.3897/zookeys.1150.93760.map9" httpUri="https://binary.pensoft.net/fig/816437" pageId="0" pageNumber="1">, 9A</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="reference_group">
<paragraph pageId="0" pageNumber="1">
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops karrawarla" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="karrawarla">Karaops karrawarla</taxonomicName>
Crews &amp; Harvey, 2011: 51, figs 39-42 (♂♀, examined).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species can be differentiated from other similar species by the genitalia. The copulatory openings are located in the middle of the epigynal plate beneath an m-shaped margin. The lateral lobes can be easily discerned (Fig.
<figureCitation captionStart="Figure 55" captionStartId="F64" captionText="Figure 55. Members of the Pilbara-Gascoyne species group, Western Australia A Karaops joehaeneri sp. nov., paratype male, Escarpment Trail, Kennedy Range National Park (sel _ 1131, WAM T 155507) B same, holotype female, epigyne (sel _ 1130 WAM T 155506) C same, endogyne D Karaops karrawarla, female paratype, epigyne, Bush Bay, Western Australia (WAM T 76700) E same, endogyne. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure55" httpUri="https://binary.pensoft.net/fig/816444" pageId="0" pageNumber="1">55B, C</figureCitation>
). The male can be distinguished by the embolus that widens slightly at the tip and the position of the conductor and median apophysis after expansion (Fig.
<figureCitation captionStart="Figure 58" captionStartId="F67" captionText="Figure 58. Karaops karrawarla, Bush Bay, Western Australia A male holotype, palp, ventral (WAM T 55001) B female paratype (WAM T 76700) C male holotype (WAM T 55001) D close up of embolus tip as indicated by arrow in E, below E palp, retrolateral F palp, expanded, light gray = tip of conductor, dark gray = median apophysis. Scale bars: 0.5 mm (A, E, F); 2 mm (B, C)." figureDoi="10.3897/zookeys.1150.93760.figure58" httpUri="https://binary.pensoft.net/fig/816447" pageId="0" pageNumber="1">58A, D-F</figureCitation>
).
</paragraph>
<caption doi="10.3897/zookeys.1150.93760.figure58" httpUri="https://binary.pensoft.net/fig/816447" pageId="0" pageNumber="1" start="Figure 58" startId="F67">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 58.</emphasis>
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops karrawarla" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="karrawarla">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops karrawarla</emphasis>
</taxonomicName>
, Bush Bay, Western Australia
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
male holotype, palp, ventral (WAM T55001)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
female paratype (WAM T76700)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
male holotype (WAM T55001)
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
close up of embolus tip as indicated by arrow in E, below
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
palp, retrolateral
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
palp, expanded, light gray = tip of conductor, dark gray = median apophysis. Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">A, E, F</emphasis>
); 2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
The description of the male and female can be found in
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.99.723" author="Crews, SC" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="1 - 103" refId="B14" refString="Crews, SC, Harvey, MS, 2011. The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region. ZooKeys 99: 1 - 103, DOI: https://doi.org/10.3897/zookeys.99.723" title="The spider family Selenopidae (Arachnida, Araneae) in Australasia and the Oriental Region." url="https://doi.org/10.3897/zookeys.99.723" volume="99" year="2011">Crews and Harvey (2011)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is only known from the type locality, Bush Bay in the Gascoyne Region, Western Australia (Map
<figureCitation captionStart="Map 9" captionStartId="F57" captionText="Map 9. Species of the Pilbara-Gascoyne species group A diamonds (juveniles): blue = Erawandoo Hill, orange = Mt. Augustus, red = Nammuldi, yellow = Hillside-Corunna-Glenn Herring, black = Marble Bar-Glacier Valley, pink = Orebody 19, 31, teal = Wheelara North, orange with thick outline = Harding Dam (maybe Karaops ngarluma, see text); Circles: dark blue = Karaops julianneae, dark green = Karaops badgeradda, red = Karaops joehaeneri sp. nov., teal = Karaops karrawarla, blue = Karaops durrantorum sp. nov., brown = Karaops burbidgei, green = Karaops martamarta, purple = Karaops nyangumarta, black = Karaops banyjima, yellow = Karaops morganoconnelli sp. nov., gray = Karaops nyiyaparli, red-brown = Karaops feedtime, bright pink = Karaops nyamal (probably, see text), white = Karaops kariyarra, bright green = Karaops yurlburr, orange = Karaops ngarluma, tan = Karaops jaburrara, peach = Karaops forteyi, teal = new species yet undescribed; small white diamonds are unidentifiable immatures for which no molecular data are available B boxed area expanded for clarity." figureDoi="10.3897/zookeys.1150.93760.map9" httpUri="https://binary.pensoft.net/fig/816437" pageId="0" pageNumber="1">9A</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="natural history">
<paragraph pageId="0" pageNumber="1">Natural history.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is known from the Carnarvon bioregion, Wooramel subregion of the Gascoyne, as is
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops joehaeneri" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="joehaeneri">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops joehaeneri</emphasis>
</taxonomicName>
sp. nov. For additional information about the bioregion and subregion, see discussion for
<taxonomicName lsidName="K. joehaeneri" pageId="0" pageNumber="1" rank="species" species="joehaeneri">
<emphasis italics="true" pageId="0" pageNumber="1">K. joehaeneri</emphasis>
</taxonomicName>
sp. nov. above.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="discussion">
<paragraph pageId="0" pageNumber="1">Discussion.</paragraph>
<paragraph pageId="0" pageNumber="1">
The genitalia of both the male and the female of
<taxonomicName authorityName="Crews &amp; Harvey" authorityYear="2011" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops karrawarla" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="karrawarla">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops karrawarla</emphasis>
</taxonomicName>
(Figs
<figureCitation captionStart="Figure 55" captionStartId="F64" captionText="Figure 55. Members of the Pilbara-Gascoyne species group, Western Australia A Karaops joehaeneri sp. nov., paratype male, Escarpment Trail, Kennedy Range National Park (sel _ 1131, WAM T 155507) B same, holotype female, epigyne (sel _ 1130 WAM T 155506) C same, endogyne D Karaops karrawarla, female paratype, epigyne, Bush Bay, Western Australia (WAM T 76700) E same, endogyne. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure55" httpUri="https://binary.pensoft.net/fig/816444" pageId="0" pageNumber="1">55D, E</figureCitation>
,
<figureCitation captionStart="Figure 58" captionStartId="F67" captionText="Figure 58. Karaops karrawarla, Bush Bay, Western Australia A male holotype, palp, ventral (WAM T 55001) B female paratype (WAM T 76700) C male holotype (WAM T 55001) D close up of embolus tip as indicated by arrow in E, below E palp, retrolateral F palp, expanded, light gray = tip of conductor, dark gray = median apophysis. Scale bars: 0.5 mm (A, E, F); 2 mm (B, C)." figureDoi="10.3897/zookeys.1150.93760.figure58" httpUri="https://binary.pensoft.net/fig/816447" pageId="0" pageNumber="1">58A-F</figureCitation>
) are re-figured for ease of comparison with
<taxonomicName lsidName="K. joehaeneri" pageId="0" pageNumber="1" rank="species" species="joehaeneri">
<emphasis italics="true" pageId="0" pageNumber="1">K. joehaeneri</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName lsidName="K. morganoconnelli" pageId="0" pageNumber="1" rank="species" species="morganoconnelli">
<emphasis italics="true" pageId="0" pageNumber="1">K. morganoconnelli</emphasis>
</taxonomicName>
sp. nov. as they closely resemble these species (Figs
<figureCitation captionStart="Figure 55" captionStartId="F64" captionText="Figure 55. Members of the Pilbara-Gascoyne species group, Western Australia A Karaops joehaeneri sp. nov., paratype male, Escarpment Trail, Kennedy Range National Park (sel _ 1131, WAM T 155507) B same, holotype female, epigyne (sel _ 1130 WAM T 155506) C same, endogyne D Karaops karrawarla, female paratype, epigyne, Bush Bay, Western Australia (WAM T 76700) E same, endogyne. Scale bars: 0.5 mm." figureDoi="10.3897/zookeys.1150.93760.figure55" httpUri="https://binary.pensoft.net/fig/816444" pageId="0" pageNumber="1">55D, E</figureCitation>
,
<figureCitation captionStart="Figure 58" captionStartId="F67" captionText="Figure 58. Karaops karrawarla, Bush Bay, Western Australia A male holotype, palp, ventral (WAM T 55001) B female paratype (WAM T 76700) C male holotype (WAM T 55001) D close up of embolus tip as indicated by arrow in E, below E palp, retrolateral F palp, expanded, light gray = tip of conductor, dark gray = median apophysis. Scale bars: 0.5 mm (A, E, F); 2 mm (B, C)." figureDoi="10.3897/zookeys.1150.93760.figure58" httpUri="https://binary.pensoft.net/fig/816447" pageId="0" pageNumber="1">58A, D-F</figureCitation>
,
<figureCitation captionStart="Figure 59" captionStartId="F68" captionText="Figure 59. Karaops morganoconnelli sp. nov. and Karaops nyamal from the Pilbara-Gascoyne species group A Karaops morganoconnelli sp. nov., paratype male, Cathedral Gorge, ~ 40 km from Newman (sel _ 1207, WAM T 155583) B same, holotype female, epigyne, Orebody 24, ~ 7 km N of Newman (WAM T 131095) C same, endogyne D same, paratype male Cathedral Gorge, ~ 40 km from Newman (sel _ 1207, WAM T 155583) E same, holotype female, Orebody 24, ~ 7 km N of Newman (WAM T 131095) F Karaops nyamal, paratype female, Mt. Webber (WAM T 107698). Scale bars: 0.5 mm (B, C); 1 mm (D); 2 mm (F); 5 mm (E)." figureDoi="10.3897/zookeys.1150.93760.figure59" httpUri="https://binary.pensoft.net/fig/816448" pageId="0" pageNumber="1">59B, C</figureCitation>
). The types were collected in a pitfall trap set in January and collected in May, so there is no way to pinpoint the time of year adults are present other than in the first five months of the year. January-May begins at the hottest and wettest time in the area, but by April and May, temperatures begin to drop, yet rainfall remains at the highest (Suppl. material 2: table S1).
</paragraph>
<caption doi="10.3897/zookeys.1150.93760.figure59" httpUri="https://binary.pensoft.net/fig/816448" pageId="0" pageNumber="1" start="Figure 59" startId="F68">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 59.</emphasis>
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops morganoconnelli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="morganoconnelli">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops morganoconnelli</emphasis>
</taxonomicName>
sp. nov. and
<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyamal" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyamal">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyamal</emphasis>
</taxonomicName>
from the Pilbara-Gascoyne species group
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
<taxonomicName authorityName="Crews" authorityYear="2023" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops morganoconnelli" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="morganoconnelli">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops morganoconnelli</emphasis>
</taxonomicName>
sp. nov., paratype male, Cathedral Gorge, ~ 40 km from Newman (sel_1207, WAM T155583)
<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
same, holotype female, epigyne, Orebody 24, ~ 7 km N of Newman (WAM T131095)
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
same, endogyne
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
same, paratype male Cathedral Gorge, ~ 40 km from Newman (sel_1207, WAM T155583)
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
same, holotype female, Orebody 24, ~ 7 km N of Newman (WAM T131095)
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
<taxonomicName authorityName="Crews" authorityYear="2013" class="Arachnida" family="Selenopidae" genus="Karaops" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Karaops nyamal" order="Araneae" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="nyamal">
<emphasis italics="true" pageId="0" pageNumber="1">Karaops nyamal</emphasis>
</taxonomicName>
, paratype female, Mt. Webber (WAM T107698). Scale bars: 0.5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">B, C</emphasis>
); 1 mm (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
); 2 mm (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); 5 mm (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
).
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">Upon visiting the type locality, it does not appear to be good habitat for selenopid species, as there are no rocks and no trees at the locality or in the vicinity. The collection data have been confirmed to be correct. It has been suggested (J. Waldock, pers. comm.) that the specimens could have washed down to the area from further inland.</paragraph>
</subSubSection>
</treatment>
</document>